Internal Anatomy Hydrostatic Equilibrium

Total Page:16

File Type:pdf, Size:1020Kb

Internal Anatomy Hydrostatic Equilibrium Internal Anatomy Hydrostatic Equilibrium • Tissues are fairly dense • Freshwater 1.0, Sea water 1.025 • Fish components: – Muscle: 1.05 – cartilage: 1.1 – bone/bony scales: 2.0 – Fats: 0.9 • Average fish – 1.06-1.09 meaning they sink • Solutions • Note – more bone = much more weight! Shark Liver Oils Air bladder anatomy • Specific gravity of oils significantly less than water – Specific gravity of 0.86 – Expensive - up to 20% of body mass – Works at any depth Ex: 10 kg fish: 80% @ 1.091 20% @ 0.860 mean: 1.045 -Phsyclistic 1.025/1.045=0.98 -Physostomus Problems with air Bladders Bohr and Root Effects • Pressure problems – Boyles law – P1V1 = P2V2 Bohr effect – Root effect – – Slows speed at which you can change depths • Accessibility to atmosphere • Increase size of fish Other causes of shift: -Increased temperature -Salting out • Secondary losses Rete Mirabile – counter current exchanger Physoclistic Air Bladder Physiology Physostome Air Bladder Physiology Gasses in bladder at high Gasses in bladder at high pressure, exposure to capillary pressure, exposure to capillary Bladder lined with guanine bed will result in absorption bed will result in absorption crystals, impermeable to gas Atmosphere Vein Vein Sphincter controls Artery Sphincter controls gas exposure to oval gas exposure to oval Capillary bed Gas Gland Gas is pumped O2 CO2 release gland. gland. Rete regulated by into bladder metabolic activity in directly from Pneumatic gas gland atmosphere or Gas Gland duct Blood Blood supply through gas gland Passive secretion Regulated by exposure Active secretion to gland Stomach Requires energy Other uses for air bladders Respiration Dorsal View • Light reflection\ • Respiratory Surface Water – Gills – Stomach • Sound production – Lungs – Cutaneous • Body • Sound reception • Buccal cavity • Moving water – Buccal pump – Ram ventilators Gill anatomy Gill anatomy Filament Arch Rakers • Arch • Rakers • Filament • Primary Lamellae • Secondary lamellae Secondary lamellae Primary lamellae Gill Surface area to body weight Cutaneous Respiration • In water: • Problems with too much – aerial uptake (lungs) 40% surface area – skin 30% – gills 30% • In air: • Problems with too little surface area – lungs 63% – skin 37% – gills (opercula closed) 0% 1000 00 Polypteriformes Air breathing fishes Air breathing anatomy • Reduced gill surface area • Most freshwater – primarily swamps – low pH and oxygen • Few marine – Gobiidae – Bleniidae – Cottidae • 374 species, 125 genera, 49 families, 17 orders Obligate vs. facultative air breathers Gill Raker Function • Obligate vs. Facultative air breather • Environmental adaptations to • Raker spacing and size indicate feeding – low oxygen conditions mode – Predator refugia (swamps or deep water) • Large, widely spaced rakers 15 14-15 14 13-14 High BOD in 12-13 13 some habitats 11-12 12 10-11 adds to 9-10 11 • Small, fine rakers problems… 8-9 10 7-8 6-7 9 5-6 4-5 8 Dissolved Oxygen (mg/L) Oxygen Dissolved 7 6 5 4 0 5 10 15 Temperature (C) 20 t) 36000 p 25 p ( y 27000 it 30 n li a 35 18000 S 0 9000 Skulls and Jaws • External representations of major bones..
Recommended publications
  • BONY FISHES 602 Bony Fishes
    click for previous page BONY FISHES 602 Bony Fishes GENERAL REMARKS by K.E. Carpenter, Old Dominion University, Virginia, USA ony fishes constitute the bulk, by far, of both the diversity and total landings of marine organisms encoun- Btered in fisheries of the Western Central Atlantic.They are found in all macrofaunal marine and estuarine habitats and exhibit a lavish array of adaptations to these environments. This extreme diversity of form and taxa presents an exceptional challenge for identification. There are 30 orders and 269 families of bony fishes presented in this guide, representing all families known from the area. Each order and family presents a unique suite of taxonomic problems and relevant characters. The purpose of this preliminary section on technical terms and guide to orders and families is to serve as an introduction and initial identification guide to this taxonomic diversity. It should also serve as a general reference for those features most commonly used in identification of bony fishes throughout the remaining volumes. However, I cannot begin to introduce the many facets of fish biology relevant to understanding the diversity of fishes in a few pages. For this, the reader is directed to one of the several general texts on fish biology such as the ones by Bond (1996), Moyle and Cech (1996), and Helfman et al.(1997) listed below. A general introduction to the fisheries of bony fishes in this region is given in the introduction to these volumes. Taxonomic details relevant to a specific family are explained under each of the appropriate family sections. The classification of bony fishes continues to transform as our knowledge of their evolutionary relationships improves.
