I I
BULLETIN DE L'INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE, ENTOMOLOGIE, 59: 5-22, 1989 BULLETIN VAN HET KONINKLIJK BELGISCH INSTITUUT VOOR NATUURWETENSCHAPPEN, ENTOMOLOGIE, 59: 5-22, 1989
Results of the Belgian 1988-expedition to the Galapagos islands Araneae.
by L. BAERT, J.-P. MAELFAIT and K. DESENDER
Contribution N° 442 of the Charles Darwin Research Foun For each of the discussed species we mention : dation. - the literature references if the species was not mentioned in both former publications; - the material sampled in 1988; Abstract - the other examined material. This section starts with the reference to the former publications (BAERT & MAEL In this paper the detailed araneological results are given of the Belgian FAIT, 1986 and BAERT, MAELFAIT & DESENDER, 1989). 1988-expedition to the Galapagos archipelago. For each of the 51 caught species, belonging to 26 families, the localities where they were found are The first author had also the opportunity to examine the given. spiders collected by Prof. A. FRANZ in 1975; Key-words : Galapagos - Araneae - Distribution. - in both former sections we give the number of males, females and juveniles collected or examined, together with the month (roman ciphers) of capture; Resume - the distribution of the species over the archipelago. The islands marked with an asterisk were not previously Ce travail donne les resultats araneologiques detailles de !'expedition beige mentioned for the species; effectuee en 1988 sur l'archipel des Galapagos. Pour chaque des 51 especes - the zoogeographic affinities are given for the species captun!es, appartenant a 26 families, les localites de capture sont donnees. Mots-clefs : Galapagos - Araneae - Distribution. not mentioned in both former publications.
The localities sampled are given in Figure 1. Introduction Some families are however still incomplete. For the Gna In 1988, the authors spent two months (February 10- April phosidae, the genus Camillina is omitted. This genus is 5) on the Galapagos Islands in the scope of their Inverte subjected to a more detailed study. The same holds for the brate survey of the archipelago, started in 1982. In a first Lycosidae. Due to a lack of good recent descriptions and/ publication (BAERT & MAELFAIT, 1986) the state of know or revisional works a large deal of the spider material ledge of the araneological fauna of the archipelago was collected during the three surveys (1982, 1986 and 1988) given based upon the material sampled by the first two still can not be identified with certainty. This is the case authors during their first stay in 1982, the available litera for the Oonopidae, some Pholcidae, Salticidae and Aranei ture data and the collections of various Institutions. dae. For the Linyphiidae six species new to science were In 1986, the authors visited the islands for the second time found in the course of the successive expeditions ( 1982, and succeeded in sampling the islands Santa Cruz, Pinzon, 1986 and 1988) and are described in a separate paper Rabida, Santiago, Pinta, Isabela (Cerro Azul, Sierra Negra (BAERT, in press). and Volcan Alcedo) and San Cristobal. The results of this expedition are given in BAERT, MAELFAIT & DESENDER (1989), together with a provisional checklist of the spiders Localities sampled in 1988 of the archipelago. In the course of our 1988-expedition, the remaining major ISLA SANTA CRUZ islands (Floreana, Espanola, Marchena, Genovesa, Daphne Mayor, Fernandina and the top of Santa Cruz) and vol Locality 1 : Puerto Ayora village, facade of building. canoes (Volcan Wolf and Darwin of Isla Isabela) were sampled. Locality 2 : The Charles Darwin Research Station (CDRS) The present publication, giving the results of this last expe located in the dry arid zone (Bahfa Academica); in the dition, is built on the same canvas as the former ones. buildings of the Station. 0\ -Y/i I I r c:l
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Fig. l : Localities sampled. I I
Araneae of the Galapagos 7
Locality 3 : Nearby the seismological station of the CDRS, tum fagara and Psychotria rufipes. Herb layer (60 % site called "Barranco". Spiders were caught under stones coverage) : the fern Blechnum occidentale and the gras and on tree cacti (Opuntia echios). Deciduous steppe forest ses Borreria /aevis, Paspalum conjugatum and Sida sa/ with a dominance of Opuntia echios and Croton scou/eri. viifolia. Lac. 14: Los Gemelos. An open pampa area within Localities 4 to 10 : Pitfall trapping and handcatches were the Scalesia wood along the western sink hole, charac carried out at different altitudes along the trail "Bellavista terized as a grassland with scattered shrubs. Shrub layer - Media Luna - top of Cerro Crocker". (10 % coverage): the dominant shrub is Hibiscus diver Loc. 4 : Altitude : 500 m (South). Miconia zone. Chop sifolius. Herb layer (100% coverage) : Paspalum ped part completely covered with grasses, ferns and conjugatum, Paspalum galapageium, Stenotaphrum low herbs. secundatum, Scleria pterota and Polypodium u·idens. Lac. 5 : Altitude : 550 m (South). Miconia-wood (35 % Loc. 15 : Altitude : 560 m (North). Typical closed forest coverage) with a dense undercover of Pteridium aquili of the transition zone (trees covered with lichens). Tree/ num ferns (35 %coverage); litter layer appr. 5 em thick. shrub layer (90% coverage) : Psidium ga/apageium, Many death Miconia trees (cfr. 1986). Tournefortia rufo-sericea, Bursera graveolens and Zan thoxylum fagara. Herb layer (20 % coverage) : Plum Loc. 6: Altitude: 700 m (Highland). Fern-sedge zone. bago scandens and Alternanthera echinocephala. Near pool along the trail to Cerro Crocker. Pitfalls located on the verge between pool and dense fern vege Lac. 16: Altitude: 500 m (North). Closed semi-dry tation (Pteridium aquilinum). deciduous forest with a brown dry soil with many seed lings. Dense green tree/shrub layer (80 % coverage) : Loc. 7 : Altitude : 750 m (Highland). Fern-sedge zone. Bursera graveolens, Walthera ovata, Zantho,tylum This sampling site was located between Cerro Puntudo fagara, Pisonia floribunda, Psidium galapageium and and the basis of Cerro Crocker; with 66 % vegetation Piscidia carthagenensis. Very diverse herb layer (20% coverage of which nearly 60% ferns. coverage). Lac. 8 : Altitude : 825 m. Sphagnum bog surrounded Lac. 17: Altitude 400 m (North). Almost closed dry by Lycopodium cernuum (100 % coverage). Northern deciduous forest of the transition type. Tree/shrub layer edge of Cerro Crocker. (80 % coverage) : Tournefortia psilostachya, Bursera Lac. 9: Altitude : 800 m. Crater floor dominated by graveolens, Pisonia floribunda, Zantho.Aylum fagara Lycopodium cernuum and Sphagnum bogs. and Waltheria ovata. Herb layer (20 % coverage) : the dominant species are Bastardia viscosa, Rhynchosia Lac. 10: Altitude 875 m. Top of Cerro Crocker (East minima and an Acanthaceae species. Brown-black soil side). 100 % coverage of grasses, ferns and herbs. between lava rocks.
