A TAXONOMIC LISTING of BENTHIC MACRO- and MEGAINVERTEBRATES from Infaunal & Epifaunal Monitoring and Research Programs in the Southern California Bight

Total Page:16

File Type:pdf, Size:1020Kb

A TAXONOMIC LISTING of BENTHIC MACRO- and MEGAINVERTEBRATES from Infaunal & Epifaunal Monitoring and Research Programs in the Southern California Bight A TAXONOMIC LISTING OF BENTHIC MACRO- and MEGAINVERTEBRATES from Infaunal & Epifaunal Monitoring and Research Programs in the Southern California Bight EDITION 11 1 July 2016 Prepared by The Southern California Association of Marine Invertebrate Taxonomists Natural History Museum of Los Angeles County Research & Collections 900 Exposition Boulevard Los Angeles, CA 90007 The editors of this list intend that it undergo regular updates. SCAMIT hopes to maintain the list as a doc- ument useful to those involved with monitoring programs within the Southern California Bight. To this end we solicit the users' assistance. Please forward any comments, corrections, or suggested additions you may have to them: Donald B. Cadien or Lawrence L. Lovell [email protected] [email protected] Marine Biology Laboratory Marine Biology Laboratory County Sanitation Districts County Sanitation Districts of Los Angeles County of Los Angeles County 24501 S. Figueroa St. 24501 S. Figueroa St. Carson, CA 90745 Carson, CA 90745 This document is available to members for downloading in Portable Document Format (pdf) at the SCAMIT web site (www.scamit.org). To view or download the document you must have the Adobe Acr- obat Reader which allows you to view or print pdf files. The reader is distributed by Adobe free of charge and may be downloaded from their web site (www.adobe.com). Copyright © 2016 by the Southern California Association of Marine Invertebrate Taxonomists This material may not be duplicated and resold. Permission to copy or use is hereby granted subject to inclusion of this copyright notice. Edition 11 i 1 July 2016 Southern California Association of Marine Invertebrate Taxonomists A TAXONOMIC LISTING OF BENTHIC MACRO- and MEGAINVERTEBRATES from Infaunal and Epifaunal Monitoring and Research Programs in the Southern California Bight Prepared by The Southern California Association of Marine Invertebrate Taxonomists Donald B. Cadien and Lawrence L. Lovell, editors EDITION 11 Taxonomic standardization, assuring that the same name is applied to an organism by different taxono- mists each time that organism is encountered, is an important quality control concern in any biological monitoring program. Compatibility of taxonomic data generated by agencies and organizations contribut- ing to regional monitoring programs is essential if the full value of these surveys is to be realized. There are two obstacles to achieving the goal of standardization: the complexity and fluidity of invertebrate tax- onomy; and the differences in expertise, experience, and opinion of taxonomists involved in monitoring within the region. While these obstacles hinder us from achieving uniformity, a great deal can be accom- plished in its pursuit. Central to that effort is regional cooperation and communication among the taxon- omists responsible for the surveys. The Southern California Association of Marine Invertebrate Taxonomists (SCAMIT) was formed in 1982 to promote the study of invertebrate taxonomy and develop a standardized taxonomy of marine inverte- brates within the region. Its membership includes most of the taxonomists responsible for conducting in- faunal and epifaunal monitoring programs in Southern California coastal waters. SCAMIT's activities include monthly meetings, workshops, and communication of taxonomic information to its membership through a newsletter and website (www.scamit.org). This list of invertebrates from benthic habitats of the Southern California Bight (SCB) is a contribution of SCAMIT toward the development of regionally coordinated monitoring within the Bight. This edition updates Edition 10 of July 2015, and continues SCAMIT's efforts to compile and maintain a list of species for which taxonomy is believed standard within the region. That is, a list of species names that are uni- formly and consistently applied to organisms in research and monitoring programs within the region. Like earlier editions, this is a conservative document constrained by both the information base from which it is derived and by the criteria for inclusion of taxa on the list. Caution is required in the use of this or any other “master species” list. Taxonomic listings do not in themselves constitute standardization of taxonomic usage. They are only tools that contribute to standard- ization, both as a product and in the process of their development and maintenance. Those responsible for developing regional monitoring should not attempt to impose on contributing surveys an artificially rigid Edition 11 1! 