    [Show full text]
  • Assessing Turbine Passage Effects on Internal Fish Injury and Delayed
    Assessing turbine passage effects on internal fish injury and delayed mortality using X-ray imaging Melanie Mueller1, Katharina Sternecker2, Stefan Milz2 and Juergen Geist1 1 Aquatic Systems Biology Unit, Department of Ecology and Ecosystem Management, Technical University of Munich, Freising, Bavaria, Germany 2 Institute of Anatomy, Faculty of Medicine, LMU Munich, Munich, Bavaria, Germany ABSTRACT Knowledge on the extent and mechanisms of fish damage caused by hydropower facilities is important for the conservation of fish populations. Herein, we assessed the effects of hydropower turbine passage on internal fish injuries using X-ray technology. A total of 902 specimens from seven native European fish species were screened for 36 types of internal injuries and 86 external injuries evaluated with a previously published protocol. The applied systematic visual evaluation of X-ray images successfully detected skeletal injuries, swim bladder anomalies, emphysema, free intraperitoneal gas and hemorrhages. Injuries related to handling and to impacts of different parts of the hydropower structure could be clearly distinguished applying multivariate statistics and the data often explained delayed mortality within 96 h after turbine passage. The internal injuries could clearly be assigned to specific physical impacts resulting from turbine passage such as swim bladder rupture due to abrupt pressure change or fractures of skeletal parts due to blade-strike, fluid shear or severe turbulence. Generally, internal injuries were rarely depicted by external evaluation. For example, 29% of individuals with vertebral fractures did not present externally visible signs of severe injury. A combination of the external and internal injury evaluation allows quantifying and comparing fish injuries across sites, and can help to identify the technologies and operational procedures which minimize harm Submitted 10 February 2020 to fish in the context of assessing hydropower-related fish injuries as well as in Accepted 26 August 2020 fi Published 16 September 2020 assessing sh welfare.
    [Show full text]
  • Sascha Mario Michel Fässler Phd Thesis
    TARGET STRENGTH VARIABILITY IN ATLANTIC HERRING (CLUPEA HARENGUS) AND ITS EFFECT ON ACOUSTIC ABUNDANCE ESTIMATES Sascha Mario Michel Fässler A Thesis Submitted for the Degree of PhD at the University of St. Andrews 2010 Full metadata for this item is available in Research@StAndrews:FullText at: https://research-repository.st-andrews.ac.uk/ Please use this identifier to cite or link to this item: http://hdl.handle.net/10023/1703 This item is protected by original copyright This item is licensed under a Creative Commons License Target strength variability in Atlantic herring ( Clupea harengus ) and its effect on acoustic abundance estimates Sascha Mario Michel Fässler Submitted in partial fulfilment of the requirements for the degree of Doctor of Philosophy University of St Andrews September 2010 ii Target strength variability in Atlantic herring ( Clupea harengus ) and its effect on acoustic abundance estimates Sascha Mario Michel Fässler iii Declarations I, Sascha Mario Michel Fässler, hereby certify that this thesis, which is approximately 39’000 words in length, has been written by me, that it is the record of work carried out by me and that it has not been submitted in any previous application for a higher degree. I was admitted as a research student in October 2006 and as a candidate for the degree of PhD in October 2007; the higher study for which this is a record was carried out in the University of St Andrews between 2006 and 2009. date ……………...…signature of candidate …………………………….. I hereby certify that the candidate has fulfilled the conditions of the Resolution and Regulations appropriate for the degree of PhD in the University of St Andrews and that the candidate is qualified to submit this thesis in application for that degree.