Locality 11 : Caseta Tortuga (altitude: 180 m). Lac. 18: Altitude 300m (North). Dry deciduous forest of the transition type. Tree/shrub layer (40 % cove Localities 12 to 21 : Pitfall trapping and handcatches were rage) : Bursera graveolens, Pisonia floribunda, carried out at different altitudes on a gradient on the Zantho.Ay lum fagara and Tournefortia psilostachya. northern slope along the main road connecting Puerto Herb layer (60% coverage) :Ipomoea triloba, an Acan Ayora to the Baltra airport. Vegetation mapping of these thaceae species, Blainvillea dichotoma, Heliotropium sites was done by Bosco NOVAK, field assistent at the angiospermium and Abutilum depauperatum. Yellow CDRS. The species of the vegetation are cited in decreas brown soil between lava blocks. ing order of dominance. Only those with a coverage (appr. Lac. /9: Altitude 250m (North). A secondary Walthe %) of 5 % or more are listed (dominant species). ria stand in a dry deciduous forest at the entrance of an exploited cerro. Tree/shrub layer (50% coverage) : Lac. 12 : Los Gemelos (altitude: 570 m). Open ever Waltheria ovata and Tournefortia psilostachya. Herb green woodland of young Scalesia trees (the oldest trees layer (50 % coverage) consisted of Merremia aegyptica ( cfr. 1982 & 1986) died off after the 1982-1983 El and Blainvillea dichotoma. Brown soil with some Nino) situated along the western sink hole. Tree/shrub gravel. layer (70 % coverage) : Scalesia pedunculata parvi flora, Psidium galapageium, Psychotria rufipes, Tour Lac. 20 : Altitude 150 m (North). Open dry deciduous nefortia rufo-sericea, Chiococca alba and Zanthoxylum forest. Palo santo wood with thorny shrubs. Tree/shrub fagara. Herb layer (30% coverage) : the dominant spe layer (40 % coverage) : Bursera graveolens, Zanthoxy cies are the grass Paspalum conjugatum and the fern lum fagara and Acacia insulae-iacobbi. Herb layer Blechnum occidentale. (20 % coverage) : Passiflora suberosa and Rhynchosia minima. Brown soil, rocky with very few litter. Loc. 13 : Los Gemelos. As former site but situated along the eastern sink hole. Tree/shrub layer (70 % Lac. 21 : Altitude 50 m (North). Open dry deciduous coverage) : Sca/esia pedunculata parviflora, Zanthoxy- forest with a tree/shrub layer (20 % coverage) domi- ''
8 L. BAERT, J.-P. MAELFAJT and K. DESENDER
nated by Bursera graveolens. Soil rocky with very few Locality 34 : Cabo Hammond; along small beach; within litter and covered with Passiflora suberosa. Some Opuntia dense Sporobolus virginicus grasses covered with ipomoea cacti (height ca. I m). triloba and with some Scutia pauciflora; along small laguna under tidal influence. Locality 22 : Playa Bachas near laguna (lighttrapping). Locality 35 :Altitude 200m. Cerro Verde; Bursera graveo lens wood with Zanthoxylumfagara, Macraea laric1jolia, ISLA FLOREANA (20.II - 22.II) Cordia sp, Waltheria ovata, Croton scouleri, Darwinio thamnus lan.cifolius glandularis , Alternanthera filifolia Locality 23 : Black beach and dry arid zone : among xero filifolia and grasses. phylous vegetation, under stones and on the beach. Locality 36 : Altitude 400 m. Beginning of encaiiada, soil Localities 24 to 28: West - northwestern gradient along covered by grasses and sedges (mainly Carex species). the road from Puerto Velasco Ibarro up to Cerro Pajas (pitfalls and hand catches). Locality 37: Altitude 600 m. Vegetation (40% coverage) mainly composed of Psychotria rufipes, Zanthoxylum Locality 24 : Altitude 100 m. A dry season deciduous fagara and Scalesia microcephala. Undergrowth of ferns, steppe forest with dense shrub vegetation, a.o. Cordia sedges and different grasses. anderssonii and Acacia ssp. Undergrowth completely dried. Locality 38: Altitude 800 m. Vegetation (total coverage) Locality 25 : Altitude 150 m. Finca of the family Cruz. mainly composed of ferns and grasses with some Zanthoxy lum fagara and Psychotria rufipes shrubs. Locality 26 : Altitude 200 m. A deciduous forest with a thin vegetation of e.g. Bursera graveolens and Acacia Locality 39 : Altitude 1000 m. Total coverage with ferns ssp. Undergrowth completely dried, soil gravelly. and sedges, some sparce Commicarpus tuberosus and Psy chotria rufipes Locality 27 : Altitude 300 m. Transition to Scalesia plants. woodland. Vegetation mainly composed of Tournefor tia rufo-sericea and Scalesia pedunculata; with thin Locality 40: Altitude 1200 m. Volcanic ashes with very shrub undergrowth; rocky soil. few Psychotria rufipes shrubs and grasses (less than 10% coverage). Locality 28 : Altitude 350m. East of Cerro Pajas. Scale sia wood. A mesophytic microphyllous evergreen forest Locality 41 :Altitude 1420 m. Near fumarole at crater rim. dominated by a Scalesia pedunculata - Croton scouleri Shrub layer (40% coverage) composed of Macraea larici community. folia, Psychotria rufipes and Scalesia microcephala. Herb layer (nearly 70% coverage) of annual grasses and few Locality 29 : Highland. Altitude 350 m. Finca of Claudio herbs. Some small patches of barren ground. Cruz. A Psidium guajava-orchard with a dense under growth of grasses and some small shrubs (height : 50 em). Locality 42 : (24.Ill). Punta Espinosa. Pools between pure pahoehoe lava with mangrove. Locality 30 : Highland. Altitude 360 m. Secondary wood of Psidium guajava, Citrus ssp and Zanthoxylum fagara. Many epiphytes (Bromeliaceae and Pipperaceae). Under ISLA ESPANOLA (3 - 9.111) growth mainly composed of ferns. Locality 43 : Punto Cevallos. Along coast. Different places Locality 31 : Highland. Altitude 350 m. At Wittmer well, were sampled : Cerro Wittmer. In litter. - old abandoned army road with a vegetation mainly com posed of Cordia lutea and Acacia rorudian.a; under Locality 32 : Highland. Altitude 340 m. Scalesia wood, stones, between dry grasses; southeast of Cerro Pajas. Mesophytic microphyllous ever - Cryptocarpus pyriformis field; green forest dominated by a Scalesia pedunculata - Croton - "Masked boobies" colony; under stones and between scouleri community. dry grasses.
Locality 3] : Punta Cormorant, eastern shore of laguna. Locality 44 : Altitude 50 m (East). Between rockboulders. Vegetation composed of Cryptocarpus pyriformis, Mayte Vegetation : Cordia lutea and Acacia rorudiana. nus octogona, Avicenna germinans, Lagun.cularia race mosa and mangroves. Locality 45 : Altitude 80 m. Vegetation composed of Lan tana peduncularis, Acacia rorudiana and Cordia lutea. Red brown soil covered with a thick litter layer, seedlings ISLA FERNANDINA (24.11 - 28.II, 24.III) and dry herbs. Araneae of the Galapagos 9
Locality 46 : Altitude 110 m. Under stones along rockwall. Locality 57: Lago Arcturus, bottom of crater, southwestern Vegetation: Cordia lutea, Parkinsonia aculeata and border of the lake. Rocks covered with mangrove and Acacia rorudiana. Cyperaceae.
Locality 47: Altitude 175 m. Subtop of island. Vegetation mainly composed of Cordia lutea, Lantana peduncularis, ISLA DAPHNE MAJOR (15.11I) Parkinsonia aculeata and Acacia rorudiana. Locality 58 : Crater bottom with a sparse vegetation mainly composed of Opuntia echios, a few Croton scouleri shrubs, ISLA MAR CHEN A (1 0 - 1l.III) dry herbs and grasses.
Locality 48: Playa Negra, along coast. Vegetation girdle Locality 59 : Outer flanks of the crater. mainly composed of Cordia lutea, Bursera graveolens, Lantana peduncularis, Opuntia helleri, Croton scouleri and CryptocaJpus pyriformis stands; under stones and in ISLA SAN CRISTOBAL (2 - 8.ill - leg. Bosco NovAK) vegetation.
