DOCSLIB.ORG

Review of Mesozoic and Cenozoic Sea Turtles from the Former Ussr

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Review of Mesozoic and Cenozoic Sea Turtles from the Former Ussr Russian Journal of Herpetology Vol. 9, No. 2, 2002, pp. 137 – 154 REVIEW OF MESOZOIC AND CENOZOIC SEA TURTLES FROM THE FORMER USSR Alexander O. Averianov1 Submitted December 17, 2001. A list of 51 localities of fossil sea turtles from the Middle Jurassic, Early and Late Cretaceous, Paleocene, Eocene and Miocene of Russia, Uzbekistan, Kazakhstan, Tajikistan, and Ukraine is provided. For four lo- calities in Tajikistan, Kazakhstan, and Ukraine presence of sea turtles is not confirmed by current data. Two new taxa, Dollochelys rogovichi sp.nov. (Cheloniidae, Eocene, Ukraine) and Turgaiscapha kushmu- runica gen. et sp.nov. (Chelonioidea indet., cf. Dermochelyidae, Late Cretaceous, Kazakhstan), are de- scribed. Key words: Cheloniidae, Chelonioidea, Dermochelyidae, Protostegidae, sea turtles, Thalassemydidae. INTRODUCTION St. Petersburg, Russia; EIMIZ, Natural History Mu- seum named after Zardabi, Baku, Azerbaijan; IPG, Turtles entered the open sea several times since Institute of Paleobiology, Georgian Academy of Sci- the beginning of their geologic history. Pronounced ences, Tbilisi, Georgia; IZK, Institute of Zoology, adaptation to this new environment (fore limbs modi- Kazakhstan Academy of Science, Alma-Ata, Ka- fied to swimming, shell lightened by costoperipheral zakhstan; IZU, Institute of Zoology, Ukraine Acad- and plastral fontanels) are found in Late Jurassic Eu- emy of Science, Kiev, Ukraine; PIN, Paleontological ropean Eurysternidae and Early Cretaceous to Recent Institute, Russian Academy of Sciences, Moscow, globally distributed Chelonioidea (monophyletic Russia; ZISP PH (or ZISP PHT), Paleoherpetologi- group, sensu Gaffney and Meylan, 1988). This re- cal collection, Zoological Institute, Russian Acad- view includes all fossil sea turtles localities for the emy of Sciences, St. Petersburg, Russia. territory of the USSR, as known previously, as well as reported here for the first time. Majority of these localities confined to the areas of the Cretaceous- REVIEW OF THE MESOZOIC Paleogene Tethys Ocean, Cretaceous Turgai Straight, AND CENOZOIC SEA TURTLES epicontinental sea of the Russian Platform, and Neo- LOCALITIES gene Paratethys basins (Fig. 1; Table 1). In the list of localities references to all descriptions and illustra- Jurassic tions of the material are given. This review does not include some unpublished materials, currently stud- 1. Peski, Moscow Region, Russia. ied by Prof. V. B. Sukhanov, by Dr. A. A. Yarkov, Geology. Black clays of Moskva Reka Forma- and by the author. Yaxartemys longicauda Riabinin, tion, Meshera Horizon, filling fissures in the Carbon- 1948, known from the juvenile specimens only from iferous limestone. Late Jurassic of Kazakhstan, originally referred to Age. Middle Jurassic, Bajocian-Bathonian. Thalassemydidae (Riabinin, 1948), possibly belongs Material and references. “Bones of skeleton to the Xinjiangchelyidae, a group of Asiatic fresh- and shell fragments of a turtle” (Krupina, 1995:129). water and estuarine waters turtles (Kaznyshkin et al., New genus and species of “Triassic generation” tur- 1990), which are not considered here. tles (Novikov et al. 1998:58). A new genus of Cryp- Collection abbreviations. CCMGE, Cherny- todira “more archaic than the Early Jurassic Kayenta- shev’s Central Museum of Geological Exploration, chelys” (Alifanov and Sennikov, 2001:75). This is an archaic sea turtle, not closely related to other sea tur- 1 Zoological Institute of the Russian Academy of Sciences, Uni- tles, recently described as Heckerochelys romani Su- versitetskaya nab. 1, St. Petersburg 199034, Russia. khanov, 2001 and referred to a new family Heckero- 1026-2296/2002/0902-0137 © 2002 Folium Publishing Company 138 Alexander O. Averianov Fig. 1. Localities of the fossil sea turtles for the territory of the USSR. Localities: 1, Peski; 2, Motnya 1; 3, Khodzhakul’ 1; 4, Gubkin; 5, Sheikhdzheili 2; 6, Khodzhakul’sai; 7, Itemir; 8, Chukhonastovka; 9, Bagaevka; 10, Saratov; 11, Nogaity; 12, Polunino; 13, Alymtau; 14, Kyrkkuduk 2; 15, Malaya Serdoba; 16, Rychkovo; 17, Zhuravlevskii; 18, Kushmurun; 19, Malaya Ivanovka; 20, Karpovka; 21, Bakh- chisarai; 22, Vostochnyi; 23, Kol’tso; 24, Darbaza 1 and 2; 25, Dzhylga 1a; 26, Dzhylga 1b and 2; 27, Andarak 2; 28, Getkilnin; 29, Dzhe- roi 1; 30, Dzheroi 2; 31, Khanabad; 32, Maisk; 33, Kentau; 35, Bakhmutovka; 36, Krasnorechenskoe; 37, Kiev; 38, Pirogovo; 39, Koktur- nak; 40, Prolom; 43, Otradnaya; 44, Abadzekhskaya; 45, Tchelkar-Teniz; 47, Perekischkyul’; 48, Agyspe; 50, Belomechetskaya; 51, Dzaudzhikau. The remaining localities cannot be located with confidence. chelyidae (Sukhanov, 2001). Material in the PIN 3. Khodzhakul’ 1, Karakalpakistan, Uzbekistan. collection. Geology. Lower or middle part of the Khodzhakul’ Formation. Early Cretaceous Age. Early Cretaceous, late Albian (possibly, Late Cretaceous, early Cenomanian). 2. Motnya 1, Buryatiya, Russia. Geology. Lower part of Khilok Formation. Material and references. Basisphenoid, CCMGE 2/11478, the holotype of Oxemys gutta Age. Early Cretaceous, Berriasian?. Nessov, 1977 (Nessov, 1977b:47, Figs. V and G; Material and references. Two bridge peripher- als (ZISP PHT B59 – 11) and part of the plastron 1987b, pl. 2, Fig. 6). “Oxemys gutta (Chelonioidea) (?hypoplastron; ZISP PHT B59-12) of “?Chelonio- (Nessov, 1977a:156). “Some Chelonioidea (genus idea fam., gen. et sp. indet.” (Nessov and Khosatzky, Oxemys) (Nessov and Khosatzky, 1977:158). “Oxe- 1981:77, pl. 4, Figs. 11 and 12). “Some poorly deter- mys gutta Ness. (Chelonioidea incertae sedis)” (Ne- mined forms (Chelonioidea?), small and middle- ssov, 1984:216). “Oxemys gutta Ness.” (Nessov, sized” (Nessov, 1984:216). “Chelonioidea (?) indet.” 1988b:96). “Turtles Chelonioidea...: Oxemys gutta” (Nessov, 1997:121, pl. 28, Figs. 13 and 14). (Nessov, 1997:135, pl. 27, Fig. 1). Review of Mesozoic and Cenozoic Sea Turtles from the Former USSR 139 Comments. The basisphenoid of O. gutta is TABLE 1. Taxa of the Sea Turtles Found in the Mesozoic and Ce- clearly of chelonioid type. The rostrum basisphenoi- nozoic of the Former USSR (Numbers of Localities Correspond to Those in the Text and Fig. 1). dale much reduced. The sella turcica obliterated. The Localities (* indicates foramina anterior canalis carotici interni lie close to- Taxon gether (separated by the 1.5 mm distance). The dor- a type locality) sum sellae high, not overhanging the sella turcica, Jurassic and separated from the latter by a bone space with a Heckerochelys romani 1* saggittal crest. The systematic position of this turtle Cretaceous remains enigmatic. Among thousands of turtle shell Oxemys gutta 3* fragments from the Khodzhakul’ locality none could cf. Oxemys sp. 5, 6 be attributed to a chelonioid. The turtle assemblage is Parathalassemys cava 7* dominated by macrobaenid-sinemydid (Kirgizemys, Chelonioidea indet. 