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Classification Des N Omiinae De La Region Orientale, De N Ouvelle-Guinee Et Des Iles De L'ocean Pacifique (Hymenoptera: Apoidea: Halictidae)

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Classification Des N Omiinae De La Region Orientale, De N Ouvelle-Guinee Et Des Iles De L'ocean Pacifique (Hymenoptera: Apoidea: Halictidae) I I BULLETIN DE L'INSTITUT ROYAL DES SCIENCES NATURELLES DE BELGIQUE ENTOMOLOGIE, 79: 151-229,2009 BULLETIN VAN HET KONINKLIJK BELGISCH INSTITUUT VOOR NATUURWETENSCHAPPEN ENTOMOLOGIE, 79: 151-229,2009 Classification des N omiinae de la Region Orientale, de N ouvelle-Guinee et des iles de l'Ocean Pacifique (Hymenoptera: Apoidea: Halictidae) par Alain PAULY Summary australiennes n' ont pas encore ete abordees, sauf lorsque leur distribution atteint la Nouvelle-Guinee eta The 282 names of species from the Oriental region, New Guinea !'exception du vaste genre Austronomia. and Pacific islands are sorted by modem genera. At least 136 valid Presque tous les types ont ete examines, les species are recognized in Oriental region and 49 species in New Guinea and Pacific islands. Keys of species are proposed for some descriptions originates etant bien souvent insuffisantes genera. ou trop imprecises pour reconnaitre les especes ou 50 new species are described: Gnathonomia boyolali, G. les placer dans un genre modeme. Des lectotypes ou cambodiana, G. mayoyaoensis, G. radiata, G. timorensis, G. neotypes ont ete designes lorsque necessaire afin de wahisi, Maculonomia sanguinea, M soekaboemiensis, M tigeri, Curvinomia submirifica, Pachynomia nathani, Steganomus liefiincki, stabiliser la nomenclature. Lipotriches aurodigitata, L. gracilis, L. notoxantha, L. postcarinata, Plusieurs groupes consideres comme sous-genre L. tubuliseta, L. vietnamensis, Macronomia anamalaiensis, M ou synonymes par MICHENER (2007) dans un esprit de angkorensis, M dilatata, M karnatakaensis, M madrasensis, regroupement sont consideres ici comme des genres a M nilgiriensis, Morbitarsis, M savannakheti, M walayarensis, part entiere. Un des arguments est la coherence entre Maynenomia chayaphumensis, M ghatensis, M indochinensis, M keralaensis, M lonavlaensis, M nathani, M packeri, M poonaensis, les genres reconnus par PAULY (1990) de la Region Austronomia arcuata, A. atromellea, A. bismarcki, A. browni, A. Afrotropicale et ceux de la Region Orientale. Les djampangi, A. gressitti, A. guadalcanalensis, A. hollandiae, A. nouvelles combinaisons ne sont pas indiquees car laminatrochanter, A. nivea, A. pilitrochanter, A. pseudoscutellata, a considerer comme provisoires jusqu'a ce qu'une A. sancristobali, A. solomonensis, A. tapinensis. analyse moleculaire pennette de mieux cemer les limites des genres et tester la validite des caracteres morphologiques utilises. Par exemple, la presence de Introduction bandes tegumentaires emaillees aux tergites est bien pratique pour distinguer le genre Nomia sensu MICHENER, Ce travail presente pour la premiere fois une tentative mais n' est pas necessairement un bon caractere pour de classification des Halictidae de la sous-famille des determiner un groupe monophylletique. La presence de Nomiinae de la Region Orientale. Ce n'est done pas une larges tegulae qui caracterise les Pseudapis est partagee monographie incluant la redescription et la distribution en Afrique par des especes appartenant a des genres de toutes les especes ni un catalogue incluant toutes aussi differents que les Steganomus, Lipotriches ou les citations dans la litterature. Ces renseignements Austronomia. Un caractere plus discret mais qui semble figureront dans un second temps dans des publications detenninant pour separer les genres semble bien etre la consacrees en detail a chaque sous-genre. conformation ouverte ou fermee du plateau basal des Nous avons inclus ici laliste des especes de Nouvelle­ tibias posterieurs des femelles. A noter que !'utilisation Guinee et des lles de l'Ocean Pacifique. Au total, nous de ce demier caractere rend parfois le classement des avons classe pas moins de 282 noms d'especes et decrit males, dont la presence d'un plateau tibial est plut6t 50 especes nouvelles. Pas moins de 136 especes sont aleatoire, complique en l'absence de femelles. reconnues comme valides pour la Region Orientale Dans les descriptions des especes nouvelles, les tandisque 49 sont reconnues pour la Nouvelle-Guinee caracteres sont repartis entre coloration et structure, et les lles du Pacifique, la distribution de la sous-famille puis generalement cites dans 1' ordre de leur importance s'etendant jusqu'en Nouvelle-Caledonie. Les especes pour faciliter la reconnaissance des especes et non I I 152 A. PAULY dans I' ordre morphologique. Des cles proviso ires sont NMV = Naturhistorisches Museum, Vienna, proposees pour la reconnaissance de la plupart des Autriche (M. VIZEK; M. MADL) . genres. L'espace reserve aux illustrations etant limite OOL = Oberosterreichs Landesmuseum, Linz, ici a 8 planches