1 July 2016 taxonomy based upon a static list of standard names. Such an approach would inevitably lead to poorer quality survey data. Some differences in taxonomy and nomenclatural usage will continue to appear in data sets originating from contributing surveys. Most of these will reflect either change in the species en- countered or new revisionary work. Some will represent non-standard usage despite efforts such as this listing and other activities of SCAMIT. Regional data-management systems must allow both growth and emendation to their species lists to conform to current usage. Ranasinghe et al. (2003b) evaluated the suc- cess of quality assurance taxonomic standardization in the second regional monitoring survey Bight ‘98 (B’98) and provided some idea of both the value of the contributions to standardization made to date and the magnitude of the unresolved issues. Edition 7 was the first edition available only on-line. It was available in Portable Document Format (pdf), which is fully searchable. This renders the Index provided in previous editions obsolete, and none is pro- vided here. Edition 11 is another step in the evolution to a fully interactive on-line list in the future, where comments and questions regarding the list contents can be made by users in real-time. Future lists will be linked to other resources on the internet, providing additional information on the species covered. We have not yet accomplished this, but look forward to doing so in the near future. COVERAGE OF EDITION 11 This document provides a list of macroinvertebrates and megainvertebrates from both soft and hard bot- tom habitats of the SCB. Holoplanktonic species are not included in the list. For the purposes of the list, macroinvertebrates are defined as those organisms retained upon a 0.3 mm or larger mesh screen. Epiben- thic megainvertebrates are defined as benthic organisms captured in otter trawls and having a body di- mension greater than 1 cm. The list draws from sam- pling programs that have been in existence since as Contributing Programs early as 1970 (see table of contributors). Over that LARGE DISCHARGE MONITORING PROGRAMS period, sampling frequencies and station locations City of Los Angeles, Bureau of Sanitation, Envi- have varied but gear types and methods have been ronmental Monitoring Division County Sanitation Districts of Los Angeles County stable. The list reflects organisms collected from Orange County Sanitation District thousands of samples within the habitats covered. City of San Diego, Public Utilities Department Most records are from soft substrate on the inner to SMALL DISCHARGE MONITORING PROGRAMS mid shelf within the SCB, but increasingly informa- Goleta Sanitary District City of Santa Barbara tion on other habitats from the intertidal to the near- Carpenteria Sanitary District shore basins is also represented. Records in the pub- City of Oxnard lished literature are not sufficient for inclusion, only South East Regional Reclamation Authority actual examined specimens form the basis for this City of Oceanside Encina Water Pollution Control Facility listing. There are 3227 species-level taxa on the list San Elijo Joint Powers Authority representing 17 phyla, 39 classes, 132 orders, and City of Avalon 706 families. International Treatment Plant MISCELLANEOUS STUDIES Geographic coverage is from Point Conception, Cal- Terminal Island Treatment Plant Outfall Study ifornia to Bahía Todos Santos, Baja California Sur, Tijuana Ocean Engineering Study Mexico in intertidal to 1000m depths. As in previous LACSD Reference Survey San Diego Sediment Mapping Study editions, the information on which the list is based San Diego Deep Benthic Pilot Study comes from survey data collected in monitoring pro- REGIONAL SURVEYS grams of publicly owned wastewater treatment Southern California Coastal Water Research works (POTWs). Coverage within the Bight is biased Project Reference Surveys 1994 Southern California Bight Pilot Project by the distribution of sampling efforts associated (SCBPP) with these programs. The largest of these are cen- Southern California Bight 1998, 2003, 2008, and tered on the metropolitan areas of Los Angeles/Or- 2013 Regional Monitoring Projects ange County and San Diego. The fauna from shelf Edition 11 2! 