2, 10, 12 14, 16, 17 Tienfucheloides) and trionychoid (Adocus, Fergane- Cheloniidae indet. 14 mys, Kizylkumemys, Trionychidae indet.) forms, all Dermochelyidae indet. 4 of which have quite distinct structure of basisphenoid Turgaiscapha kushmurunica 18* Protostegidae indet. 9, 11, 15 (known for trionychoids mentioned, for macrobae- Teguliscapha rossica 4* nids Hangaiemys, Ordosemys, Dracochelys, and si- Teguliscapha(?) sp. 8 nemydid Sinemys). Cenozoic Chelonioidea indet. 21–23, 28, 29, 33–36, 38, 39, 4. Gubkin and Staryi Oskol, 41, 42, 49, 50 Belgorod Region, Russia. Cheloniidae indet. 27, 30 32, 40, 43, 44, 48, 51 Geology. Phosphorite conglomerate within yel- Osteopyginae indet. 20 low and gray sands of the Sekmenov Formation, out- Tasbacka aldabergeni 24, 25*, 26 cropped in the ore quarries Lebedinskyi and Stoilen- cf. Tasbacka sp. 19 Dollochelys rogovichi 37* skii near the cities Gubkin and Staryi Oskol respec- Glarichelys gwinneri 45–47 tively. Age. Early – Late Cretaceous, late Albian — early Cenomanian. 1980 – 1994, by A. O. Averianov and his colleagues Material and references. “Sea turtles in 1991 – 2000. Chelosp[h]arginae” (Nessov, 1985:148). “Plastron, carapace and skull elements, dentary (with moder- Late Cretaceous ately elongated symphysis, without saggittal crest)... of small-sized, primitive, ... oldest known Protostegi- 5. Sheikhdzheili 2, Karakalpakistan, Uzbekistan. dae (Chelospharginae)” (Nessov, 1987a:77). “Chelo- Geology. Upper part of the Khodzhakul’ Forma- spharginae (Teguliscapha)” (Glickman et al., 1987: tion. 258). “Sea turtles of the genus Teguliscapha Nessov, Age. Late Cretaceous, early Cenomanian. 1987 [sic]... relatively advanced in lower jaw struc- Material and references. “Testudines indet.: cf. ture... but retained primitive humerus of thalassoid Oxemys sp.” (Nessov, 1997:139). type” (Nessov, 1987b:89, Fig. 4). “Chelosphargi- 6. Khodzhakul’sai, Karakalpakistan, Uzbekistan. nae... with narrow long shell, thalassoid humerus... Geology. Upper part of the Khodzhakul’ Forma- referred to a new genus Teguliscapha Nessov, tion. gen.nov.” (Nessov et al., 1988:130, Figs. 1(17), 2). Age. Late Cretaceous, early Cenomanian. “Turtles Protostegidae and other [sea turtles]” (Ne- Material and references. “Testudines indet.: cf. ssov, 1997:126). The frontal, parietal, opisthotic, Oxemys sp.” (Nessov, 1997:140). some other cranial fragments, dentary, a number of shell fragments of medium-sized Teguliscapha rossi- 7. Itemir, Bukhara Viloyat, Uzbekistan. ca Nessov, 1988 (Protostegidae, Chelospharginae) Geology. Kulbecke Formation. and much rarer shell fragments of a gigantic archaic Age. Early(?)-Late Cretaceous, late Albian? Dermochelyidae indet. The material (CCMGE
Load more

Recommended publications
  • First Turtle Remains from the Middle-Late Jurassic Yanliao Biota, NE China
