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6I u86t)zt 6t:it)9?rsrInN NuytsiaVol. No. 1 (19871 20 6,

3cm 5mm Fi.Jne L Acaciaalcockt: A - Floweringbranchlet. B to D Phyllodevariation E - !9gfrye (mltv:e):| . L€"sume(immature). G - See.dshowing light red-brownfunicle encircling seed in a double{old. A frolllC.R. Alcock lbI2:BfromCR AlcarAl256tctrcmJ.ll. u tiRle)\N At68 7045; Dfrom C.R.41rc,k lo56:EfromcR,4/,o'( 1682:F {rom ,tri.Hollidav s.n.lAD 98422719514Cfrom E.M. Canning5390 and S Corbett Bushysftrub to c. 3 m tall, often suckering(especially if rootsare disturbed).,Balk mostly smooth,grey or brown at baseof trunks,reddish on young branches.Branchlets terete but slightly angledat extremities,very finely nerued,straight to very slightly flexuose,rather slender,glabrous, sometimes marked with prominent,distant, raisedleaf baseswhere phyllodes havefallen. Stipules pemistent, triangular to deltate,minute, c.0.5 mm long.Pft yllodes usually narrowlyelliptic to oblanceolate,slightly asymmetric, gradually narrowed towards the base, attenuateor abruptlynarrowed to acuteor obtuseapices, mucro usually present and uncinate, mostly6'9(10) cm longand 8'21(30)mm widebut frequentlyinterspersed with a few smaller phyllodes(4-5 cm x 5-8 mm), l:w : (2)3-10,straight to shallowlyrecurved, very thinly coriaceous,flexible, glabrous, dark greenfmidrib nol oYerlyprominent but slightly raisedwhen dry, usuallyyellowish to light brown, centralto slightly excentric'lateralnerves mther obscure, divergingfrom midrib at an acuteangle, sparsely and openly longitudinallyanastomosing; marginalnerves narrow, yellowishto light brown; pulvinus 2-6mm long, drying finely wrinkled anddark brown or reddishbrown. G/rndsituated on uppermargin of the phyllode{3)8 l2(20) mm abovethe pulvinus,circular or oblong,0.3-0.6mm long, sometimesvery slightlyraised abovethe marginwhich is sometimesvery slightlyindented at the gland,orifice shallow but distinct.Inflolescences l(2) per node,racemose, very rarely interspersedwith somesimple axillary peduncles,racemes sometimes paniculately arranged. Racemes (1.512'4(5) cm long with {2)5'11heads. the lowermosthead inserted 8-18 mm abovethe baseiraceme axes straight to veryslightly flexuose, glabrous, very finely wrinkledwhen dry, baseebracteale. Peduncles '^eg norssnlJeou pue e^oJ,{Jouew pu€ lurod lsoiA ruoJJpeliolloc s; 'uloJur.I uooqs?q lr lrBd aqt Jo .{l[uortxa uroqtnoseql uorC uod leeu lulod lqBI.I ,(lllg 1r le pue IJed leuorl€Nulorurl oql urqlr,r oIe'I enlrd pue elnsuruaduol8uruuo( edeJ llrH al proJrrl€ls1€ Jnrco suortelndoduJeqlJou oqJ lred€ uI 0Z lnoqEs?ere o,rt rxo{ polcello, ueeqs?q sarcods oql eroq,r?lFllsny qlnos'?lnsuruedar g lspe-qtnosoruoJlxg uounEnsle l1 pl '(pBolq,{llEullouqe (c sopoll^qd pl OV) Si9l 991y76 {a13u14/11't'e^oJ ^roual,{ pre,t\oluloour-I lrod urorJ[rul gt] rul 'Zosgtt :(HJUAd tB sdnp ot u s uoadntJ puBtrl pue 0€ 66t9bt't'n'E SNVJ) 's J trrA t J \8It gNvl) u s,{.laryrr4 puorleNulorur.I u '?psuruod6rt, L0E6rt. ..tt,00 09fI ,9t ott.'ryed'turod uolEuruocod€J:(Cy) uotu{S lrod lq8r-I ^llrg l?oN ^ '.G6tLZZV86 t699 ?'O'ulorurl (7y) us t'putoH )'t86t 1IH proJruBrs:(0600t28 pue J; 'JINE'naqtoS 1800tZ8CSt) 06tS .: pue /8t9 g pueSutuuoj Wg'\rvrt e^oJ froruot{ SuoFulorur.I trod uo{rulZt o'lr?d ulocurT:())(09€? ou roU),Mol€r'}r'o^oJ ^Jourol,^l 'S.0tIBUoTIBN 'lllH '{r8d lerlsnv :4,,0€ ,9SoS .9t ott proJruEts IBuortBNulorurl'.(Ay) 6LZt )lrutv ,{ 'y, '{r?d ulosurl'?ar? proJur?ls:(clv) )poqy s: '.eorel€uor}eN IIrH 8/z€ a'J'{r?d leuorlBN ulmul'I iutod tsoiA :(CV) t99Z ryoTv tr?'lllrH apnqtroN rsou] ulocur-IUod Jo Ji gss url t c'{00rJ 1l?s:(cY) z89l lrulv tr]'ulorur.I lrod Jo gss [onp]u{ 0t lnoq? ll 'lurod 'ulocurl tseld :(TAI,{'(V) 999I lroqv yJ trod Jo iASSu.r{ 0f c'Ued IeuoItBN ,{ 'lurod ulocurl lsel[ :(HJUgd'(JV) 99ZI )trntv tr )'(llrH proJuerspu? urr?J srapulld 'ulotul'I 'a^rased uoa,{}oq Uod JoS rul SI t srr{cu{,,|\) ,,sel€'I o{[d,, :((V) TZZI\roqy tr J I 'o^oJ 1 ?un?C pu? erolC ulo.ur-I ,{roruol\ :vI'IvUJSOv HJ.}OS paauDxa suaw0ads