1 July 2016 and upper slope depths in these areas is well represented. North of the Los Angeles region, monitoring programs are fewer in number and smaller in scale. Consequently, the fauna of the northern Bight is un- der-represented. Records from Regional Monitoring Surveys in 1994, 1998, 2003, 2008 and 2013 (Allen et al. 1998, 2002, 2007, 2011; Bergen et al. 1998; Ranasinghe et al. 2003a, 2007, 2012) improved cover- age in this northern region as well as between POTW monitoring areas. Coverage is poorer south of the US/Mexico border where the only records come from monitoring of the South Bay Ocean Outfall (south to Punta Banderas) and the Mexican component of the Bight ’98 Regional Survey (south to Bahía Todos Santos). Records from estuaries,
Recommended publications
  • Bryozoan Studies 2019
    BRYOZOAN STUDIES 2019 Edited by Patrick Wyse Jackson & Kamil Zágoršek Czech Geological Survey 1 BRYOZOAN STUDIES 2019 2 Dedication This volume is dedicated with deep gratitude to Paul Taylor. Throughout his career Paul has worked at the Natural History Museum, London which he joined soon after completing post-doctoral studies in Swansea which in turn followed his completion of a PhD in Durham. Paul’s research interests are polymatic within the sphere of bryozoology – he has studied fossil bryozoans from all of the geological periods, and modern bryozoans from all oceanic basins. His interests include taxonomy, biodiversity, skeletal structure, ecology, evolution, history to name a few subject areas; in fact there are probably none in bryozoology that have not been the subject of his many publications. His office in the Natural History Museum quickly became a magnet for visiting bryozoological colleagues whom he always welcomed: he has always been highly encouraging of the research efforts of others, quick to collaborate, and generous with advice and information. A long-standing member of the International Bryozoology Association, Paul presided over the conference held in Boone in 2007. 3 BRYOZOAN STUDIES 2019 Contents Kamil Zágoršek and Patrick N. Wyse Jackson Foreword ...................................................................................................................................................... 6 Caroline J. Buttler and Paul D. Taylor Review of symbioses between bryozoans and primary and secondary occupants of gastropod
    [Show full text]
  • Federal Register / Vol. 62, No. 182 / Friday, September 19, 1997 / Proposed Rules
    49398 Federal Register / Vol. 62, No. 182 / Friday, September 19, 1997 / Proposed Rules DEPARTMENT OF THE INTERIOR time. The information on candidate taxa Massachusetts, New Hampshire, New will be revised and updated continually Jersey, New York, Pennsylvania, Rhode Fish and Wildlife Service by the Regional Offices identified as Island, Vermont, Virginia, and West having lead responsibility for the Virginia. 50 CFR Part 17 particular taxa. The Service anticipates Regional Director (TE), U.S. Fish and Wildlife Service, 300 Westgate Center Endangered and Threatened Wildlife publishing annually an update of the Drive, Hadley, Massachusetts 01035± and Plants; Review of Plant and candidate notice of review, annual notice of findings on recycled petitions, 9589 (413/253±8615). Animal Taxa That Are Candidates or Region 6. Colorado, Kansas, Montana, Proposed for Listing as Endangered or and annual description of progress on listing actions. Nebraska, North Dakota, South Dakota, Threatened, Annual Notice of Findings Utah, and Wyoming. on Recycled Petitions, and Annual ADDRESSES: Interested persons or organizations should submit comments Regional Director (TE), U.S. Fish and Description of Progress on Listing Wildlife Service, P.O. Box 25486, Actions regarding a particular taxon to the Regional Director of the Region Denver Federal Center, Denver, AGENCY: Fish and Wildlife Service, identified as having the lead Colorado 80225±0486 (303/236±7398). Interior. responsibility for that taxon. Comments Region 7. Alaska. Regional Director (TE), U.S. Fish and ACTION: Notice of review. of a more general nature may be Wildlife Service, 1011 East Tudor Road, submitted to the Chief, Division of Anchorage, Alaska 99503±6199 (907/ SUMMARY: In this document, the Fish Endangered Species, U.S.