Locality 49: Playa· Negra, altitude 25 m, inland. Dense Locality 60 : Highland, altitude 540 m. Miconia robinso shrub/bush layer mainly composed of Bursera graveolens, niana - and Psidium guajava forest with Panicum laxum, Croton scouleri, Lantana peduncularis, Chamaesyce vimi Centella asiatica grass and Pteridium ferns. nea and Caste/a galapageia. Herblayer richer and thicker than in following locality. Locality 61 : Highland, altitude 600 m. Along border of the "El Junco" lake. Vegetation: Polygonum sp., Rubus Locality 50 : Playa Negra, altitude 50 m inland. Dense sp., Centella asiatica, Panicum laxum and Psidium gua shrub/bush layer composed of Bursera graveolens, Croton java. scouleri, Lantana peduncularis and Waltheria ovata, some Opuntia helleri and a few sedges. Very thin litter layer. Locality 62 : Near the house of Senor RosiLLO, some 2 km from El Junco, altitude appr. 550 m. Locality 51 : Punta Mejia. Different stands along the coast were sampled : - CryptocaJpus pyriformis belt; ISLA SANTIAGO (19.III) - Ash slope covered with Coldenia sp., Tiquilia sp. and Opuntia helleri; Locality 63 : Playa Espumila, laguna with mangrove (light - Croton scouleri belt located behind Cryptocarpus belt trapping). with a vegetation mainly composed of Opuntia helleri, Croton scouleri, Bursera graveolens, Cordia lutea, some Waltheria ovata, Lantana peduncularis and Caste/a ISLA ISABELA - VOLCAN WOLF (20 - 24.III) galapageia, few grasses and Tiquilia herbs. Locality 64 : Landing site north of Pun to Bravo. 1° Camp site with Maytenus octogona and Bursera graveolens vege ISLA GENOVESA (12 - 14.III) tation; 2° In Sporobolus pyramidatus grass behind man grove girdle; 3° On the beach (lighttrapping). Locality 52 : Beach of Bahia Darwin. Cryptocarpus pyri formis stand along small pool subject to tides; under Man Locality 65 : Lowland; Bursera graveolens wood with Lan grove. tana peduncularis, Alternantherafilifolia, Cordia lutea and Locality 53 : Along Northern Barranco at Bahia Darwin, Scutia parciflora. Herb layer completely dry. bare lava with Opuntia helleri and Cordia lutea as main vegetation. Locality 66: Altitude 125 m. Under Pisonia floribunda tree. Vegetation composed of Bursera graveolens, Zan Locality 54 : Bahfa Darwin, eastern barranco. Main vegeta thoxylum fagara, Tournefortia sp., Waltheria ovata and tion : Bursera graveolens, Opuntia helleri and Cordia Gossypium sp.; undergrowth (20% coverage) with herbs, lutea. Passiflora sp.,sedges and grasses; few litter. Locality 55 : Lago Arcturus. First platform inside the cra ter, altitude 60 m. Vegetation composed of Cordia lutea, Locality 67 : Altitude 350 m. Under Pisonia jloribunda Bursera graveolens, few Croton scouleri and Opuntia hel tree in dense Croton scouleri wood with an undergrowth leri. Rocky. (40 % coverage) of annual grasses.
Locality 56 : Lago Arcturus, bottom of crater, northern Locality 68 : Altitude 400 m. Under Pisonia floribunda border of the lake, between mangrove. tree. Bare soil with moderate litter layer. 10 L. BAERT, J. -P. MAELFAIT and K. DESENDER
Locality 69 : Altitude 600 m. Shrublayer (70 % coverage) Locality 81 : Altitude 400 m. Shrub layer (60 % coverage) composed of Pisoniafloribunda, Bursera graveolens, Psi dominated by Bursera graveolens, Croton scouleri and dium galapagaeium and Zanthoxylum fagara; with green Waltheria ovata; furthe1more a few Pisonia floribunda, veil vegetation. Thick litter layer. Scalesia affinis and Opuntia insularis. Bare lava soil with a few grasses, sedges and herbs. Locality 70: Altitude 675 m. Vegetation comparable with previous site. Locality 82 : Altitude 500 m. Shrub layer (15 % coverage) with Bursera graveolens, Croton scouleri, Macraea larici Locality 71 : Altitude 825 m. Vegetation comparable with folia, Zanthoxylum fagara, Opuntia insularis and Walthe both previous sites but here with rich herblayer of grasses ria ovata. Some lichens hanging on the shrubs. Stony and ferns. underground, half covered with grasses, sedges and various herbs. Locality 72 : Altitude 1000 m. Appr. 50 % coverage with Zanthoxylum fagara, Toumefortia sp., Cordia lutea, Locality 83 : Altitude 600 m. Shrub layer (85 %coverage) Macraea laricifolia and Psidium galapagaeium. Total dominated by Scalesia microcephala, Bursera graveolens coverage with fern undergrowth (cfr. Pteridium) and creep and Croton scouleri; furthermore a few Zantho;rylum ing plants. Rocky (lava boulders) with thick litter layer, fagara and Tournefortia sp .. Rocky underground nearly no soil. half covered with grasses, creeping plants, ferns and a few herbs (e.g. Darwiniothamnus tenuifolius); A lot of wood Locality 73: Altitude 1200 m. 60-70 % coverage with debris. Zanthoxylum fagara, Croton scouleri, Scalesia sp., Locality 84 : Altitude 800 m. Shrub layer (70% coverage) Macraea laricifolia, Psidium galapagaeium, Ipomoea sp., dominated by Croton scouleri, Scalesia microcephala and ferns and Opuntia sp. Rocky (lava boulders) with soil but Macraea laricifolia; further a few Caste/a galapageia and less litter than in previous site. Dodonaea viscosa. Herb layer (80% coverage) dominated Locality 74 : Altitude 1425 m. Vegetation mainly com by grasses, a few sedges and herbs (e.g. Lantana peduncu posed of Opuntia sp., Scalesia sp.; herb layer (40-50% laris, Darwiniothamnus tenuifolius), ferns and creeping coverage) with Commicarpus sp., Lantana peduncularis, plants. Zanthoxylum fagara, various fern species and grasses. Locality 85 : Altitude 900 m. Shrub layer (85 % coverage) dominated by Macraea laricifolia (shrub), Dodonaea vis Locality 75: Altitude 1625 m. Vegetation (20% coverage) cosa and Zanthoxylum fagara; some Opuntia insularis and mainly composed of two Scalesia species, Macraea larici Scalesia microcephala. A herb layer (40-50% coverage) folia in tree form, Opuntia sp. and "Garrapatero"-grasses. of grasses. Locality 76 : Altitude 1700 m, top of vulcano, crater rim. Locality 86: Altitude 1000 m. Shrub layer (70% coverage) Shrub layer (20% coverage) with Opuntia sp., Macraea dominated by Macraea laricifolia (shrub) and Dodonaea laricifolia in tree form, two Scalesia species and Tow-ne viscosa; also with some Opuntia insularis, Zantho.l)•lum fortia sp .. Herb layer (70 % coverage) composed of an fagara, the two Scalesia species and Croton scouleri. A Alternanthera sp. and "Garrapatero"-grasses. herb layer (25 % coverage) of grasses and ferns. Locality 87 : Altitude 1100 m. Shrub layer ( 1.5 m high, ISLA ISABELA - VOLCAN DARWIN (25 - 28.lll) 60 % coverage) dominated by Macraea laricifolia (shrub) with Dodonaea viscosa, Scalesia microcephala and Opun Locality 77 : Beagle crater, landing site, beach (lighttrap tia insularis. Soil gravelly and covered with lichens and a ping). herb layer (35 % coverage) of grasses, ferns and some herbs. Locality 78 : Beagle crater; outer western crater flank. Locality 88: Altitude 1200 m. Shrub layer (60% coverage) Shrub layer (80% coverage) mainly composed of Cordia dominated by Macraea laricifolia (shrub) and Dodonaea lutea and Bursera graveolens. Herb layer : a few dry gras viscosa; with some Scalesia sp., Opuntia insularis and ses. Soil gravelly. Zanthoxylum fagara. Herb layer (30% coverage) domi Locality 79 : Beagle crater, eastern border of inner lake. nated by ferns but with a few grasses. Total coverage with Spirobolus pyramidatus grasses. A Locality 89 : Altitude 1200 m. Lava tunnel. Fine brown Cordia lutea shrub was also sampled. humid soil mainly covered by ferns (30% coverage). Locality 80: Altitude 200m. Shrub layer (80% coverage) Locality 90 : Altitude 1300 m. Shrub layer (80% coverage) dominated by Bursera gra veolens, with Zanthoxylum dominated by Scalesia and Dodonaea viscosa; furthern1ore fagara, Scalesia affinis, Croton scouleri, Waltheria ovata, a few Zanthoxylumfagara, Macraea laricifolia (shrub) and Caste/a galapageia, Opuntia insularis and Cordia leuco Opuntia insularis. Black soil covered with a herb layer phlyctis. Herb layer (80 % coverage) consisting of dry gras (50% coverage) dominated by grasses and with some ferns ses. and herbs. ,,
Araneae of the Galapagos 11
ISLA ISABELA - VOLCAN ALCEDO (28.III) ESPANOLA (III): Locality 45 U), Locality 46 U), Locality 47 (2 j). Locality 91 : Bahfa Urvina. Vegetation along black beach MARCHENA (III) : Locality 48 (2 'i? 'i?, 2 j), Locality 50 mainly composed of Cordia lutea and Waltheria ovata. (I SA 'i?), Locality 51 (1 SA 'i? , 2 j 'i? 'i?, j). GENOVESA (III) : Locality 53 (2 j 'i? 'i? ), Locality 54 (I o, 1 SA 'i?, 13 j). ISLA ISABELA - VOLCAN SIERRA NEGRA (29.III) DAPHNE MAJOR (III) : Locality 58 ( 1 o, 4 'i? 'i?, 5 SA 'i? 'i?, 2 j), Locality 59 (3 'i? 'i? ). Locality 92 : Puerto Villamil. Lighttrapping along dunes SANTIAGO (Ill) : Locality 63 ( 1 SA o, 1 'i?, 2 SA and lagoons with mangrove. 'i? 'i?, 2 j). ISABELA : Volcan Wolf (Ill) : Locality 64 (1 SA 'i? ), Locality 65 (2 SA o o , j), Locality 66 (SA o), Locality SYSTEMATIC ACCOUNT 74 U). Volcan Darwin (III) : Locality 79 (2 j), Locality 80 (1 SA o ), Locality 81 ( 1 SA o, j 'i? ), Locality 82 ANYPHAENIDAE ( 1 'i? ), Locality 83 U), Locality 88 U).