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Affinities. Acacia alcockii is placed in sectlonPhyllodineae DC. on account of its globular flower-headsand 1-nervedphyllodes. Taxonomically the new speciesis placedbetween.'4' ancepslXJ. andA. leiophyilaBenth. from whtch it can be distinguishedby a combination of its racemoseinflorescinces and transverse seeds (see key under,4.cnceps). All threespecies occurwithin the . From dataderived from comparativemorphology it is possiblethal A. alcockiiis of hybrid origin. However,it seemsthat the northernand southernpopulations of the speciesmay have had different origins

Thesetwo setsof populationscan generallybe recognizedby the following characten'

Northernpopulations: phyttodes 8'11(14) mm wide,l:w : 6-10,usually shallowly recurued (sometimesstraight); Iegumes to 7-9cm long,8-12mm wide,thinly to moderatelycoriaceous, itraighfedgedoi slightlyto moderatelyconstricted between the seedsalthough random deep constdctionsdo occur.

Southernpopulatio ns phyllodest 2-21(30)mm wide,l:w : (2)3-7,usually straight (rarely shallowlyreiurved); legumes to 3.5-5cm long, 12-17mm wide, moderatelycoriaceous to slightlyirustaceous, oclasionally more or lesssub'woody, straight edged (occasionally few or many deepconstrictions on somelegumes).