    [Show full text]
  • Relationship Between Gill Raker Morphology and Feeding Habits of Hybrid Bigheaded Carps (Hypophthalmichthys Spp.)
    Knowledge and Management of Aquatic Ecosystems (2015) 416, 36 Knowledge & c I. Battonyai et al., published by EDP Sciences, 2015 Management of DOI: 10.1051/kmae/2015031 Aquatic Ecosystems www.kmae-journal.org Journal fully supported by Onema Relationship between gill raker morphology and feeding habits of hybrid bigheaded carps (Hypophthalmichthys spp.) I. Battonyai(1),,A.Specziár(1),Z.Vitál(1),A.Mozsár(1), J. Görgényi(2), G. Borics(2),L.G.Tóth(1),G.Boros(1) Received July 6, 2015 Revised October 26, 2015 Accepted October 27, 2015 ABSTRACT Key-words: Bigheaded carps and especially silver carp have been considered as an bigheaded carp, effective biological control for algal blooms, thus were introduced to sev- gill raker eral countries in the last decades, including Hungary. Our aim was to ex- morphology, plore the feeding habits of bigheaded carps in Lake Balaton (Hungary), filter-feeding, where the stock consists mainly of hybrids (silver carp × bighead carp). food size, We examined the relationship between filtering apparatus (gill raker) mor- plankton phology and size-distribution of planktonic organisms in the food. We failed to find any significant relationship between gill raker parameters and plankton composition in the filtered material. Bigheaded carps with vari- ous types of gill rakers consumed food within the same size-spectrum, independently of the rate of hybridization. However, the linkage between the proportion of different planktonic size classes in the water and in the diet of fish was detectable in case of both phytoplankton and zooplankton consumption, suggesting that the seasonally variable availability of differ- ent food items was an important factor in determining the food composi- tion of bigheaded carps.
    [Show full text]
  • Pictorial Guide to the Gill Arches of Gadids and Pleuronectids in The
    Alaska Fisheries Science Center National Marine Fisheries Service U.S. DEPARTMENT OF COMMERCE AFSC PROCESSED REPORT 91.15 Pictorial Guide to the G¡ll Arches of Gadids and Pleuronectids in the Eastern Bering Sea May 1991 This report does not const¡Ute a publicalion and is for lnformation only. All data herein are to be considered provisional. ERRATA NOTICE This document is being made available in .PDF format for the convenience of users; however, the accuracy and correctness of the document can only be certified as was presented in the original hard copy format. Inaccuracies in the OCR scanning process may influence text searches of the .PDF file. Light or faded ink in the original document may also affect the quality of the scanned document. Pictorial Guide to the ciII Arches of Gadids and Pleuronectids in the Eastern Beri-ng Sea Mei-Sun Yang Alaska Fisheries Science Center National Marine Fisheries Se:nrice, NoAÀ 7600 Sand Point Way NE, BIN C15700 Seattle, lÍA 98115-0070 May 1991 11I ABSTRÀCT The strrrctures of the gill arches of three gadids and ten pleuronectids were studied. The purPose of this study is, by using the picture of the gill arches and the pattern of the gi[- rakers, to help the identification of the gadids and pleuronectids found Ín the stomachs of marine fishes in the eastern Bering Sea. INTRODUCTION One purjose of the Fish Food Habits Prograrn of the Resource Ecology and FisherY Managenent Division (REF![) is to estimate predation removals of cornmercially inportant prey species by predatory fish (Livingston et al. 1986).