    Vol.7, No.1 pages 1-11 Science Technology and Engineering Journal (STEJ) Research Article First Turtle Remains from the Middle-Late Jurassic Yanliao Biota, NE China Lu Li1,2,3, Jialiang Zhang4, Xiaolin Wang1,2,3, Yuan Wang1,2,3 and Haiyan Tong1,5* 1 Laboratory of Vertebrate Evolution and Human Origins of the Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China 2 CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China 3 College of Earth and Planetary Sciences, University of China Academy of Sciences, Beijing 100044, China 4 State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Beijing 100083, China 5 Palaeontological Research and Education Centre, Mahasarakham University, Kantarawichai, Maha Sarakham 44150, Thailand * Corresponding author: [email protected] (Received: 14th August 2020, Revised: 11th January 2021, Accepted: 9th March 2021) Abstract - The Middle-Late Jurassic Yanliao Biota, preceding the Early Cretaceous Jehol Biota in NE China has yielded a rich collection of plant, invertebrate and vertebrate fossils. But contrary to the Jehol Biota which is rich in freshwater vertebrates, in the Yanliao Biota the aquatic reptiles are absent, and turtles have not been reported so far. In this paper, we report on the first turtle remains from the Yanliao Biota. The material consists of a partial skeleton from the Upper Jurassic Tiaojishan Formation of Bawanggou site (Qinglong, Hebei Province, China). Characterized by a broad skull with a pair of sulci carotici and a remnant of an interpterygoid vacuity, a well-developed anterior lobe of the plastron with mesiolaterally elongated epiplastra and a relatively large oval entoplastron; it is assigned to Annemys sp.
    [Show full text]
  • A New Xinjiangchelyid Turtle from the Middle Jurassic of Xinjiang, China and the Evolution of the Basipterygoid Process in Mesozoic Turtles Rabi Et Al
    A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles Rabi et al. Rabi et al. BMC Evolutionary Biology 2013, 13:203 http://www.biomedcentral.com/1471-2148/13/203 Rabi et al. BMC Evolutionary Biology 2013, 13:203 http://www.biomedcentral.com/1471-2148/13/203 RESEARCH ARTICLE Open Access A new xinjiangchelyid turtle from the Middle Jurassic of Xinjiang, China and the evolution of the basipterygoid process in Mesozoic turtles Márton Rabi1,2*, Chang-Fu Zhou3, Oliver Wings4, Sun Ge3 and Walter G Joyce1,5 Abstract Background: Most turtles from the Middle and Late Jurassic of Asia are referred to the newly defined clade Xinjiangchelyidae, a group of mostly shell-based, generalized, small to mid-sized aquatic froms that are widely considered to represent the stem lineage of Cryptodira. Xinjiangchelyids provide us with great insights into the plesiomorphic anatomy of crown-cryptodires, the most diverse group of living turtles, and they are particularly relevant for understanding the origin and early divergence of the primary clades of extant turtles. Results: Exceptionally complete new xinjiangchelyid material from the ?Qigu Formation of the Turpan Basin (Xinjiang Autonomous Province, China) provides new insights into the anatomy of this group and is assigned to Xinjiangchelys wusu n. sp. A phylogenetic analysis places Xinjiangchelys wusu n. sp. in a monophyletic polytomy with other xinjiangchelyids, including Xinjiangchelys junggarensis, X. radiplicatoides, X. levensis and X. latiens. However, the analysis supports the unorthodox, though tentative placement of xinjiangchelyids and sinemydids outside of crown-group Testudines. A particularly interesting new observation is that the skull of this xinjiangchelyid retains such primitive features as a reduced interpterygoid vacuity and basipterygoid processes.