Anyphaenoides pacifica (Banks, 1902) Other material examined
Material sampled in 1988 - cfr BAERT & MA ELFAIT (1986): p. 97, Map 3; - cfr BAERT, MAELFA IT & DESENDER (1989); SANTA CRUZ: Locality 2 (1 'i? : Ill), Locality 20 (1 SA - FRANZ, 1975: SANTA CRUZ: Cerro Colorado (1 'i? : 'i? : IV). V), Playa Tortuga (1 'i? :VI). FLOREANA: Post Office FERNANDINA (II): Locality 35 (1 o).MARCHENA Bay (I 'i?, j : V), Punta Cormorant (I 'i? : V). SAN (Ill) : Locality 48 (2 o o ), Locality 50 (2 j), Locality 51 CRISTOBAL : Puerto Grande (I 'i? , j : VI). SANTA (2 j). FE : U: V). SANTIAGO : Bucanero camp U: VI). ISABELA: Vo lcan Wolf (III) : Locality 65 (3 'i? 'i? ), Loca lity 66 (1 o, j), Locality 73 (2 j), Locality 74· (1 o, 1 SA Distribution o, j 'i? ), Locality 75 (1 o, 1 SA 'i?, 2 j), Locality 76 (3 o o, 1 'i?, j). Vo lcan Darwin (III) : Locality 80 (1 'i? ), Baltra, Daphne maior, Espanola, Fernandina, Floreana, Locality 82 (1 'i?, j), Locality 84 (1 'i? ), Locality 85 (1 SA Genovesa, lsabela (Darwin, Alcedo, Sierra Negra, Cerro 'i? ), Locality 86 (3 o o , 2 'i? 'i?, 3 j), Locality 87 (4 j), Azul, Wolf*), Marchena, Pinta, Pinzon, Rabida, San Cris Locality 88 (1 SA o, 1 SA 'i? ), Locality 90 ( 1 o, 1 'i?, tobal, Santa Cruz, Santa Fe, Santiago and Seymour Norte. 6 j).
Other material examined Cyclosa turbinata (Walckenaer, 1841)
- cfr BAERT & MAELFA IT (1986): p. 97, Map 7; Material sampled in 1988 - cfr BAERT, MAELFAIT & DESENDER (1989); - FR ANZ, 1975: PINZON: crater rim (1 o, 1 'i? : VI). FLOREANA (II): Locality 29 (1 'i? ). SAN CRISTOBAL : Puerto Grande (1 'i? : VI). FERNANDINA (II): Locality 34 (2 'i? 'i?). MARCHENA (III): Locality 51 (1 SA 'i?). Distribution Other material examined Floreana, Fernandina*, Isabela (Alcedo, Darwin*, Wolf*), Marchena*, Pinta, Pinzon, San Cristobal, Santa Cruz, San - cfr. BAERT & MAELFA IT (1986) : p. 98, Map 4; tiago and Seymour Norte. - cfr. BAERT, MAELFAIT & DESENDER (1989).
Distribution ARANEIDAE Baltra, Espanola, Fernandina, Floreana, Genovesa, Isabela Argiope argentata (Fabricius, 1775) (Darwin), Marchena *, Rabida, San Cristobal, Santa Cruz, Santa Fe and Santiago. Material sampled in 1988
SANTA CRUZ (II) : Locality 3 (1 'i? , j). Gaster acantha cancriformis (Linn aeus, 1767) FLOREANA (II) : Locality 24 (1 'i? , j), Locality 25 U 'i? ), Locality 26 (1 'i?, 1 SA 'i? ), Locality 33 (l SA 'i?) . Material sampled in 1988 FERNANDINA (II) : Locality 34 (1 o, 7 'i? 'i?, j 'i?, 2 j), Locality 36 (1 o ). SANTA CRUZ (II) : Locality 11 (SA o, 2 'i? 'i? ). ''
12 L. BAERT, J.-P. MAELFAIT and K. DESENDER
FLOREANA (II) : Locality 25 (2 - cfr. BAERT & MAELFAIT (1986): p. 100, Map 6; Metepeira desenderi Baert, 1987 - cfr. BAERT, MA ELFAIT & DESENDER (1989); - FRANZ, 1975: FLOREANA: Post Office Bay (j o : V). Material sampled in 1988 SAN CRISTOBAL: Cerro San Joaquin (1 Other material examined CLUBIONIDAE - cfr. BAERT (1987): p. 145; - cfr. BAERT, MAELFAIT & DESENDER (1989). Corinna wollebooki Banks, 1930 Distribution Material sampled in 1988 Daphne maior, Espanola, Fernandina, Floreana, Genovesa, SANTA CRUZ (III) : Locality 2 (8 o o, 2 Other material examined Neoscona cooksoni (Butler, 1877) - cfr. BAERT & MAELFAIT (1986): p. 100, Map 7; Material sampled in 1988 - cfr. BAERT, MAELFAIT & DESENDER (1989). SANTA CRUZ: Locality 3 (j Araneae of the Galapagos 13 CTENIDAE Other material examined Odo insularis Banks, 1902 - cfr. BAERT, MAELFAIT & DESENDER (1989); - FRANZ, 1975 : SANTIAGO : Bucanero camp (3 2 2, Material sampled in 1988 j 2 : VI). SANTIAGO (III) : Locality 63 (2 2 2, j). Distribution ISABELA: Volcan Wolf (III): Locality 64 (1 2, 2 SA 2 2 ), Locality 74 (1 2, j 2 ). Volcan Darwin (III) : Loca Fernandina, Espanola*, Isabel a (Aicedo, Darwin*, Sierra lity 79 (1 2 ). Negra, Wolf*), Marchena*, Rabida, Santa Cruz and San tiago. Other material examined - cfr. BAERT(l987): p. 154; DYSDERIDAE - cfr. BAERT, MAELFA IT & DESENDER(l989); - FRANZ, 1975: SANTIAGO : James Bay (1 2 : VI). Ariadna tarsalis (Banks, 1902) Distribution Material sampled in 1988 Baltra, Eden, Fernandina, Isabela (Alcedo, Darwin, FERNANDINA (III) : Locality 42 (1 o, 1 2, 2 j 2 2 ). Wolf*), Pinzon, Santa Cruz and Santiago. ISABELA: Volcan Wolf (III): Locality 66 (1 2, 3 j 2 2 ), Locality 67 (1 o ), Locality 68 (1 2 ). Other material examined DICTYNIDAE - cfr. BAERT, M AELFA IT & DESENDER(l989): - FRANZ, 1975 : SANTA FE : (j 2 : V). SAN CRISTO- Emblyna formicaria Baert, 1987 BAL : Cerro El Junco (2 j 2 2 : V); - LUBIN, 1982: PINTA: !500ft alt. (1 o :II). SANTIA Material sampled in 1988 GO : Los Jaboncillos, 2700ft (I 2 : IV). ISABELA : Vo lcan Wolf (III) : Locality 75 (1 2 ). Distribution Other material examined Baltra, Culpepper, Fernandina, Isabela (WolP), Pinta, Pin zon, San Cristobal*, Santa Cruz, Santa Fe* and Santiago. - cfr. BAERT(l987) : p. 149. Distribution EUSPARRASIDAE Isabela (Beagle crater, Wolf*). Heteropoda venatoria Linnaeus, 1767 Tivyna spathula (Gertsch & Davis, 1937) Material sampled in 1988 Material sampled in 1988 SANTA CRUZ (II): Locality 2 (I 2). FLOREANA (III) : Locality 29 (2 2 2 + egg cocoon). SANTA CRUZ : Locality 2 (SA o, 3 2 2, 4 j : III), SANTIAGO (III) : Locality 63 (I o). Locality 19 (1 o, 2 2 2 : II; 3 o o, 2 2 2 : IV), Locality 20 (2 o o : III; 2 2 2 : IV), Locality 2 1 (I o : IV). Other material examined FERNANDINA (II) : Locality 34 (3 o o, 3 2 2, 3 j), Locality 35 (1 o, SA o, j). - cfr. BAERT & MAELFAIT (1986): p. 102, Map 9; ESPANOLA (III): Locality 43 (SA o). - cfr. BAERT, MAELFA IT & DESENDER (1989); MARCHENA (III) : Locality 48 (3 SA o o, 7 j), Locality - FRANZ, 1975: SANTA CRUZ : CDRS-building (I o : 51 (6 o o, 2 SA o o, 7 2 2, j). V). SANTIAGO (III) : Locality 63 (3 o o, SA o, 2 2 2 ). ISABELA: Vo lcan Wolf(III): Locality 65 (SAo). Volcan Distribution Darwin (Ill) : Locality 79 (I 2 ), Locality 80 (1 o ,j o, j 2 ). Volcan Alcedo (III) : Locality 91 (7 o o, 2 2 2, 2 j). Floreana, Isabela (S ierra Negra), San Cristobal, Santa Cruz Volcan Sierra Negra (III): Locality 92 (1 o, SA o). and Santiago*. ' ' ' 14 L. BAERT, J.