    [Show full text]
  • Poissons De La Cote Atlantique Du Canada 110180 12039370 C.1
    comprennent des Êperlans, des Perches, des Civelles et des jeunes de leur propre espèce. 482 En eau salée, la Perche blanche consomme de tout petits Poissons, des Crevettes, des Crabes et le frai de Poisson disponible. 49 Bien qu'elle soit modérément abondante, la Perche blanche n'est pas utilisée au Canada. Un lac de la Nouvelle-Êcosse, d'une superficie de 52 acres, renfermait plus de 23,000 Perches blanches. 435 Des prises de commerce de près de 2 millions de livres ont été faites annuellement dans la baie Chesapeake, où les pêcheurs à la ligne considèrent la Perche blanche comme un bon Poisson de sport. 4° Bar d'Amérique Striped bass Roccus saxatilis (Walbaum) 1792 AUTRES NOMS VULGAIRES: striper, rock, rockfish DIAGNOSE: Corps quelque peu allongé mais fort, la plus grande hauteur, sous le milieu de la dorsale épineuse, entre environ 4 fois dans la longueur totale, légèrement comprimé, pédoncule caudal fort. Tête entrant 4 fois dans la longueur totale, en pointe arrondie, légèrement comprimée, bouche terminale, mâchoire inférieure légèrement débordante, angle de la bouche sous le devant de dents petites, 2 plages parallèles sur la base de la langue, aussi présentes sur les mâchoires, le vomer et les os palatins; 2 épines faibles dirigées vers l'arrière sur la marge de chaque opercule, préopercule faiblement serratulé le long du bord. Diamètre de l'oeil entrant 8 fois dans la longueur de la tête. Nageoires: dorsales (2), 1", XIII—X, épines fortes réunies par une membrane, quatrième épine plus longue que les autres, entrant 21 fois dans la longueur de la tête, première et dernière épines très courtes, les autres de longueur intermédiaire, nageoire située au-dessus de l'extrémité de la pectorale, r dorsale, 10-13, premiers rayons égalant en longueur la plus grande épine, dé- croissant postérieurement à moins de la moitié du rayon le plus long, insérée à faible distance derrière la dorsale épineuse, base d'une longueur égale à celle de la dorsale épineuse et représen- tant les 3 de la longueur de la tête.
    [Show full text]
  • Fine Structure of the Gas Bladder of Alligator Gar, Atractosteus Spatula Ahmad Omar-Ali1, Wes Baumgartner2, Peter J
    www.symbiosisonline.org Symbiosis www.symbiosisonlinepublishing.com International Journal of Scientific Research in Environmental Science and Toxicology Research Article Open Access Fine Structure of the Gas Bladder of Alligator Gar, Atractosteus spatula Ahmad Omar-Ali1, Wes Baumgartner2, Peter J. Allen3, Lora Petrie-Hanson1* 1Department of Basic Sciences, College of Veterinary Medicine, Mississippi State University, Mississippi State, MS 39762, USA 2Department of Pathobiology and Population Medicine, College of Veterinary Medicine, Mississippi State University, Mississippi State, MS 39762, USA 3Department of Wildlife, Fisheries and Aquaculture, College of Forest Resources, Mississippi State University, Mississippi State, MS 39762, USA Received: 1 November, 2016; Accepted: 2 December, 2016 ; Published: 12 December, 2016 *Corresponding author: Lora Petrie-Hanson, Associate Professor, College of Veterinary Medicine, 240 Wise Center Drive PO Box 6100, Mississippi State, MS 39762, USA, Tel: +1-(601)-325-1291; Fax: +1-(662)325-1031; E-mail: [email protected] Lepisosteidae includes the genera Atractosteus and Lepisosteus. Abstract Atractosteus includes A. spatula (alligator gar), A. tristoechus Anthropogenic factors seriously affect water quality and (Cuban gar), and A. tropicus (tropical gar), while Lepisosteus includes L. oculatus (spotted gar), L. osseus (long nose gar), L. in the Mississippi River and the coastal estuaries. Alligator gar platostomus (short nose gar), and L. platyrhincus (Florida gar) (adversely affect fish )populations. inhabits these Agricultural waters andrun-off is impactedaccumulates by Atractosteus spatula [2,6,7]. Atractosteus are distinguishable from Lepisosteus by agricultural pollution, petrochemical contaminants and oil spills. shorter, more numerous gill rakers and a more prominent accessory organ. The gas bladder, or Air Breathing Organ (ABO) of second row of teeth in the upper jaw [2].