Detailedfield studiesof threeof the northernpopulations were recently conducted by J' Briggsand F. Vickery. Thesepopulations are locatedat , StamfordHill (which coi-prisesseveral hundred plants) and Cape DonningtonPeninsula. Briggs (pers. comm ) ."ported thut plants from thesepopulations, do not show the rangeof morphological_variatio,n that one would expectfrom hybrid swarms.Furthermore, n eitherA. ancepsnor A leiophylk occurwithin seveialkilometres (? at least15 km) of thesepopulations. Nine seedlingsraised by Briggsfrom seedcollected from thesepopulations showed no apparentmorphological ,"gr.gitlon. Seedlingsof l. ancepsand A. leiophyllgwere not examined.Pollen from the (Stamford foitoriing 15 specimenswas examlned, Alcock 1256(Pillie Lake), Bnggs 121 1 Eill), Vickery\.n. i,cNg :,lg:ol to 349318(Cape Donnington Peninsula) and Svmon 6693 (Billv Light Foint). The only abnormalitieswere seen in the polyadsof Symon6693 where most weleeither asymmetric in shapeor possessedgrains with irregular,vestigial or no cytoplasm.

Comparablefield studieshave not beenconducted on the southempopulations. However, Briggsfias visited Memory Covewhere he observedA. ancepsand A leiophyllabt not A' alciikii. V ickery,who knowsthe southernLincoln National Parkwell, reported(via Briggs) that he has neverseen anything resemblingA. alcockii in the alea. Theseobsewations suggest that the plantswhich arehere attributed to A. alcockiilrom Memory Coveand WestPoint arenot c,ommonand may indeedsimply be spontaneous hybrids (probably between A' anceps andA. teiophylk). The pollenof ca[n ing 5390and Corbeft.howev€r. showed no iregulalities. The other-specimensfrom thesesouthern areas were either in fruit or very young bud' r

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Hybridity. Jndgtngprincipallyfrom morphologicalevidence derived from herbariumspecimens it appearsas thoughA. ancepshybridizes with a numberof relatedtaxa. Only rarelywere we ableto examinethese taxa in the field. Our purposehere is to draw attentionto these putative hybridsand thus lay a foundationfor future studies.

L A. ancepsx nematophylla.See A. nematophyl/abelow. 2. ? A. ancepsx leiophylla.See A. alcockii above. 3. ? A. ancepsx notabilis.