    [Show full text]
  • Annelida, Serpulidae
    Graellsia, 72(2): e053 julio-diciembre 2016 ISSN-L: 0367-5041 http://dx.doi.org/10.3989/graellsia.2016.v72.120 SERPÚLIDOS (ANNELIDA, SERPULIDAE) COLECTADOS EN LA CAMPAÑA OCEANOGRÁFICA “FAUNA II” Y CATÁLOGO ACTUALIZADO DE ESPECIES ÍBERO-BALEARES DE LA FAMILIA SERPULIDAE Jesús Alcázar* & Guillermo San Martín Departamento de Biología (Zoología), Facultad de Ciencias, Universidad Autónoma de Madrid, calle Darwin, 2, Canto Blanco, 28049 Madrid, España. *Dirección para la correspondencia: [email protected] RESUMEN Se presentan los resultados de la identificación del material de la familia Serpulidae (Polychaeta) recolectado en la campaña oceanográfica Fauna II, así como la revisión de citas de presencia íbero-balear desde el catálogo de poliquetos más reciente (Ariño, 1987). Se identificaron 16 especies pertenecientes a 10 géneros, además de la primera cita íbero-balear de una quimera bioperculada (Ten Hove & Ben-Eliahu, 2005) de la especie Hydroides norvegicus Gunnerus, 1768. En cuanto a la revisión del catálogo se mencionan 65 especies, actuali- zando el nombre de 20 de ellas y añadiendo cinco especies ausentes en el catálogo de Ariño (1987): Hydroides stoichadon Zibrowius, 1971, Laeospira corallinae (de Silva & Knight-Jones, 1962), Serpula cavernicola Fassari & Mòllica, 1991, Spirobranchus lima (Grube, 1862) y Spirorbis inornatus L’Hardy & Quièvreux, 1962. Se cita por primera vez Vermiliopsis monodiscus Zibrowius, 1968 en el Atlántico ibérico y a partir de la bibliografía consultada, se muestra la expansión en la distribución íbero-balear
    [Show full text]
  • Polychaete Worms Definitions and Keys to the Orders, Families and Genera
    THE POLYCHAETE WORMS DEFINITIONS AND KEYS TO THE ORDERS, FAMILIES AND GENERA THE POLYCHAETE WORMS Definitions and Keys to the Orders, Families and Genera By Kristian Fauchald NATURAL HISTORY MUSEUM OF LOS ANGELES COUNTY In Conjunction With THE ALLAN HANCOCK FOUNDATION UNIVERSITY OF SOUTHERN CALIFORNIA Science Series 28 February 3, 1977 TABLE OF CONTENTS PREFACE vii ACKNOWLEDGMENTS ix INTRODUCTION 1 CHARACTERS USED TO DEFINE HIGHER TAXA 2 CLASSIFICATION OF POLYCHAETES 7 ORDERS OF POLYCHAETES 9 KEY TO FAMILIES 9 ORDER ORBINIIDA 14 ORDER CTENODRILIDA 19 ORDER PSAMMODRILIDA 20 ORDER COSSURIDA 21 ORDER SPIONIDA 21 ORDER CAPITELLIDA 31 ORDER OPHELIIDA 41 ORDER PHYLLODOCIDA 45 ORDER AMPHINOMIDA 100 ORDER SPINTHERIDA 103 ORDER EUNICIDA 104 ORDER STERNASPIDA 114 ORDER OWENIIDA 114 ORDER FLABELLIGERIDA 115 ORDER FAUVELIOPSIDA 117 ORDER TEREBELLIDA 118 ORDER SABELLIDA 135 FIVE "ARCHIANNELIDAN" FAMILIES 152 GLOSSARY 156 LITERATURE CITED 161 INDEX 180 Preface THE STUDY of polychaetes used to be a leisurely I apologize to my fellow polychaete workers for occupation, practised calmly and slowly, and introducing a complex superstructure in a group which the presence of these worms hardly ever pene- so far has been remarkably innocent of such frills. A trated the consciousness of any but the small group great number of very sound partial schemes have been of invertebrate zoologists and phylogenetlcists inter- suggested from time to time. These have been only ested in annulated creatures. This is hardly the case partially considered. The discussion is complex enough any longer. without the inclusion of speculations as to how each Studies of marine benthos have demonstrated that author would have completed his or her scheme, pro- these animals may be wholly dominant both in num- vided that he or she had had the evidence and inclina- bers of species and in numbers of specimens.