    [Show full text]
  • Interpreting Character Variation in Turtles: [I]Araripemys Barretoi
    A peer-reviewed version of this preprint was published in PeerJ on 29 September 2020. View the peer-reviewed version (peerj.com/articles/9840), which is the preferred citable publication unless you specifically need to cite this preprint. Limaverde S, Pêgas RV, Damasceno R, Villa C, Oliveira GR, Bonde N, Leal MEC. 2020. Interpreting character variation in turtles: Araripemys barretoi (Pleurodira: Pelomedusoides) from the Araripe Basin, Early Cretaceous of Northeastern Brazil. PeerJ 8:e9840 https://doi.org/10.7717/peerj.9840 Interpreting character variation in turtles: Araripemys barretoi (Pleurodira: Pelomedusoides) from the Araripe Basin, Early Cretaceous of Northeastern Brazil Saulo Limaverde 1 , Rodrigo Vargas Pêgas 2 , Rafael Damasceno 3 , Chiara Villa 4 , Gustavo Oliveira 3 , Niels Bonde 5, 6 , Maria E. C. Leal Corresp. 1, 5 1 Centro de Ciências, Departamento de Geologia, Universidade Federal do Ceará, Fortaleza, Brazil 2 Department of Geology and Paleontology, Museu Nacional/Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil 3 Departamento de Biologia, Universidade Federal Rural de Pernambuco, Recife, Brazil 4 Department of Forensic Medicine, Copenhagen University, Copenhagen, Denmark 5 Section Biosystematics, Zoological Museum (SNM, Copenhagen University), Copenhagen, Denmark 6 Fur Museum (Museum Saling), Fur, DK-7884, Denmark Corresponding Author: Maria E. C. Leal Email address: [email protected] The Araripe Basin (Northeastern Brazil) has yielded a rich Cretaceous fossil fauna of both vertebrates and invertebrates found mainly in the Crato and Romualdo Formations, of Aptian and Albian ages respectively. Among the vertebrates, the turtles were proved quite diverse, with several specimens retrieved and five valid species described to this date for the Romualdo Fm.
    [Show full text]
  • Post-Cranial Skeleton of Eosphargis Breineri NIELSEN
    On the post-cranial skeleton of Eosphargis breineri NIELSEN. By EIGIL NIELSEN CONTENTS Preface 281 Introduction 282 Eosphargis breineri NIELSEN 283 Material and locality 283 Measurements 283 Skull 284 Vertebral column 284 Carapace 284 Plastron .291 Shoulder girdle 294 Fore-limb 296 Pelvis 303 Hind-limb 307 Remnants of soft tissues 308 Remarks on the relationship of Eosphargis 308 Literature 313 Plates 314 Preface. From the outstanding collector of fossils from the Lower Eocene marine mo clay deposits in Northern Jutland, Mr. M. BREINER JENSEN, leader of the Fur museum, I have received both in 1959 and 1960 for investiga- tion further remnants of turtles from the locality at Knudeklint on the island of Fur, where Mr. BREINEB. JENSEN in 1957 collected the almost complete skull of Eosphargis breineri NIELSEN described by me in 1959. I owe Mr. BREINER JENSEN a sincere thank for the permission to investigate also this new important material, the tedious preparation of which has been carried out in the laboratory of vertebrate paleontology in the Mineralogical and Geological Museum of the University of Copen- hagen mainly by stud. mag. BENTE SOLTAU, who also is responsible for the photographs in this paper. I hereby thank Mrs SOLTAU for her carefull work. Moreover I thank the artists Mrs BETTY ENGHOLM and Mrs RAGNA LARSEN who have drawn the text-figures, as well as stud. mag. SVEND 21 282 EIGIL NIELSEN : On the post-cranial skeleton of Eosphargis breineri NIELSEN E. B.-ALMGREEN, who in various ways has assisted in the finishing the illustrations. To the CARLSBERG FOUNDATION my thanks are due for financial support to the work of preparation and illustration of the material and to the RASK ØRSTED FOUNDATION for financial support to the reproduction of the illustrations.