Somespecimens from StreakyBay and FowlersBay (northernEyre Peninsula)possess charactemwhich suggest they are somewhat intermediate between A. ancepsand A. notabilis. Many of thesespecimens have previously been determined as l. notabilisat herb.AD and mappedas such by Whibley(1980: 122). Their coarselyangular branchlets, thickly coriaceous phyllodes,golden fimbriolate bracteole laminae and calyx lobesand more or lesswoody legumes clearlyrelate them Io A. anceps.However, a few characterssuggest intergradation with -4. notabilis,viz. inflorcscencesa mixture of axillary pedunclesand racemes,phyllodes occasionally with 2 glandsand./or sub-attenuate. Although l. ancepsocciJrs in the samegeneral area as theseputative hybrids, current collections show A. notabilisto be distributedconsiderably fudher east.Specimens referable to this possible80i72 hybrid are C.R. Alcock 3395 (AD), N. Hall 80i72 (MEL, PERTH), A.E. Orchard3142, (AD), Veitchs.n. (AD 96716196)- all StreakyBay area,A.E.Orchard 3144 (AD) - FowlersBay. Specimens with oblanc€olate, obtusephyllodes superficially resemble ,4. alcockii lsoulhernEyre Peninsula- seeabove) but are distinguishedby their largerflower-heads (50-60'flowered), golden fimbriolate bracteoles and more or lesswoody legumes. Key to A, ancepsand someallied taxa (includingputative hybrids) 1a. Flowering pedunclesnever racemosely arranged; legumes mriaceous crustaceous to woody...... 2 b. Floweringpeduncles all or somearranged in racemesof 1(2)or more flower- heads;legumes as aboveor firmly chartaceous ...... 3 2a. Phyllodes0.7- 1.5 mm wide,flat or tercte,never continuous with branchlets; Ilowers28-41 per head;legumes5-6 mm wide;seed longitudinal (S.A., not colnmon.)...... A.nematoohvlla b. Phyllodesl-3 cm wide,flat, sometimesmntinuous; flowen 60-130per head; legumes10-15 mm wide;seeds transverse (W.A., S.A.)...... -...... A. anceps c. Someor all abovecharacten intermediate (S.A., rare) A. ancepsx nematophya 3a. Phyllodes1-5 mm wide, 3,10 cm long, flat or terete;racemes wilh (l)2-4(13) flower-heads; flowers 25,40 per head; legumes 3-6 mm wide, coflaceous to crustaceous;seeds longitudinal (S.A., N.S.W., vic.)...... A.caramiforia b. Phyllodesall or mostlymore than 5 mm wide,flal...... 4 4a. Legumes5-6 mm wide,firmly chartaceousto slightlycoriaceous; seedslongitudinal; bracteole laminae c. I mm diam., golden, fimbriolate;phyllodes falcate, thinly coriaceous,l0-13 cm long and 15-22mm wide (S.A.)...... A. Ieionhvlla 'uerp 'pesr?r 'I urx 9'0 c [pq311sf,lpuorszcco s^oqeuru t o] snur^lnd puo le ']uouruord Jo ',{lpIetsp epofl,tqdaql Jo urAJEruJoddn uo polenirs ^ro^ lou prrlg osoumldua]Jo ueq,r 'Euol 'e!er4 polluu,{r.pup u,rorq {l?p ot tq8rT ulru Z-g l snumfld'.3usoruolseue fluodo'el8u€ '^lp 'snoJounu elnce,tue^B IB quprruoql rxoq Sur8lo^tp ueq^\luapr,ls ,4lqeuosee: lnq eurJ 'pecunouoJd se^rouIerolel :(u.{oJq}q8IT 8ur^Jp ueuo) i olle,{ ,{lqeuos?alsattau putztout :(u,^.{oJqtq8rt 8ur,{rpuelJo q8noqlp) ,rollo,{ uo$o',{Jp ueq,r pest?l,{llq8rls 'pmunouold 'crJtuotxa ^lqeuos?or ^llq8lls fre^ ol l?Jluoc q]ptw :?unot uoq,r{snocnelS tnq uaoJt 'unrpsur qsr.{oJ8'snorq€I8 'p?orq ueq^\ ,tl?rcadso ot?lnpun ,{[q8rTs 'po^lnrur-ol?ncl€ ^lelel '}JeJo eJou ro pe^JnoeJ-elenrr?^ltq8rJs , Jo^,roJ E ,{lleuorsecaoq8noqlp lq8r?qs,{llensn 01 'pop,i '^Jp SupusJs? oJcJoqlBJ ueq,r pe11uu,r[1aug ,{:e.r.sourlouros 'snoooeum fl3l?Jepouj ol 'uJru ,{Iuqt'S Z ol,/(eJe tnq 6-9t ^Fnsn orlerq1pl. ol qfuel Surqr"al^roJ ? ,{ll?uorse3.o '8uol 'oJcnu St 'olnulu lnq opr,r ruru ^llensn rur (l oleuDunto lq8r?Jts ? qlr,{ 'gsnlqo 8l-6 I)01,/(9)'J?ourt pu? po,rorreu flldnJq? reql?r serd? ^peolq fllEuorsemoouos lnq oleloJouBlqo 'olnulru 'Jeln8ueul ,tl,rouEu 01 rqd{lo tlr orJeusapo ttld Suotrutu g 0 c salndlrs qsrppal sioot4sMaN UalleJe^Eq sopoflfqd oJoq,r seqcu€lq alnl?u uo lueprlo eJeqcrqia sos?q 'luelslp 'osournjd J?el posr€Jlueulruojd qtlr pa{J?rll ,{lsnoncrdsuoo'snoJq?[E 'srJde :raq1 sp:e,no1 Jeln8u€^lluau[uoJd 01 fllq8rTs lnq eloJol'pa^lou ^lourJ sJalq)uDJg 's\unq slJ€dlo,rol uo 'qiootus Jo u,\{orq,peJ01 ,{er8 Zqg soqcueJqsnoFpued-qns ot Surqcjesourleuos aql Jo spuaoqt spJ?,rolpoleJlueJuoc sepoll^qd'oseq tuo4 SurJo{onssorunouos 'lepuels s{unJl ',{qsnq flolel 'f133?4s 'll?t 'fpurds puBuodo su,roJo u (S)t-g'€aa0 llolus ro qruqsWruuols-of8urs ^[msn