    [Show full text]
  • Oxygen, Ecology, and the Cambrian Radiation of Animals
    Oxygen, Ecology, and the Cambrian Radiation of Animals The Harvard community has made this article openly available. Please share how this access benefits you. Your story matters Citation Sperling, Erik A., Christina A. Frieder, Akkur V. Raman, Peter R. Girguis, Lisa A. Levin, and Andrew H. Knoll. 2013. Oxygen, Ecology, and the Cambrian Radiation of Animals. Proceedings of the National Academy of Sciences 110, no. 33: 13446–13451. Published Version doi:10.1073/pnas.1312778110 Citable link http://nrs.harvard.edu/urn-3:HUL.InstRepos:12336338 Terms of Use This article was downloaded from Harvard University’s DASH repository, and is made available under the terms and conditions applicable to Other Posted Material, as set forth at http:// nrs.harvard.edu/urn-3:HUL.InstRepos:dash.current.terms-of- use#LAA Oxygen, ecology, and the Cambrian radiation of animals Erik A. Sperlinga,1, Christina A. Friederb, Akkur V. Ramanc, Peter R. Girguisd, Lisa A. Levinb, a,d, 2 Andrew H. Knoll Affiliations: a Department of Earth and Planetary Sciences, Harvard University, Cambridge, MA, 02138 b Scripps Institution of Oceanography, University of California San Diego, La Jolla, CA, 92093- 0218 c Marine Biological Laboratory, Department of Zoology, Andhra University, Waltair, Visakhapatnam – 530003 d Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, MA, 02138 1 Correspondence to: [email protected] 2 Correspondence to: [email protected] PHYSICAL SCIENCES: Earth, Atmospheric and Planetary Sciences BIOLOGICAL SCIENCES: Evolution Abstract: 154 words Main Text: 2,746 words Number of Figures: 2 Number of Tables: 1 Running Title: Oxygen, ecology, and the Cambrian radiation Keywords: oxygen, ecology, predation, Cambrian radiation The Proterozoic-Cambrian transition records the appearance of essentially all animal body plans (phyla), yet to date no single hypothesis adequately explains both the timing of the event and the evident increase in diversity and disparity.