-P. MAELFAIT and K. DESENDER Olios galapagoensis Banks, 1902 vesa, Isabela (Alcedo, Darwin, WolF), Marchena, Pinta, Pinzon, Santa Cruz, Santa Fe, Santiago, Wolf. Material sampled in 1988 SANTA CRUZ: Locality 2 (1 o : II; 1 'il : III), Locality GNAPHOSIDAE 8 U: III), Locality 10 (1 o : II), Locality 12 (SA o : III), Locality 13 (1 S? : III), Locality 16 U S? : IV) Locality 17 Poecilochroa bifasciata Banks, 1902 U : IV), Locality 18 U 'il : III). FERNANDINA (III) : Locality 42 U). Material sampled in 1988 MARC HENA (III) : Locality 48 ( 1 o, SA o), Locality 49 (1 o, 1 S? ), Locality 51 (SA o). MARCHEN A (III) : Locality 50 (1 <;? ). ISABELA: Volcfm Wolf (III): Locality 72 U), Locality ISABELA: Volcan Wolf (Ill): Locality 65 (1 o, 2 j). 73 (SA 'il, 2 j), Locality 75 (SA o), Locality 76 (1 Distribution Trachyzelotes kulckzynski (Bosenberg, 1902) Baltra, Eden, Espanola (Gardner), Fernandina, Isabela Material sampled in 1988 (Darwin, Sierra Negra, WolF), Marchena*, Pinta, Pinzon, San Cristobal, Santa Cruz and Santiago. SANTA CRUZ: Locality 17 (1 o : II; 2 o o : IV). Other material examined FILISTATIDAE - cfr. BAERT, MAELFAIT & DESENDER(l989). Filistatoides fasciatus (Banks, 1902) Distribution Material sampled in 1988 ESPANOLA (Ill): Locality 43 (2 <;? <;?, 3 j), Loca San Cristobal and Santa Cruz. lity 47 (2 j). MARCHEN A (III) : Locality 49 (5 <;? <;? ), Locality 50 (3 <;? <;?, 14 j), Locality 51 (6 <;? <;?, 6 j). LINYPHIIDAE GENOVESA (III) : Locality 52 U). DAPHNE MAJOR (III): Locality 59 (1 <;?). Laminacauda dentichelis Millidge, 1985 ISABELA : Volcan Wolf (III) : Locality 65 (1 <;?, 3 j), Locality 66 U). Volcan Datwin (III) : Locality 78 (1 S?, Material sampled in 1988 2 j). SANTA CRUZ : Locality 5 (I o : II), Locality 6 (1 o : Other material examined ill), Locality 7 (3 o o, 6 S? <;? : II; 2 o o, 1 <;? : III; 2 <;? <;? : IV), Locality 8 (2 o o, SA o, 2 Negra, crater rim (fern-sedge zone), alt. 1060 m, 23.III LOXOSCELIDAE (9 o o, 18 S? S?, 10 j). SAN CRISTOBAL: Along road to El Junco, alt. 600 m (near swamp), 3.III Loxosceles laeta (Nicolet, 1849) (5 o o, 6 S? S? ); crater rim of El Junco (fern-sedge zone), 3.1II o o); Material sampled in 1988 - FRANZ, 1975 : ISABELA : Santo Tomas, Cueva de Zueca, 29.V (1 o, I SA S? ); ESPANOLA (III) : Locality 45 U <;> ), Locality 46 (2 j - PECK,1985: SANTA CRUZ: Media Luna, alt. 620 m, <;> <;> ). Miconia vegetation, 14.V-13.VII (5 o o, I S? ); ISABELA: Voldm Wolf (III): Locality 64 U). - ABEDRABO S. ,1986: ISABELA: Volcan Sierra Negra: alt. 900 m, 10-14.II (8 oo, 6 S? S?), 15-17.IV (82 Other material examined o o, 18 S? <;> ), 8-20.VII (140 o o, 54 <;> <;> ). - cfr. BAERT & MAELFAIT (1986): p. 104, Map 17; Distribution - cfr. BAERT, MAELFAIT & DESENDER (1989). Isabela (Sierra Negra), San Cristobal and Santa Cruz. Distribution Floreana, Espanola, Isabela (Wolf"), Pinta, Pinzon, Rabida, Santa Cruz, Santa Fe and Santiago. Notiohyphantes excelsa (Keyserling, 1886) Material sampled in 1988 METIDAE SANTA CRUZ : Locality 4 (SA S? : III), Locality 5 (SA Leucauge bituberculata Baert, 1987 o : II; 1 o, 1 <;> : III), Locality 6 (1 S? : IV), Loca- lity 7 (2 <;> <;> : II; SA S? : III), Locality 10 (2 S? S? : Material sampled in 1988 II), Locality 11 (I o, 4 S? S?, 2 SA o o : II). FLOREANA (II) : Locality 29 (1 o, 3 S? S? ), Locality 30 SANTA CRUZ: Locality 5 (1 <;> : II), Locality 8 (1 S? : (1 o, 2 SA o o, 2 S? S?, 2 SA S? S? ), Locality 31 III), Locality 9 ( 4 S? S? : III), Locality 10 (2 o o, 3 S? S? : (2 j <;> <;> ). II; SA o, ·2 SA S? <;>, 3 j: III; 1 <;> : IV), Locality 11 MARCHENA (III): Locality 48 (SA o). (3 S? S?, j S? : II). SAN CRISTOBAL (III) : Locality 60 (SA S? ). FLOREANA (II) : Locality 29 (2 S? <;>, j), Locality 30 (3 <;> <;> ' 4 j). Other material examined FERNANDINA (II) : Locality 35 (2 o o, 2 <;> S? ), Locality 36 (SA o, I S?, 6 j), Locality 37 (7 S? S?, 3 j), - cfr. BAERT, MA ELFAIT & DESENDER (1989); Locality 38 (SA o, SA S? ). - BAERT & MA ELFA IT, 1982 : SANTA CRUZ: Caseta IS ABELA : Volcan Wolf (III) : Locality 65 U), Locality 66 Occidente, alt. 700 m, 17-18.III ( 4 S? S?, 4 j.). El Chato (SA o, 1 <;> ), Locality 67 U o), Locality 68 (3 S? S?, j), trail : alt. 200 m, 30.III (9 o o, I 0 S? S? ); alt. Locality 69 U), Locality 71 (SA S? ), Locality 73 (SA o) , 280 m, 30.III (3 o o, 3 S? S? ). Media Luna trail : Locality 74 (1 <;> ), Locality 76 (3 j). Voldm Darwin alt. 200-250 m (Culture zone), lO.III (1 o, 10 S? S?, 4 j.); (III) : Locality 89 (2 S? S? ), Locality 90 U). alt. 600 m (Media Luna, Sphagnum), lO.III (2 S? S?, 1 SA o); alt.600 m (pampa), 8.III (1 S? ). SAN