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'tr3lqlqA aoaulpolltqJuogxes Dlrrrtl uerl?llsnv qtnos A'I C T urts€l^trU g 28 NuytsiaVol.6, No. | (19871

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A Figute.3.Acacia crcracer. A - Floweringbranchlet. B - Legume.C Seedshowing reddish brown funicle % encjrclingseed. A [ron M. &mmons 1722;. B ftotl D.J.E. Whibtey7 3161 C lrcm B. Coptey3898.

Iryflorescencesl-2 p9l,nod9,very fragrant,racemose, rarely interspersedwith a few simple WdLrncles.Racemes (l)2-4(6) cm long,with (3)5-9(14,20:B. Copley,pers. comm.)heads; raceme axes.straightor slighflyflexuose, somewhat angular, glabrous, pruinose, sometimes growing out' baseebracteate. Peduncles 4 I l(17)mm long,mostly alternate but the lower2 sometimei opposite,,finelylongitudinally sulcatewhen dry, glabrous,pruinose; receptacle c apitatelbasal peduncularbract l, persistent,ovate to narrowiy ovate,b.5-l mm long, glabrous.Flower- headsglobtl1ar,4-5 mm diam.when dry, to l0 mm diam.when fresh, lemon yellow to golden yellow'with.34-45155) densely arranged flowers, the budsconspicuously pruinose. Braiteores sub-peltate,1.2,1.5 mm long,glabrous; c/aws narrowly linear, glabrousilaminae observable betweenflowers but not overly prominentin the bud, not thiakened,more or lesscircular and usually minutely apiculate, pruinose. F/owers 5 merous, glabrous.Calyx % lhe length of corolla,gamosepalous, divided for c. % its lengthinto oblong,rounded iobes; calyx tribe 'sarunEol esoumld(,{nq8lls ,{luo Jo)lou slr pu? '(8uol ,Jouoqs,epo[^qd saloepErqetBlorjquu'{cqq ot u,rolq UBp urx S-t)solrunpod lalnols 'V .U radspuBIE t-t(Z) sll ,iq DaJoptJ ruog paqsrn8uqsrps u,wpaa$ ,re^o,r\oH.sornt?eJ 'opollfqd Iecl8olodJ?cpuB murJsalouu letaueEpue slotqcu8Jqesourntd slr ul solcods,reu aql salquesoJ(BrT?"qsnV ujolserl\) ,{lo{?lg pue ueplP.lNttuDwpaals o!?oJV .speos osle^SuEtl ,sopoll,{qd qllr\ (apl,laruru ZI-8) soun8elpeoJq pue s]elqcu?Jqosoutnrd,uou snoec?rrorstr ^q pazuSocoJ^lrpeol sr (o l?du^s lou am o,{\1oql lnq) oaJDDtJ V se ee:eIe:aueE eues eql rn s,ro$ qcrqa slllqopu oporv )soutnld lou alp qJrq,{\(rur gl o, sorun8alJo8uol pu? (p0re,!tou'Zr-[t)speoq-reaou :e6:e1 'pe,trnc ,{l,ro[eqs pu? ue3lE-pluorB qcrqa ({uc 'slolqJuelq l,l-t) sepoll,{qdre8uol ^lpnsn osoumld-uou'osonxeU ^lluoururoJd str ^q peqsln8ultsrp ^lrpEorsr rlcLq/f.4113 polnqulsp ,tlloqlnos etorx oql ol l?lrr s sr ql,rol? sU 'sournS€l U 'slaHcu8Jq ltq€q pu" $cueosolouur osournldflluoutruold slr ^q senadsoseql tuo4 'V .Llr/0lqJxe Ip poqsm8uusrps ,arrp./, arnv ]I.'enIN'dslllqqou pue .IenW.gDlltuldopruau.V .tltu.g 'fla{Elg .V,talpurlryle DIItqdoDt X pueuop rz141t111t8 peh\SDlloluiDlo| 'V ")( V,^aIqrtq1t pue urTsBtrAtrlltpoqo sdaJun t' :soroedsSul,rolloJ eql ^q p4ueseldotst Elnsutued sj,(g eql uo pue dno$ opI^\-BrJEJlsnVu? srsrqJ .elctunJuuoJrlrJ .qsrppe.l rreql ,{q pelcrrcue .{lloq^\ro.{l ?d el? qclq,r spoosfq pue secusJsoJouutosouocel f,lpnsn fq pozuel3?lBqt .sag1u4[y saneds JC aDampoKqd uon:,esDpDrV Jo dno$ B ot savoleqDarDptJ olcocy 'Joqopo puB ^l?nJqod ',{J?nue[ur polcollorueeq a^Bq soun8el eJnletrAl '^l€nrqod ol ^ln[ uorJ paloelloruooq e^?q (seunSoloJnl?uur ro oJnlaurqll^r oq? uouo) 'pouod ro,tlolJur suorurcods SuuaA\oUSuol ? qlr,r [t8uruaag 'pouad3q4n{ puo Satauolg