    [Show full text]
  • TREBALLS 1 DEL MUSEU DE ZOOLOGIA Illustrated Keys for The,Classification of Mediterranean Bryozoa
    / AJUNTAMENT DE BARCELONA TREBALLS 1 DEL MUSEU DE ZOOLOGIA Illustrated keys for the,classification of Mediterranean Bryozoa M. Zabala & P. Maluquer 1 BARCELONA 1988 NÚMERO 4 Drawing of the cover: Scrupocellaria reptans (Linnaeus), part of a branch with bran- ched scutum, ovicells, frontal avicularia and lateral vibracula. Treb. Mus. 2001. Barcelona. 4. 1988 Illustrated keys for the classification of Mediterranean Bryozoa Consell de Redacció: O. Escola, R. Nos, A. Ornedes, C. Prats i F. Uribe. Assessor científic: P. Hayward M. ZABALA, Dcpt. de Ecologia, Fac. de Biologia, Univcrsitat de Barcelona, Diagonal, 645 08028 Barcelona. P. MALUQUER, Dept. de Biologia Animal, Fac. de Biologia, Lniversitat de Barcelona, Diagonal, 645 08028 Barcelona. Edita: Museu de Zoologia, Ajuntament de Barcelona Parc de la Ciutadclla, Ap. de Correus 593,08003 -Barcelona O 1987, Museu de Zoologia, Ajuntament de Barcelona ISBN: 84-7609-240-7 Depósito legal: B. 28.708-1988 Exp. ~0058-8k'-Impremta Municipal Composición y fotolitos: Romargraf, S.A. FOREWORD Bryozoansare predominantly marine, invertebrate animals whose curious and often attractive forms have long excited the interest of naturalists. In past times they were regarded as plants, and the plant- !ike appearance of some species was later formalized in the term "zoophyte", which also embraced the hydroids and a few other enigmatic animal groups. As "corallines" they were considered to be close to the Cnidaria, while "moss animals" neatly described the appearance of a feeding colony. Establishing their animal nature did not resolve the question of systematic affinity. It is only comparatively recently that Bryozoa have been accepted as a phylum in their own right, although an early view of them as for- ming a single phylogenetic unit, the Lophophorata, with the sessile, filter-feeding brachiopods and pho- ronids, still persists.
    [Show full text]
  • CNIDARIA Corals, Medusae, Hydroids, Myxozoans
    FOUR Phylum CNIDARIA corals, medusae, hydroids, myxozoans STEPHEN D. CAIRNS, LISA-ANN GERSHWIN, FRED J. BROOK, PHILIP PUGH, ELLIOT W. Dawson, OscaR OcaÑA V., WILLEM VERvooRT, GARY WILLIAMS, JEANETTE E. Watson, DENNIS M. OPREsko, PETER SCHUCHERT, P. MICHAEL HINE, DENNIS P. GORDON, HAMISH J. CAMPBELL, ANTHONY J. WRIGHT, JUAN A. SÁNCHEZ, DAPHNE G. FAUTIN his ancient phylum of mostly marine organisms is best known for its contribution to geomorphological features, forming thousands of square Tkilometres of coral reefs in warm tropical waters. Their fossil remains contribute to some limestones. Cnidarians are also significant components of the plankton, where large medusae – popularly called jellyfish – and colonial forms like Portuguese man-of-war and stringy siphonophores prey on other organisms including small fish. Some of these species are justly feared by humans for their stings, which in some cases can be fatal. Certainly, most New Zealanders will have encountered cnidarians when rambling along beaches and fossicking in rock pools where sea anemones and diminutive bushy hydroids abound. In New Zealand’s fiords and in deeper water on seamounts, black corals and branching gorgonians can form veritable trees five metres high or more. In contrast, inland inhabitants of continental landmasses who have never, or rarely, seen an ocean or visited a seashore can hardly be impressed with the Cnidaria as a phylum – freshwater cnidarians are relatively few, restricted to tiny hydras, the branching hydroid Cordylophora, and rare medusae. Worldwide, there are about 10,000 described species, with perhaps half as many again undescribed. All cnidarians have nettle cells known as nematocysts (or cnidae – from the Greek, knide, a nettle), extraordinarily complex structures that are effectively invaginated coiled tubes within a cell.