CRISTO Other material examined BAL : Along road to El Junco : Between El Progreso and the El Junco lake, alt. 500 m (Aguayava vegetation, - cfr. BA ERT (1987): p. 143; 3.lll (6 o o, 9 <;> <;> ); alt. 600 m (road verge, near - cfr. BAERT, MAELFAIT & DESENDER (1989). swamp), 3.III (1 o, 7 S? S?, 8 j); El Junco lake (pampa), alt. 700 m, 3.III (8 S? S?, 6 j.); Distribution - ScHATZ, 1985 : SAN CRISTOBAL : Miconia zone, 29.III o o); Fernandina, Floreana*, Isabel a (Cerro Azul, Darwin*, - C.A.S. : SANTA CRUZ : Cerro Mesa, alt. ca 440 m, Sierra Negra, Wolf*), Pinta, San Cristobal, Santa Cruz and 16.IV.I964 (2 o o); Santiago. - C.D.R.S.: SANTA CRUZ: Media Luna, 16.III.l982 (2 o o). MIMETIDAE Distribution Ero gemelosi Baert & Maelfait, 1984 Floreana*, Isabela (Sierra Negra), Marchena*, San Cristo- bal and Santa Cruz. Material sampled in 1988 I I 16 L. BAERT, J.-P. MAELFAIT and K. DESENDER ISABELA: Volcan Wolf (III): Locality 72 (1 SA 2), OECOBIIDAE Locality 73 (1 2 ). Oecobius concinnus Simon, 1892 Other material examined Material sampled in 1988 ' - cfr. BAERT & MAELFAIT (1986) : p. 106; - cfr. BAERT, MAELFAIT & DESENDER (1989). SANTA CRUZ : Locality 2 (1 9 : II; 1 o : III). FLOREANA (II): Locality 24 (1 o). Distribution ESPANOLA (III) : Locality 43 (9 o o, 6 2 9 ), Locality 45 (1 o), Locality 46 (1 SA o), Locality 47 U 2). Isabe1a (Wolf)* and Santa Cruz. IS ABELA : Volcan Alcedo (III) : Locality 91 (1 9 ). Other material examined MYSMENIDAE - cfr. BAERT & MAELFAIT (1986): p. 106, Map 17; - cfr. BAERT, MAELFAIT & DESENDER (1989). Calomyspoena santacruzi Baert & Maelfait, 1983 Distribution Material sampled in 1988 Espaiiola*, Floreana, Isabela (Alcedo)*, Pinzon, San Cris SANTA CRUZ : Locality 5 (2 2 2 : III), Locality 7 tobal, Santa Cruz, Santa Fe and Santiago. (1 2 : II), Locality 13 (1 o : IV). FLOREANA (II) : Locality 29 (I 2 ). ISABELA : Volcan Wolf (III) : Locality 72 (1 2 ). Val OXYOPIDAE can Darwin (III) : Locality 82 ( 1 2 ), Locality 87 (1 2 ). Oxyopes gracilis Keyserling, 1877 Other material examined Material sampled in 1988 - cfr. BAERT & MAELFAIT (1986): p. 106, Map 13; SANTA CRUZ : Locality 6 U : III; 1 o, 1 2 : IV), - cfr. BAERT, MAELFAIT & DESENDER (1989); Locality 7 (1 o, 2 j : II), Locality 10 U : IV), Locality - FRANZ, 1975 : PINZON : crater rim (1 2 : VI). 14 U: II; 1 o, 3 SA o o, 1 SA 2, 12 j: III; 1 SA o, 1 SA 9 : IV). Distribution FERNANDINA (II) : Locality 39 (1 9 ), Locality 41 (1 o, 1 9' 2 j). . Floreana*, Isabela (Darwin*, Sierra Negra, Wolf~), Pin MAR CHEN A (III) : Locality 49 (1 9 ). zon*, San Cristobal, Santa Cruz and Santiago. ISABELA : Volcan Wolf (III) : Locality 65 U 2 ), Loca lity 74 U), Locality 76 U). Volcan Darwin (III) : Locality 90 U). OCHYROCERATIDAE Other material examined Theotima galapagosensis Baert & Maelfait, 1986 - cfr. BAERT & MAELFAIT (1986): p. 107, Map 15; - cfr. BAERT, MAELFAIT & DESENDER (1989). Material sampled in 1988 Distribution SANTA CRUZ (II): Locality 5 (4 2 2). FLOREANA (II) : Locality 27 (15 9 9 ), Locality 31 Fernandina, Isabela (Alcedo, Cerro Azul, Darwin*, (11 9 9 ), Locality 32 (I 9 ). Wolf*), Marchena*, Pinta, Santa Cruz and Santiago. Other material examined PHOLCIDAE - cfr. BAERT & MAELFA IT (1986) : p. 106, Map 14; - cfr. BAERT, MAELFAIT & DESENDER (1989). Coryssocnemis conica Banks, 1902 Distribution Material sampled in 1988 Floreana*, Jsabela (Cerro Azul), San cristobal and Santa SANTA CRUZ : Locality 6 U : IV), Locality 7 (1 o, Cruz. 1 9 : II; 1 SA o : IV), Locality 8 (1 o, 2 j : III; 1 9 : '' Araneae of the Galapagos 17 IV), Locality 9 (2 Espanola, Fernandina, Floreana, Isabela (Cerro Azul, Dar win*, Sierra Negra*, Wolf*), Pinta, Pinzon, Rabida, San Frigga crocuta (Taczanowski, 1879) Cristobal, Santa Cruz, Santa Fe, Santiago and Seymour Norte. Material sampled in 1988 SANTA CRUZ (III): Locality I7 ( I o). Coryssocnemis insularis Banks, 1902 FERNANDINA (II) : Locality 34 (I SALTICIDAE Baltra, Fernandina, Floreana, Isabela (Alcedo, Darwin, Sierra Negra, Wolf*), Pinzon, San Cristobal, Santa Cruz Darwinneon crypticus Cutler, 1971 and Santiago. Material sampled in 1988 Metacyt·ba insularis (Banks, 1902) SANTA CRUZ : Locality 3 (5 o o, 2 18 L. BAERT, J.-P. MAELFAJT and K. DESENDER Other material examined Scytodes hebraica Simon, 1891 - cfr. BAERT & MAELFAIT (1986): p. 110, Map 19; Material sampled in 1988 - cfr. BAERT, MAELFAIT & DESENDER (1989); - FRANZ, 1975 : SAN CRISTOBAL : Playa Ochova SANTA CRUZ : Locality 2 (1 'i' : III; 1 'i' : IV), Locality (1 o : V); Puerto Grande (1 o, 1 SA o, j : VI). 10 (1 o : II; 1 o : IV), Locality 13 (1 ~ : III). PINZON : Crater rim (j ~ : VI). SANTA FE : (1 o, FLOREANA (II): Locality 23 (1 o). 