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eur srncco rreraqa'Err,rrsnv qlnos'?rnsuruod rr^.J'i#:?,r'&uJi"rYr"jJX:Tyitfitr; '(Cy '1699 .S.t.O II€) gtgl pue EItl tatq!q,/!l :G.E092086pue '2961fnnrqed rcnE9L6 (Iy)'u's p.tooglDtoisDd uolptlsny qjnos pue 'u's 'W't 'SNYJ) t..(ZIZgtl L6 I I Z9€I16 CV) qrlws L6I tllr.t ZZ:(HIUAd ZZ suowutls xa 'u's 'S'11'296t 7tl:(nAOv.fl ,896t 180609/6Cv) platr ^r€ruqed € :(S00t0696CIy).us n33ne ^Iry :(Cv) '.ElJIg.d '(v) '(tg]Ar '(Iy) '(cv) 6'.rgn 'csJZ9t taqq d 8689 ,68€ 96st naqq s:(HJLdg.d 'CV) '(JV) .pauw1ra Z6€I pu? (Cgl 06€I s33ug e.t :vI.IyUJSnV HL|IOS suawnadg ',{o[0,{ 'pouo{rql IHD ele^elc e ur Suu?urui]ol pu? {oeq SurlqnopeJoJaq 1r olcJrcue ^[oq,t Jo % ol pees,{aoloqBupuelxo uorll puBouln8el ol tuatxqc?tlel€ou (Iu? oql Jo dol te^o spuelxo^ltuenbo4 q.rq,r) ploJlroqs € qtt,r ^ll€nsn ',fip ,(ql3uel ueqa olullq fra^ pue u,rotq qsppoJot u,to:q 1q8r1 popu?dxo)Buol utru gl .c 'uuoJrtg '8uol .i ^Hcrqla1rlunt'.eqll'ww g'I-C I utul g t-9.2 apan||]rn!le\ , sp:e,lno1iuruedo rAoJJ?u? qll/$.ro snonurluocJeqlro'otnrsqo ^Ja^ uotSotnald :\cElq ,,{utqs,ipq8ns ,(Icrql .opl,r luur S'Z-Z)posseJduoc lEq,{euos ol pr8rnl Jaq}el luut g.E-t ,Buolu]ru ,pro8uolqo 'atun8el .8uot .c r-9 Jo xod?SuroBJ Irre qlr^r purpnlEuol qpras rurug edqspseq,otnre ^lldnrq? xede'oleFcllar floJnrsqo ^Ja^'8uno,{ uaq,{ lseol1e asoumld,{lsnon3dsuoc ,u,to:q eled 'snorqelS'p^rno 'snoareuoo ,{llq8rJstuo ol iq8lells {pq8rJsol snooJ" Bqc,{1rurr; ,arunEel :ad 'epr,r '8uol ,rncco spess0l ol dn qtl,r rutu 9,9 uo (01)6ol suorlculsuocdeop ruopuEt,roJ e,{[?uors?ccoq8noqtle u]oql uoe,rtoqpetou]suor fllq8rls ^te^.,(lpnsnpu? spooseql te^o 'Suolqo .ss3[e^Jeu,srsaq]us pesreJ,{llr 8 ^lt{lotJzu iJs^ sawnSaTelsses,{./r,1() 1?pexauol 1ou 'olnce 'os?q .8uol .r .uoqt ^lldnJq€puB elz^oqo fl.rouBu ol oarJ turu /.1 s1r7ad uoo,rloq 'elq,r ,snouelquau.eleurqJnl enssqsnoueqderp q1l,\4(so^lau pdos) saduls u,l olq lq8rTS qlh\ ',telqrq/{ aDaapotrqd vo\ies DlrrrF uprl€ltsnv qlnos a f (l t urtset\l U.g NuytsiaVol. b. No. I (1q87)