couleur permettant la reconnaissance Autriche (F. GusENLEITNER; M. ScHWARZ). des genres, !'illustration photographique des caracteres OUMNH = Oxford University Museum of Natural speci:fiques seront visibles en attendant publication History, Oxford (anciennement Oxford dans les articles subsequents sur le site web: http:// university Museum), UK. (J. HoGAN) www.alashymenoptera.net/Hakictidae.htm RMNH = Rijksmuseum van Natuurlijke Historie, Leiden, Pays-Bas (C. VAN ACHTERBERG; R. Acronymes des collections etudiees (entre parentheses DE VRIES). le nom des personnes ayant aide au pret de materiel): SDEI = Senckenberg Deutsches Entomologisches Institut, Eberswalde, Allemagne (H. AMNH =American Museum of Natural History, DATHE; A. TAEGER; C. KUTZCHER). New York, USA (J. S. AscHER; E. L. SMUK = University of Kansas, Lawrence, Kansas, QUINTER). USA (C.D. MICHENER; M.S. ENGEL). BBMH = Bishop Museum, Honolulu, Hawai, USA TIGER = project: Hymenoptera Institute, University (T. GONSALVES). of Kentucky, Lexington, USA (S. CLUTTS BMNH =Natural History Museum, London, UK & M. SHARKEY viaL. PACKER). [anciennement British Museum (Natural UP = University of Peradeniya, Sri Lanka History)] (G. ELsE; D. NoTION). (J. EDIRISINGHE; P. KARUNARATNE; W.A. CAS = California Academy of Sciences, San INOKA). Francisco, USA (W.J. PULAWSKI). USNM = National Museum of Natural History, FSAG = Faculte Universitaire des Sciences Smithsonian Institution, Washington D.C., Agronomiques, Gembloux, Belgique (E. USA. (B. BARrus) HAUBRUGE). UZMK = Universitets Zoologiske Museum, HNM = Magyar Nemzeti Museum, Budapest, Copenhagen, Danmark (L. VrLHELMSEN). Hongrie. Baldock David., collection privee (UK.) HYAS = Entomological Laboratory, Hyogo University of Agriculture, Sesayama, Japan. IRSNB = Institut royal des Sciences naturelles Cle des genres de Nomiinae de Ia Region Orientale et de Belgique, Bmxelles, Belgique (P. de Nouvelle-Guinee GROOTAERT; J.L. BOEVE; J. CONSTANT). ITZA = Instituut voor Taxonomische Zoologie, (1) Tergites avec des bandes apicales tegumentaires Amsterdam, Pays-Bas (W. HoGENEs; G. .. .............. ....... ... ... .. .... .. .. ... ..... ...... .... ..... (2) SCHULTEN) .. - Tergites sans bandes apicales tegumentaires (seulement IZB = Institute of Zoology, Academia Sinica, parfois des bandes pubescentes) ..................... (8) Beijing, Chine. IZK = Institute of Zoology, Academi Sinica, (2) Metanotum avec tme double projection .. ...... (3) Kunming, Chine. - Metanotum sans double projection ....... .... ..... .. (4) KU = Kyushu University (0. TADAUCHI; Y. HIRASHIMA) . (3) Metanotum et scutellum avec des projections ........ LACM = Los Angeles County Museum of natural ........ ... ........... ...... ...... .... ... ....... Crocisaspidia History, Los Angeles, California, USA - Seulement le metanotum avec une projection bi:fide ... (R.R. SNELLING). ....................... ..... .... .. ....... ..... .Hoplono1nia MCSNG = Museo Civico di Storia Naturale, Genova, ltalie (F. PENATI). (4) Femelles avec le plateau basal des tibias posterieurs MNHNP = Museum National d 'Histoire N aturelle, complet ....... ... ... .. .. .. .. ... ....................... .... (5) Paris, France (C. VILLEMANT). - Femelles avec le plateau basal des tibias posterieurs MNHUB = Museum fur Naturkunde an der Humboldt incomplet .. ..... ................... ....... ........... ... (7) Universitat zu Berlin, Allemagne (F. KocH). (5) Tegulae ovales, plus petits. Milles avec des II Classification des Nomiinae 153 mandibules souvent tres developpees ou presentant un (13) Metanotum avec un tubercule median ou une crochet sur le bord interne ............... Gnathonomia projection. Mandibules des femelles tridentees. Calcar - Tegulae «auriforme», la partie apicale etant retrecie, interne des tibias posterieurs des femelles avec deux plus grands. Males a mandibules simples ............ (6) grandes dents ................................. .. .Mellitidia - Metanotum fortement convexe, deprime au milieu, sans (6) Femelles: calcar externe des tibias posterieurs avec structure mediane. Mandibules des femelles bidentees. une forte dent subapicale et perpendiculaire. Plateau Calcar interne des tibias posterieurs des femelles avec basal des tibias posterieurs borde par une carene en de petites dents ciliees et serrees ........ .Ptilonomia forme d'ecusson, le centre largement pilifere. Deux sexes: marge apicale de l'aile anterieure plus ou moins (14) Ocelles fortement developpes. Grandes especes de enfumee sur toute sa largeur. Ponctuation du sctum couleur testacee, parfois totalement noires. Nouvelle- generalement plus enfoncee et double. Males: pattes Gurnee et lies voisines. .......................Reepenia posterieures generalement plus epaisses et plus courtes - Ocelles peu developpes ou bien corps plus petit ou bien ................................................... Curvino1nia autre distribution ...... ......................... ........ (15)
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