1 SA o: V). GENOVESA (Ill) : Locality 54 (j ~ ), Locality 55 (1 o, 2 SA o o, 2 ~ ~, 2 SA ~ ~, j), Locality 56 (1 o, Distribution 2 SA o, 3 ~ ~, 4 j), Locality 57 (1 o, 1 SA o, 4 ~ ~' 3 j ~ ~). Daphne, Espanola*, Fernandina, Floreana, Isabela (Dar win, Wolf*), Pinzon, San Cristobal*, Santa Cruz, Santa Other material examined Fe* and Santiago. - cfr. BAERT & MAELFAIT (1986): p 113, Map 21; - cfr. BAERT, MAELFAIT & DESENDER (1989); Philaeus pacificus Banks, 1902 - FRANZ, 1975: FLOREANA: Post Office Bay (1 SA o : V). ISABELA: Cerro Azul, Caleta Iguana (1 o, Material sampled in 1988 3 Other material examined Scytodes longipes Lucas, 1845 - cfr. BAERT & MAELFAIT (1986): p. 110; Material sampled in 1988 - cfr. BAERT, MAELFA IT & DESENDER (1989); - FR ANZ, 1975 : FLOREANA: Post Office Bay (1 o : SANTA CRUZ (II): Locality 2 (1 o, 1 SA o). V). PINZON: Crater rim (2 o o : VI). SANTIAGO: FLOREANA (II) : Locality 23 (j Material sampled in 1988 Distribution FLOREANA (11) : Locality 23 (2 J 'i' ), Locality 27 Floreana, Isabela (Darwin*, Sierra Negra), San Cristobal (1 ~ ). and Santa Cruz. Other material examined SELENOPIDAE - cfr. BAERT & MAELFAIT (1986): p.113, Map 21; - cfr. BAERT, MAELFAIT & D ESENDER (1989). Selenops galapagoensis Banks, 1902 Distribution Material sampled in 1988 Floreana, Genovesa, Isabela (Sierra Negra), Rabida, San SANTA CRUZ: Locality 2 (I SA ~ , 2 j ~ : U; 1 SA Cristobal and Santa Cruz. o : III), Locality 16 (j : Ill). I I Araneae of the Galapagos 19 FLOREANA (II) : Locality 23 (2 j). Tetragnatha nitens (Audouin in Savigny, 1825) ISABELA: Volcan Wolf (III): Locality 65 (2 S? Distribution Fernandina, Isabela (Sierra Negra) and Santiago. Fernandina, Floreana, Isabela (Darwin*, Sierra Negra*, Wolf*), San Cristobal and Santa Cruz. THERIDIIDAE Achaearanea hirta (Taczanowski, 1872) SICARIIDAE Material sampled in 1988 Sicarius ultriformis (Butler, 1877) SANTA CRUZ (II): Locality 3 (1 Other material examined - cfr. BAERT & MAELFAIT (1986) : p. 115, Map 23; - cfr. BAERT, MA ELFA IT & DESENDER (1989). - cfr. BAERT & MAELFAIT (1986): p. 115, Map 21; - cfr. BAERT, MAELFAIT & DESENDER (1989). Distribution Distribution Isabela (Darwin), Marchena*, Rabida, San Cristobal, Santa Cruz and Santiago. Eden, Espanola, Floreana, Rabida and Santiago. Argyrodes elevatus Taczanowski, 1872 TETRAGNATHIDAE Material sampled in 1988 Glenognatha maelfaiti Baert, 1987 SANTA CRUZ (II) : Locality 11 (I S? ). Material sampled in 1988 FLOREANA (ii) : Locality 23 (1 S? ), Locality 24 (I o ), Locality 27 (2 j S? S? ). SANTA CRUZ: Locality 4 U: IV), Locality 5 (I S? : II; FERNANDINA (III) : Locality 42 (1 o). l o, 3 D i strihu t ion Other material examined Floreana*, Isabela (Cerro Azul, Sierra Negra) and Santa - cfr. BAERT & MA ELFAIT (1986): p. 11 5, Map 24; Cruz. - cfr. BAERT, MA ELFA IT & DESENDER (1989). I I 20 L. BAERT, J.-P. MAELFAIT and K. DESENDER Distribution - cfr. BAERT, MAELFAIT & DESENDER (1989); - FRANZ, 1975: FLOREANA: Post Office Bay (1 c : Fernandina, Floreana, Isabela (Alcedo, Darwin, Sierra V). SAN CRISTOBAL : Along road to El Progreso Negra), Pinta, Pinzon, Rabida, San Cristobal, Santa Cruz, (1 9 : V); Along road to Bahfa del Agua Dulce Santiago and Seymour Norte. (1 9 : V). SANTA CRUZ: Along road in Scalesia zone (1 c, 6 Other material examined Latrodectus apicalis Butler, 1877 - cfr. BAERT & MAELFAIT (1986): p. 115, Map 25; Material sampled in 1988 - LUBIN, 1982 (CDRS): SANTA CRUZ: La caseta (2 :I); c c FLOREANA (II): Locality 23 (1 9, 2 j 9 9), Locality - LuBIN, I983 (CDRS) : SANTIAGO : Los Guyabillos, 33 (1 9' j). southeast (1 c : III); El mirador, alt. 240 m (1 c : III). MARCHENA (III): Locality 51 (1 c, 3 9 9). ISABELA : Volcan Wolf (III) : Locality 69 U), Locality 7 5 Distribution (1 9, j). Volcan Darwin (III) : Locality 78 (1 9 ), Loca Isabela (Darwin, Sierra Negra, Wolf*), Santa Cruz and lity 90 (1 Coleosoma floridanum Banks, 1900 - cfr. BAERT & MAELFAIT (1986) : p. 117, Map 27; - cfr. BAERT, MAELFAIT & DESENDER (1989); Material sampled in 1988 - FRANZ, 1975 : FLOREANA : Post Office Bay (1 c, 1 Other material examined Distribution - cfr. BAERT & MAELFAIT (1986): p. 115, Map 26; San Cristobal and Santa Cruz. I I Araneae of the Galapagos 21 Theridion calycynatum Holmberg, 1876 FLOREANA (II): Locality 23 (1 o, 3 2 2). FERNANDINA (II): Locality 42 (1 o). Material sampled in 1988 GENOVESA (III) : Locality 52 (I 2 ). ISABELA: Voldm Wolf (III) : Locality 75 (2 Q Q ). Other material examined Volcan Darwin (III) : Locality 89 (3 o o, 5 Q Q, js), Locality 90 (2 o o, 1 Q, 6 j o o, 9 j Q Q). - cfr. BAERT & MAELFAIT (1986) : p. 119, Map 28; - cfr. BAERT, MAELFAIT & DESENDER (1989); Other material examined - FRANZ, 1975: ISABELA : Sierra Negra, Jaboncillo- wood, culture zone (1 o, 1 2 : VI). SANTA - cfr. BAERT & MAELFAIT (1986) : p. 117, Map 28; CRUZ : Along the road in Scalesia zone (1 o, 2 2 2, - cfr. BAERT, MAELFAIT & DESENDER (1989); js: V). - FRANZ, 1975: SANTIAGO: Forest east of "El Tunnel" (1 Q : VI). Distribution Distribution Fernandina*, Floreana, Genovesa*, lsabela (Darwin, Sierra Negra), San Cristobal and Santa Cruz. Fernandina, lsabela (Cerro Azul, Darwin*, Sierra Negra, Wolf*), Pinta, Santa Cruz and Santiago*. Tidarren sisyphoides ( walckenaer, 1841) Theridion coldeniae Baert & Maelfait, 1986 Material sampled in 1988 Material sampled in 1988 ISABELA : Volcan Wolf (III) : Locality 65 (3 2 2, 8 j), Locality 66 (3 2 2, 4 j 2 2 ), Locality 67 (1 o, j), SANTA CRUZ (IV): Locality 21 (2 Q Q). Locality 68 (I o, 1 2, 2 j). Volcan Da/win (III) : Loca MARCHEN A (III) : Locality 49 (1 o, 1 SA o, 2 Q Q), lity 78 (1 2 ), Locality 79 (2 2 2 ), Locality 80 Locality 51 (4 oo, 1 SA o, 3 Q Q). (3 o o, 2 2 2 ), Locality 81 (1 o, 1 SA o, 2 2 2, j), GENOVESA (III) : Locality 55 (1 2 ). Locality 89 (2 2 2, 7 j). ISABELA: Volcan Wolf (III): Locality 65 (1 o, 3 SA o o, 5 2 2 ), Locality 74 (2 o o, 1 SA o, 2 2 2, j), Other material examined Locality 75 (1 2, j). Volcdn Darwin (III) : Locality 78 (1 o, 3 2 2, 1 SA 2 ), Locality 80 (1 2 ), Loca - cfr. BAERT & MAELFAIT ( 1986) : p. 119; lity 83 (1 SA o), Locality 84 (1 o, 12 2 2, js), Locality - cfr. BAERT, MAELFAIT & DESENDER (1989). 