Conservationstatus. Endangered (Leigh et al. 1981).Known only from a few hundredplants occurringalong a road vergeand on adjacentlease-hold land within a very rest cted area (about4x2km).

Etymology.Cretaceus lL.) - chalk-white.Refen to the conspicuouslypruinose branchlets, inflorescencesand legumes.

4. Acacianematophylla F. Muell.ex Benth.,Lrnnaea 26:612 (1S55).Figure 2F-H. "Boston TyW citation. Point NovaeHollandiae australioris (F. Miill.)". Zype:Boston point, 5.A., C. Wilhelmi(holo: K; iso:C, K, MEL 615302,6\ 5306,615307,615310 - seenote below).

Bushyslirub to c. 2-5m talI.Branchlets angled at extremities,glabrous. pftyl/odes narrowly linear,(14) 20-40 mm long,0.7-1.5(2) mm wide,ascending to erect,straight to shallowlycurved, uncinate,flat to terete,usually drying finely wrinkled,glabrous, grey-green, 4-nerved in all; midrib tmpressedor sometimes(when phyllodesflat) slightly raisedpulvinus distinct and articulate.Peduncles axtllary, I per node,4-15 mm long,glabrous, base ebra cteate. Flower. heodsglobrlar,23 4l-flowered.Flowers 5-merous.Calyx gamosepalous.Icgumes hnear, raised overseeds and slightly to moderatelyconstricted between them, to 1I cm long,5-6 mm wide, coriaceouscrustaceous to sub-woody,glabrous, longitudinally rugose. Seeds longitudinal, oblongoidto ellipsoid,4.5-5 mm long,c. 2.5 mm wide,black, Vz-%encircledby the filiform, reddish brown funicle.

Distribution.Coastal dunes of S.A. from near DrummondPoint to Point Boston(southern )and Innestone to Sturt Bay (southemYorke Peninsula).Also recordedfrom Venus Bay (c. 125km N of Drummond Point) and Mypongaon the FleurieuPeninsula.

Selectedspecimens examined.SOUTH AUSTRALIA: Near SleafordMere, about 15km SW of Port Lincoln,9 Nov. 1968,J.B.Cleland s.n. (AD 96921030);Myponga, R.-8. Filson 3002 (MEL); About 2 km S of WhalersWell Swamp, Peninsula,T.R.N Lothian 3'709 (AD); Sturt Bay, c. 15 km S of Warooka,B.R. Maslin 4538 (PERTH); 13.5km S of Port Lincolnsilo, Z.D. Wlliams9'l13 (AD);12mi [9km] from Port Lincolnalong road, J. W. Wrigley W Al 68 7632 (AD 91302081J .