    [Show full text]
  • A Phylogenomic Analysis of Turtles ⇑ Nicholas G
    Molecular Phylogenetics and Evolution 83 (2015) 250–257 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev A phylogenomic analysis of turtles ⇑ Nicholas G. Crawford a,b,1, James F. Parham c, ,1, Anna B. Sellas a, Brant C. Faircloth d, Travis C. Glenn e, Theodore J. Papenfuss f, James B. Henderson a, Madison H. Hansen a,g, W. Brian Simison a a Center for Comparative Genomics, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA b Department of Genetics, University of Pennsylvania, Philadelphia, PA 19104, USA c John D. Cooper Archaeological and Paleontological Center, Department of Geological Sciences, California State University, Fullerton, CA 92834, USA d Department of Biological Sciences, Louisiana State University, Baton Rouge, LA 70803, USA e Department of Environmental Health Science, University of Georgia, Athens, GA 30602, USA f Museum of Vertebrate Zoology, University of California, Berkeley, CA 94720, USA g Mathematical and Computational Biology Department, Harvey Mudd College, 301 Platt Boulevard, Claremont, CA 9171, USA article info abstract Article history: Molecular analyses of turtle relationships have overturned prevailing morphological hypotheses and Received 11 July 2014 prompted the development of a new taxonomy. Here we provide the first genome-scale analysis of turtle Revised 16 October 2014 phylogeny. We sequenced 2381 ultraconserved element (UCE) loci representing a total of 1,718,154 bp of Accepted 28 October 2014 aligned sequence. Our sampling includes 32 turtle taxa representing all 14 recognized turtle families and Available online 4 November 2014 an additional six outgroups. Maximum likelihood, Bayesian, and species tree methods produce a single resolved phylogeny.
    [Show full text]
  • Basal Turtles Shell Bone Histology Indicates Terrestrial Palaeoecology Of
    Downloaded from rspb.royalsocietypublishing.org on May 22, 2012 Shell bone histology indicates terrestrial palaeoecology of basal turtles Torsten M Scheyer and P.Martin Sander Proc. R. Soc. B 2007 274, 1885-1893 doi: 10.1098/rspb.2007.0499 Supplementary data "Data Supplement" http://rspb.royalsocietypublishing.org/content/suppl/2009/03/13/274.1620.1885.DC1.ht ml References This article cites 26 articles, 5 of which can be accessed free http://rspb.royalsocietypublishing.org/content/274/1620/1885.full.html#ref-list-1 Article cited in: http://rspb.royalsocietypublishing.org/content/274/1620/1885.full.html#related-urls Receive free email alerts when new articles cite this article - sign up in the box at the top Email alerting service right-hand corner of the article or click here To subscribe to Proc. R. Soc. B go to: http://rspb.royalsocietypublishing.org/subscriptions Downloaded from rspb.royalsocietypublishing.org on May 22, 2012 Proc. R. Soc. B (2007) 274, 1885–1893 doi:10.1098/rspb.2007.0499 Published online 22 May 2007 Shell bone histology indicates terrestrial palaeoecology of basal turtles Torsten M. Scheyer*,† and P. Martin Sander Institute of Palaeontology, University of Bonn, Nussallee 8, 53115 Bonn, Germany The palaeoecology of basal turtles from the Late Triassic was classically viewed as being semi-aquatic, similar to the lifestyle of modern snapping turtles. Lately, this view was questioned based on limb bone proportions, and a terrestrial palaeoecology was suggested for the turtle stem. Here, we present independent shell bone microstructural evidence for a terrestrial habitat of the oldest and basal most well- known turtles, i.e.