85 (1 2 ), Locality 86 (1 o, 4 2 2 , 2 SA o o, 2 SA 2 2 ), Locality 87 (2 o o, 4 2 2, js), Locality 90 Distribution (2 0 0' 2 2 2 ). Fernandina, lsabela (Aicedo, Darwin, Wolf*), Santa Cruz Other material examined and Santiago. - cfr. BAERT & MAELFAIT (1986): p. 119, Map 28; - cfr. BAERT, MAELFAIT & DESENDER (1989); THOMISIDAE - FRANZ, 1975: FLOREANA : Post Office Bay (1 2 : V). SANTIAGO : Bucanero camp (2 o o, 5 2 2 : Misumenops inclusus (Banks, 1902) VI); forest east of "El Tunnel" (1 2 : VI). Material sampled in 1988 Distribution ISABELA : Voldm Darwin (III) : Locality 84 (j 2 ). Floreana*, Genovesa*, lsabela CAicedo, Cerro Azul, Dar- win, Sierra Negra, Wolf*), Marchena*, Pinta, Santa Cruz, Other material examined Santa Fe and Santiago. - cfr. BAERT & MAELFAIT (1986) : p. 119, Map 27; - cfr. BAERT, MAELFAIT & DESENDER (1989); Theridion rufipes Lucas, 1846 - FRANZ, 1975: SANTA CRUZ: Playa Tortuga (l 2 : VI). Material sampled in 1988 Distribution SANTA CRUZ : Locality 1 (1 2 : III), Locality 2 (2 2 2 : II; I o : III; 1 o : IV). Isabela (Alcedo, Darwin, Sierra Negra) and Santa Cruz. '' 22 L. BAERT, J.-P. MAELFAIT and K. DESENDER Tmarus stolzmanni Keyserling, 1892 Discussion Material sampled in 1988 The checklist published in BAERT et al.(l989) can be com pleted with six linyphiid species (BAERT, in press) and a FERNANDINA (III) : Locality 42 ( 1 ~ ). theridiid species Theridion strepitus described by PECK & MARCHENA (III): Locality 48 (l o). SHEAR ( 1987), all of them only known from the islands. Camillina cruz PLATNICK & SHADAB, 1982 has been syno Other material examined nymized with Camillina galapagoensis (B ANKS, 1902) by PLATN ICK & MuRPHY(l987), while the same authors des - cfr. BAERT & MAELFAIT (1986) : p. 119, Map 29; cribed a new species C.isabela.This brings the total number - LuBt N, 1982 (CDRS): SANTIAGO: Los Guyabillos, alt of known spiders for the archipelago to 80 of which 46 900 ft (l o, 2 ~ ~, 1 SA o, j: IV, V); are for the moment only known from the islands. A good - ZMO: FLOREANA: Post Office Bay (1 ~ : Xl.l925). deal of them might probably be through endemics, but it is still to early to generalize, as the spider fauna of the Distribution southern American mainland is not yet well known.The zoogeopgraphical affinities of the Galapagos spider fauna Fernandina*, Floreana, Isabel a (Darwin), Marchena*, with the American mainland is at the moment analyzed Santa Cruz and Santiago. and will be subject of a future publication. ULOBORIDAE Acknowledgements Zosis geniculatus (Olivier, 1789) This investigation was financed by (1) the Belgian Ministry of Education, (2) the National Foundation for Scientific Material sampled in 1988 Research (NFWO) and (3) the Leopold III Foundation. We are deeply grateful for the generous support of these Insti SANTA CRUZ (II) : Locality 3 (3 2 2 ). tutions. We received excellent cooperation from the Char les Darwin Research Station (Dir. Dr. Gi.inther RECK and Other material examined his staff), the Parque Nacional de Galapagos (lntendente lr. Humberto OcHOA and Lcdo Fausto CEPEDA). We wish - cfr. BAERT & MAELFAIT (1986): p. 122, Map 29; to thank the Ministerio de Agricultura y Ganaderia, Pro - cfr. BAERT, MAELFAIT & DESENDER (1989). grama Nacional Forestal for the permission they gave us to carry out our investigation programm. We also thank Distribution our field assistant Marfa GALARZA for her valuable help in the fieldwork, Dr. J. GR UBER of the Naturhistorisches Santa Cruz. Museum Wien for the loan of the material collected on the Galapagos islands by Prof. Dr. FRA NZ in 1975 and Patrick CH EVALIER for the drawing of the map. References BAERT, L. - 1987 - Spiders of the Galapagos Islands. Part IV . PLATNtCK, N. & MuRPHY, J. - 1987- Studies on Malagasy spiders, Miscellaneous families II. Bulletin de l'lnstitut royal des Sciences 3. The Zelotine Gnaphosidae (Araneae, Gnaphosidae), with a Naturelles de Belgique : Entomologie, 57 : 141-155. review of the genus Camillina. American Museum Novitates, 2874 : l-33. BAERT, L. - in press - Spiders of the Galapagos. Part Y. Linyphii dae. Bulletin of the Britisch Arachnological Society. L.BAERT, BAERT, L. & MAELFAIT, J.-P. - 1986 - A contribution to the Koninklijk Belgisch Instituut knowledge of the spider fauna of the Galapagos (Ecuador). Bulle voor Natuurwetenschappen, tin de I' Institut royal des Sciences Naturelles de Belgique : Ento Afdeling Entomologie, mologie, 56 : 93-123. Vautierst-raat 29, B-1040 Brussel (Belgium). BAERT, L. , MAELFA tT, J. -P. & DESE NDER, K. - 1989 - Results of the Belgian 1986-expedition : Araneae, and provisional checklist J.-P. MA ELFAIT, of the spiders of the Galapagos archipelago. Bulletin de I' fn stitut Wetenschappelijk medewerker K.B.I.N., royal des Sciences Naturelles de Belgique : Entomologie, 59 : 29- Afdeling Entomologie. 54. K. DES EN DER , PECK, S. & SH EAR, W. - 1987 - A new eyeless, stridulating Laboratorium voor Ekologie, Theridion spider fro m caves in the Galapagos islands (A raneae, K.L. Ledeganckstraat 35, Theridiidae). Canadian EntomologisT, 119 : 88 1-885 . B-9000 Gent (Belgium).