    [Show full text]
  • An Annotated Checklist of the Marine Macroinvertebrates of Alaska David T
    NOAA Professional Paper NMFS 19 An annotated checklist of the marine macroinvertebrates of Alaska David T. Drumm • Katherine P. Maslenikov Robert Van Syoc • James W. Orr • Robert R. Lauth Duane E. Stevenson • Theodore W. Pietsch November 2016 U.S. Department of Commerce NOAA Professional Penny Pritzker Secretary of Commerce National Oceanic Papers NMFS and Atmospheric Administration Kathryn D. Sullivan Scientific Editor* Administrator Richard Langton National Marine National Marine Fisheries Service Fisheries Service Northeast Fisheries Science Center Maine Field Station Eileen Sobeck 17 Godfrey Drive, Suite 1 Assistant Administrator Orono, Maine 04473 for Fisheries Associate Editor Kathryn Dennis National Marine Fisheries Service Office of Science and Technology Economics and Social Analysis Division 1845 Wasp Blvd., Bldg. 178 Honolulu, Hawaii 96818 Managing Editor Shelley Arenas National Marine Fisheries Service Scientific Publications Office 7600 Sand Point Way NE Seattle, Washington 98115 Editorial Committee Ann C. Matarese National Marine Fisheries Service James W. Orr National Marine Fisheries Service The NOAA Professional Paper NMFS (ISSN 1931-4590) series is pub- lished by the Scientific Publications Of- *Bruce Mundy (PIFSC) was Scientific Editor during the fice, National Marine Fisheries Service, scientific editing and preparation of this report. NOAA, 7600 Sand Point Way NE, Seattle, WA 98115. The Secretary of Commerce has The NOAA Professional Paper NMFS series carries peer-reviewed, lengthy original determined that the publication of research reports, taxonomic keys, species synopses, flora and fauna studies, and data- this series is necessary in the transac- intensive reports on investigations in fishery science, engineering, and economics. tion of the public business required by law of this Department.
    [Show full text]
  • Of Polychaetes (Annelida: Polychaeta) from the Atlantic and Mediterranean Coasts of the Iberian Peninsula: an Annotated Checklist E
    López and Richter Helgol Mar Res (2017) 71:19 DOI 10.1186/s10152-017-0499-6 Helgoland Marine Research ORIGINAL ARTICLE Open Access Non‑indigenous species (NIS) of polychaetes (Annelida: Polychaeta) from the Atlantic and Mediterranean coasts of the Iberian Peninsula: an annotated checklist E. López* and A. Richter Abstract This study provides an updated catalogue of non-indigenous species (NIS) of polychaetes reported from the conti- nental coasts of the Iberian Peninsula based on the available literature. A list of 23 introduced species were regarded as established and other 11 were reported as casual, with 11 established and nine casual NIS in the Atlantic coast of the studied area and 14 established species and seven casual ones in the Mediterranean side. The most frequent way of transport was shipping (ballast water or hull fouling), which according to literature likely accounted for the intro- ductions of 14 established species and for the presence of another casual one. To a much lesser extent aquaculture (three established and two casual species) and bait importation (one established species) were also recorded, but for a large number of species the translocation pathway was unknown. About 25% of the reported NIS originated in the Warm Western Atlantic region, followed by the Tropical Indo West-Pacifc region (18%) and the Warm Eastern Atlantic (12%). In the Mediterranean coast of the Iberian Peninsula, nearly all the reported NIS originated from warm or tropi- cal regions, but less than half of the species recorded from the Atlantic side were native of these areas. The efects of these introductions in native marine fauna are largely unknown, except for one species (Ficopomatus enigmaticus) which was reported to cause serious environmental impacts.