    [Show full text]
  • Gill Rakers of Basking Sharks (Lamniformes: Cetorhinidae) from the Tertiary of Sakhalin Island, Russia
    ZOOSYSTEMATICA ROSSICA, 23(2): 269–275 25 DECEMBER 2014 Gill rakers of basking sharks (Lamniformes: Cetorhinidae) from the Tertiary of Sakhalin Island, Russia Жаберные тычинки гигантских акул (Lamniformes: Cetorhinidae) из третичных отложений острова Сахалин, Россия M.V. NAZARKIN М.В. НАЗАРКИН M.V. Nazarkin, Zoological Institute, Russian Academy of Sciences, 1 Universitetskaya Emb., St Petersburg 199034, Russia. E-mail: [email protected] Isolated gill rakers of basking sharks (Lamniformes: Cetorhinidae) were collected from the upper Oligocene Holmsk Formation and Middle-Upper Miocene Kurasi Formation of the Sakhalin Island. This is the second finding of fossil basking shark remains in Russia. Fossil gill rakers are similar to those in recent representatives of the genus Cetorhinus, but differ from the latter by wider and shorter medial processes and higher bases. These features are more typi- cal for basking sharks in the fossil genus Keasius. Additional material is needed for the exact taxonomic identification of the Tertiary basking sharks from Sakhalin. Сообщается о находке отдельных жаберных тычинок гигантских акул (Lamniformes: Cetorhinidae) в верхнее-олигоценовых отложениях холмской свиты и средне-верхне-ми- оценовых отложениях курасийской свиты острова Сахалин. Это второе обнаружение ис- копаемых остатков гигантских акул на территории России. Рассматриваемые тычинки напоминают таковые современного рода Cetorhinus, но отличаются более короткими и широкими медиальными отростками и более высокими основаниями. Эти признаки бо- лее характерны для гигантских акул вымершего рода Keasius. Необходимы более много- численные дополнительные материалы для точной идентификации третичных гигант- ских акул Сахалина. Key words: Oligocene, Miocene, Sakhalin, fossil basking shark, Cetorhinidae Ключевые слова: олигоцен, миоцен, Сахалин, ископаемая гигантская акула, Cetorhinidae INTRODUCTION (Compagno, 2002).
    [Show full text]
  • RESEARCH ARTICLE Bottles As Models: Predicting the Effects of Varying Swimming Speed and Morphology on Size Selectivity and Filtering Efficiency in Fishes
    1643 The Journal of Experimental Biology 214, 1643-1654 © 2011. Published by The Company of Biologists Ltd doi:10.1242/jeb.048702 RESEARCH ARTICLE Bottles as models: predicting the effects of varying swimming speed and morphology on size selectivity and filtering efficiency in fishes E. W. Misty Paig-Tran1,*, Joseph J. Bizzarro2, James A. Strother3 and Adam P. Summers1 1Friday Harbor Laboratories, University of Washington, 620 University Road, Friday Harbor, WA 98250, USA, 2School of Aquatic and Fishery Sciences, University of Washington, Box 355020, Seattle, WA 98195-5020, USA and 3University of California Irvine, 321 Steinhaus, Irvine, CA 92697, USA *Author for correspondence ([email protected]) Accepted 30 January 2011 SUMMARY We created physical models based on the morphology of ram suspension-feeding fishes to better understand the roles morphology and swimming speed play in particle retention, size selectivity and filtration efficiency during feeding events. We varied the buccal length, flow speed and architecture of the gills slits, including the number, size, orientation and pore size/permeability, in our models. Models were placed in a recirculating flow tank with slightly negatively buoyant plankton-like particles (~20–2000m) collected at the simulated esophagus and gill rakers to locate the highest density of particle accumulation. Particles were captured through sieve filtration, direct interception and inertial impaction. Changing the number of gill slits resulted in a change in the filtration mechanism of particles from a bimodal filter, with very small (≤50m) and very large (>1000m) particles collected, to a filter that captured medium-sized particles (101–1000m). The number of particles collected on the gill rakers increased with flow speed and skewed the size distribution towards smaller particles (51–500m).
    [Show full text]
  • A Preliminary Assessment of Barotrauma Injuries and Acclimation Studies for Three Fish Species Final Report
    PNNL-24720 A Preliminary Assessment of Barotrauma Injuries and Acclimation Studies for Three Fish Species Final Report February 2016 RS Brown RW Walker JR Stephenson Prepared for Manitoba Hydro under an Interagency Agreement with the U.S. Department of Energy under Contract DE-AC05-76RL01830 PNNL-24720 A Preliminary Assessment of Barotrauma Injuries and Acclimation Studies for Three Fish Species Final Report RS Brown RW Walker JR Stephenson February 2016 Prepared for Manitoba Hydro, Winnipeg, Manitoba under an Interagency Agreement with the U.S. Department of Energy under Contract DE-AC05-76RL01830 Pacific Northwest National Laboratory Richland, Washington 99352 Summary Fish that pass hydro structures either through turbines, deep spill, or other deep pathways can experience rapid decreases in pressure that can result in barotrauma. In addition to the morphology and physiology of the fish’s swim bladder, the severity of barotrauma is directly related to the volume of undissolved gas in fish prior to rapid decompression and the lowest pressure the fish experience as they pass through hydro structures (termed the “nadir”). The volume of undissolved gas in fish is influenced by the depth of acclimation (the pressure at which the fish is neutrally buoyant); therefore, determining the depth at which fish are neutrally buoyant is a critical precursor to determining the relationship between pressure changes and injury or mortality. White Sturgeon (Acipenser transmontanus), Walleye (Sander vitreus), and Tiger Muskie (Esox lucius X E. masquinongy), were studied to better understand the likelihood of barotrauma when they pass through hydro structures. The pressure at which Walleye and Tiger Muskie were able to become neutrally buoyant was easily measured in pressure chambers as most fish hovered within the water column after being held overnight at a consistent acclimation pressure.