    [Show full text]
  • Glarichelys Knorri (Gray)-A Cheloniid from Carpathian Menilitic Shales (Poland)
    ACT A P A L A E 0 ~ T 0 LOG I CA P 0 LON IC A Vol. IV I 9 5 9 No . 2 MARIAN MLYN ARSKI GLARICHELYS KNORRI (GRAY) - A CHELONIID FROM CARPATHIAN MENILITIC SHALES (POLAND) Abstract . - The fossil remains here described belonged to a young indiv id ual of Glarichelys knorri (Gray), a sea turtle. They were collected from Carpathian me­ nilitic shales at Winnica near Jaslo. Its systematic position is discussed a nd general comments are made on some fossil and recent sea turtles, on problems concerning their mo rphology, on the taxonomic significance of p halanges in fossil sea turtles, a nd on the presence in cheloniids of foramina praenucha lia. Biological and ecological notes concernin g G. knorri (Gray) are likewise given. INTRODUCTlION The fossil sea turtle remains here described have been collected from an outcrop in the steep bank of the Jasiolka stream, near the Winnica farm, about 10 km to the east of Jaslo (P olish Carpathians). The specimen was found in greyish-brown menilitic shales intercalating the Kr osno sandstone beds, about 30 m above the foot of the men tioned bank. Unfortunately, the geological age of these beds has not, as yet, been definitely established. On their microfauna it is probably Lower Oligocene or Upper Eocene 1. The vertebrate fauna from the Jaslo area has lately attracted the attention of palaeontologists. Abundant and we ll preserved bon y fish remains have been collected there.They belong to several families, mostly to Clupeidae and Gadidae. They are n ow being worked out by A .J erz­ manska (1958) of the Wroclaw University.
    [Show full text]
  • Tetrapod Track Assemblages from Lower Cretaceous Desert Facies In
    Palaeogeography, Palaeoclimatology, Palaeoecology 507 (2018) 1–14 Contents lists available at ScienceDirect Palaeogeography, Palaeoclimatology, Palaeoecology journal homepage: www.elsevier.com/locate/palaeo Tetrapod track assemblages from Lower Cretaceous desert facies in the Ordos Basin, Shaanxi Province, China, and their implications for Mesozoic T paleoecology ⁎ Lida Xinga,b,c, Martin G. Lockleyd, , Yongzhong Tange, Anthony Romiliof, Tao Xue, Xingwen Lie, Yu Tange, Yizhao Lie a State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences, Beijing 100083, China b School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China c State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China d Dinosaur Trackers Research Group, CB 172, University of Colorado at Denver, PO Box 173364, Denver, CO 80217-3364, USA e Shaanxi Geological Survey, Xi'an 710054, Shaanxi, China f School of Biological Sciences, The University of Queensland, Brisbane, Queensland 4072, Australia ARTICLE INFO ABSTRACT Keywords: Tetrapod ichnofaunas are reported from desert, playa lake facies in the Lower Cretaceous Luohe Formation at Ichnofacies Baodaoshili, Shaanxi Province, China, which represent the first Asian example of an ichnnofauna typical of the Brasilichnium Chelichnus Ichnofacies (Brasilichnium sub-ichnofacies) characteristic of desert habitats. The mammaliomorph Sarmientichnus tracks, assigned to Brasilichnium,
    [Show full text]
  • 71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
    ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas.
    [Show full text]
  • Universidad Nacional Del Comahue Centro Regional Universitario Bariloche
    Universidad Nacional del Comahue Centro Regional Universitario Bariloche Título de la Tesis Microanatomía y osteohistología del caparazón de los Testudinata del Mesozoico y Cenozoico de Argentina: Aspectos sistemáticos y paleoecológicos implicados Trabajo de Tesis para optar al Título de Doctor en Biología Tesista: Lic. en Ciencias Biológicas Juan Marcos Jannello Director: Dr. Ignacio A. Cerda Co-director: Dr. Marcelo S. de la Fuente 2018 Tesis Doctoral UNCo J. Marcos Jannello 2018 Resumen Las inusuales estructuras óseas observadas entre los vertebrados, como el cuello largo de la jirafa o el cráneo en forma de T del tiburón martillo, han interesado a los científicos desde hace mucho tiempo. Uno de estos casos es el clado Testudinata el cual representa uno de los grupos más fascinantes y enigmáticos conocidos entre de los amniotas. Su inconfundible plan corporal, que ha persistido desde el Triásico tardío hasta la actualidad, se caracteriza por la presencia del caparazón, el cual encierra a las cinturas, tanto pectoral como pélvica, dentro de la caja torácica desarrollada. Esta estructura les ha permitido a las tortugas adaptarse con éxito a diversos ambientes (por ejemplo, terrestres, acuáticos continentales, marinos costeros e incluso marinos pelágicos). Su capacidad para habitar diferentes nichos ecológicos, su importante diversidad taxonómica y su plan corporal particular hacen de los Testudinata un modelo de estudio muy atrayente dentro de los vertebrados. Una disciplina que ha demostrado ser una herramienta muy importante para abordar varios temas relacionados al caparazón de las tortugas, es la paleohistología. Esta disciplina se ha involucrado en temas diversos tales como el origen del caparazón, el origen del desarrollo y mantenimiento de la ornamentación, la paleoecología y la sistemática.