    [Show full text]
  • Bathyal and Abyssal Polychaetes (Annelids) from the Central Coast of Oregon
    AN ABSTRACT OF THE THESIS OF DANIL RAY HANCOCK for the MASTER OF SCIENCE (Name) (Degree) in OCEANOGRAPHY presented on (Major) (Date) Title: BATHYAL AND ABYSSAL POLYCIiAETES (ANNELIDS) FROM THE CENTRAL COAST OF OREGON Abstract approved Redacted for Privacy Andtew G. Ca4ey, Jr. Polychaete annelids from 48 benthic samples containing over 2000 specimens were identified.Samples were taken with either an anchor dredge or an anchor-box dredge from a 15 station transect (44° 39. l'N) that ranges from 800 to 2900 meters in depth.Sediment subsamples were collected and analyzed for organic carbon and sedi- ment particle size using standard techniques.Temperature and oxygen of the water near the bottom were taken with a modified Smith- McIntyre grab; however, these measurements were not taken simultaneously with the dredged biological samples. The results indicated that at least 115 species in 53 families of the class Polychaeta were represented in this transect line.This study found an absence of the families Serpulidae and Syllidae and a reduction of the number of speciesin the families Nereidae, Cirratulidae and Capitellidae.Only five genera had not previously been reported from the deep sea.The depth distribution of the polychaetous annelids recovered in this study, coupled with limited physical data, suggest that five faunal regions can be distinguished. Nine new forms of polychaeteous annelids are tentativelydescribed, and others are anticipated in future collections.Suggestions for future studies are also indicated. Bathyal and Abyssal Polychaetes
    [Show full text]
  • Northern Adriatic Bryozoa from the Vicinity of Rovinj, Croatia
    NORTHERN ADRIATIC BRYOZOA FROM THE VICINITY OF ROVINJ, CROATIA PETER J. HAYWARD School of Biological Sciences, University of Wales Singleton Park, Swansea SA2 8PP, United Kingdom Honorary Research Fellow, Department of Zoology The Natural History Museum, London SW7 5BD, UK FRANK K. MCKINNEY Research Associate, Division of Paleontology American Museum of Natural History Professor Emeritus, Department of Geology Appalachian State University, Boone, NC 28608 BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY CENTRAL PARK WEST AT 79TH STREET, NEW YORK, NY 10024 Number 270, 139 pp., 63 ®gures, 1 table Issued June 24, 2002 Copyright q American Museum of Natural History 2002 ISSN 0003-0090 2 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY NO. 270 CONTENTS Abstract ....................................................................... 5 Introduction .................................................................... 5 Materials and Methods .......................................................... 7 Systematic Accounts ........................................................... 10 Order Ctenostomata ............................................................ 10 Nolella dilatata (Hincks, 1860) ................................................ 10 Walkeria tuberosa (Heller, 1867) .............................................. 10 Bowerbankia spp. ............................................................ 11 Amathia pruvoti Calvet, 1911 ................................................. 12 Amathia vidovici (Heller, 1867) ..............................................
    [Show full text]
  • 1 Class Stenolaemata Order Cyclostomata Philip E
    1 Class Stenolaemata Order Cyclostomata Philip E. Bock, Paul D. Taylor, Peter J. Hayward and Dennis P. Gordon 1.1 Definition and general description may be encrusting or erect and branching or foli- Stenolaemata is the most ancient bryozoan class, ose. They are typically dense, opaque white in with a fossil record beginning in the earliest Ordo- colour, occasionally flushed pink or purple, and vician, ~500 million years ago (Taylor and Ernst the calcification can appear speckled because of the 2004; Ma et al. 2015). Seven orders are recognised presence of numerous tissue-plugged pseudopores. currently (Taylor and Waeschenbach 2015), of In the Crisiidae, exemplifying the erect, branching which only Cyclostomata survives and includes all Articulata, the zooids are arranged in narrow rows living stenolaemate species. Globally, the order with openings on only one side of the slender, flex- Cyclostomata includes some 543 species assigned ible colony of branches linked by cuticular joints to 98 genera and 23 families (Bock and Gordon (nodes). Erect colonies of species of Tubuliporina, 2013). The group comprises, on average, ~11% of Cancellata and Cerioporina are unjointed (with the the species in any Recent bryozoan fauna (range single exception of the tubuliporine genus Crisuli- 0–24%, Banta 1991) and only rarely dominates in pora), gracile to robust, and have zooids arranged terms of numbers of colonies or biomass. evenly, in clusters or in ordered transverse rows. Stenolaemates are commonly termed ‘tubular Many species of Tubuliporina have encrusting col- bryozoans’, in reference to their elongate, slender, onies, occasionally taking the form of simple, uni- usually cylindrical zooids.
    [Show full text]