    [Show full text]
  • Comparison of Gill Raker Morphology of Alewife from Recent and Bygone Introductions
    Comparison of gill raker morphology of alewife from recent and bygone introductions Daniel Stich1 and Chase Ducey2 INTRODUCTION The alewife (Alosa pseudoharengus) is an anadromous species of herring native to North America, but along the east coast of the United States, landlocked alewife populations have become well established in inland lakes mostly due to fish stocking. Anadromous adults typically reach 10 to 12 inches in length, whereas adult size in landlocked populations is highly variable but tends to be much smaller (Palkovacs et al. 2007). A high temperature threshold allows them to comfortably live nearly anywhere and helps them survive as they move along streams and rivers to other bodies of water. A reproductive reaction to predation allows them to establish quickly in a new environment, which can sometimes interfere with the pre-existing ecosystem (Gibson and Myers 2003). Alewives are planktivores, eating large zooplankton and fry (juvenile fish that no longer feed off their own yolk sacs) by filter feeding, which utilizes gill rakers (bony or cartilaginous processes that project off the branchial arch, which supports the gills). As alewife populations grow, competition for food increases, and the abundance of large zooplankton decreases, resulting in the need for alewife to adapt to smaller types of zooplankton. The need to retain smaller food sizes during filter feeding may exert selective pressure on the morphology of gill rakers used in feeding (Palkovacset al. 2007). Landlocked alewife tend to have more gill rakers that are more closely spaced than fish of similar size in anadromous populations (Palkovacs et al. 2007), and the number and spacing of gill rakers in landlocked population is correlated with prey sizes (Palkovacs and Post 2009).
    [Show full text]
  • Gill Rakers and Teeth of Three Pleuronectiform Species (Teleostei) of the Baltic Sea: a Microichthyological Approach
    Estonian Journal of Earth Sciences, 2017, 66, 1, 21–46 https://doi.org/10.3176/earth.2017.01 Gill rakers and teeth of three pleuronectiform species (Teleostei) of the Baltic Sea: a microichthyological approach Tiiu Märssa, Mark V. H. Wilsonb, Toomas Saata and Heli Špileva a Estonian Marine Institute, University of Tartu, Mäealuse St. 14, 12618 Tallinn, Estonia; [email protected], [email protected], [email protected] b Department of Biological Sciences and Laboratory for Vertebrate Paleontology, University of Alberta, Edmonton, Alberta T6G 2E9, Canada, and Department of Biology, Loyola University Chicago, Chicago, Illinois, USA; [email protected] Received 16 September 2016, accepted 14 November 2016 Abstract. In this microichthyological study the teeth and bony cores of gill rakers of three pleuronectiform species [European plaice Pleuronectes platessa Linnaeus, 1758 and European flounder Platichthys flesus trachurus (Duncer, 1892), both in the Pleuronectidae, and turbot Scophthalmus maximus (Linnaeus, 1758) in the Scophthalmidae] of the Baltic Sea are SEM imaged, described and compared for the first time. The shape and number of teeth in jaws and on pharyngeal tooth plates as well as the shape, size and number of the bony cores of gill rakers in these taxa differ. The European plaice and European flounder carry incisiform teeth anteriorly in their jaws and smoothly rounded, molariform teeth on pharyngeal tooth plates; the teeth of the plaice are more robust. The gill rakers have similar gross morphology, occurring as separate conical thornlets on gill arches. The bony cores of these thornlets (rakers) consist of vertical ribs with connective segments between them.
    [Show full text]