    [Show full text]
  • Osseous Growth and Skeletochronology
    Comparative Ontogenetic 2 and Phylogenetic Aspects of Chelonian Chondro- Osseous Growth and Skeletochronology Melissa L. Snover and Anders G.J. Rhodin CONTENTS 2.1 Introduction ........................................................................................................................... 17 2.2 Skeletochronology in Turtles ................................................................................................ 18 2.2.1 Background ................................................................................................................ 18 2.2.1.1 Validating Annual Deposition of LAGs .......................................................20 2.2.1.2 Resorption of LAGs .....................................................................................20 2.2.1.3 Skeletochronology and Growth Lines on Scutes ......................................... 21 2.2.2 Application of Skeletochronology to Turtles ............................................................. 21 2.2.2.1 Freshwater Turtles ........................................................................................ 21 2.2.2.2 Terrestrial Turtles ......................................................................................... 21 2.2.2.3 Marine Turtles .............................................................................................. 21 2.3 Comparative Chondro-Osseous Development in Turtles......................................................22 2.3.1 Implications for Phylogeny ........................................................................................32
    [Show full text]
  • Presence of the Jehol Biota Turtle Ordosemys Liaoxiensis in the Early Cretaceous Hengtongshan Formation of Southern Jilin Province, China
    Foss. Rec., 22, 57–64, 2019 https://doi.org/10.5194/fr-22-57-2019 © Author(s) 2019. This work is distributed under the Creative Commons Attribution 4.0 License. Presence of the Jehol Biota turtle Ordosemys liaoxiensis in the Early Cretaceous Hengtongshan Formation of southern Jilin Province, China Chang-Fu Zhou1, Wen-Hao Wu2,3, and Márton Rabi4,5 1College of Earth Science and Engineering, Shandong University of Science and Technology, Qingdao 266590, Shandong, China 2Key Laboratory for Evolution of Past Life and Environment in Northeast Asia, Ministry of Education, Changchun 130026, Jilin, China 3Research Center of Palaeontology and Stratigraphy, Jilin University, Changchun 130056, Jilin, China 4Central Natural Science Collections, Martin Luther University Halle-Wittenberg, Halle (Saale), Germany 5Department of Geosciences, University of Tübingen, Tübingen, Germany Correspondence: Chang-Fu Zhou ([email protected]) and Wen-Hao Wu ([email protected]) Received: 25 May 2019 – Revised: 15 August 2019 – Accepted: 19 August 2019 – Published: 12 September 2019 Abstract. Recently, a vertebrate assemblage of the Jehol ence of the Eosestheria (crustacean)–Ephemeropsis trisetalis Biota has been reported from the Early Cretaceous Heng- (mayfly insect)–Lycoptera (fish) community (T. Shao et al., tongshan Formation of Xingling Town, Meihekou City, Jilin 2017). Freshwater sinemydid turtles are among the most Province, China. It is dominated by the fishes Lycoptera and common tetrapod fossils of the Jehol Biota, which makes Sinamia and the sinemydid turtle Ordosemys. Here, we de- their biostratigraphic potential of interest (e.g., Ji, 1995; Tong scribe the turtle specimens and referral to Ordosemys liaox- et al., 2004; Zhou, 2010a, b; Zhou and Rabi, 2015; S.
    [Show full text]