"Boston-point, Typespecimens. The holotypeof,4. nematophyllaat herb.Kew is annotated Dr. F. Mueller". The specimenoriginated from the SonderHerbarium and the sheetis stamped HerbariumBenthamianum 1854. An isotypeof thistaxon, also at herb.K (duplicatesat MEL), showsthat the plantwas in fact collectedby C. Wilhelmi,not F. Mueller.This Kew isotype "Boston sheetis stampedHerbarium Hookerianum 1867 and is annotated(in Mueller'shand) Point, Port Lincoln. Legit. Carl Wilhelmii exam. Dr. ferd. Mueller". Although Bentham determinedthis specimenas A. nematophyllahe seeminglydid not do so prior to the publicationof the protologueotherwise C. Wilhelmi would presumablyhave been given as ",4. the collector.Above Bentham's annotation someone has written calamifoliaSweelvar." and underlinedthe speciesname in red. This underliningindicates that the specimenwas examinedby Benthamprior to the publicationof Acacia in Fl. Austral. vol. 2 (seefootnote on p. 8 of Fl. Austral.vol. 1, 1863).In Fl. Austral.vol. 2 (1864)Bentham included the name A. nematophyllaunder A. calamifolra,presumably following Mueller (1863:l2).

The Murray scrub specimensreferred to in the protologueof A. nematophyllaarc A. wilhelmianaF. Muell. (fideCourt 1972:.\60\. '(6 ,0SI9l9 9t9.Igru uo4 luoruAB{,HIUAd pue -IAIN'){)'u's ',4'lggl ,oAuEU 6lIgB n anw loo'V srapurld:sad{ppaLo.ttd.(lSIglg 'u "V 'ulocurl S TAr{) s twpqlrll J S ttod Jeou sloouods:(patceles oroq) ad(rcJraT JInD'J ,,.o1,resep ur e8ueu sropurldsoluoru pv Iulor{ll,\\ ulorur.I snuod ol?tlutcr^ uL,.uo\Djlr adtJ 'ollt?lt\ ,.llsntr{ .C .S sropulg :enlB^\ olqEtoN (gsgl) 9:l [UBEll sllqBlou ep8Jv 'sorJnu lBcrd?pe^lnool qcrq,r sopoll,{qdJroql ,{q osp pu? soruoJ?l ur peAuEJl? lcel ',{m[rye lJoqs lou oreqclq,r selJunpodoleolceJqa Jleql,{q poqsrn8uDstp^lrp€oJ eJe lnq oztspu? adBqs epollfqd u (Ic€Ifl I snaut re^ DdtDJotrltu V pellec^Fnoeuoro ueuo ore qcy4n\ 'tr .(CV \1oltuolot Jo srxroJopoll^qd aleloecuelqo olquasot spHq^q e^rlqnd osoqJ ,l.gZI pue g 'y'J) 'e^lasa{ J Iioqv elnsuruodoJ,{A ?un?J puB?lolg ulocur.I:(qy ,6Zn ggNS sd\Uqd '^?g lr'Jry)€lnsuruad o)trol lnod spr?,rolu,roto{rotr ruo4 gZ :(HJUAd ,0tst [ur{ ,9fl90086S}] lur altsDw V'g:,^[SN'tt6z pu€ tatdoJ g:/€t90086 ,€fl90086 ''u's ,4[SN'6262',{efl CV uMo.tg7) BlnsuruodeUoA nls :ol? potcollo3ueoq o^eq spuqfq elqzqo:deseqt qcq,r ruo4 serlrl?coloqJ etn8tC)pBoq Jod sre,rou Jequnu,qlpr,r orun8al ,oJnlxol pu? 'slueJed[,IZ Jo qlphr opoll^qd zrL o^rlBlndo,rl oql uaa,rloqelerpeuJolur ole qcrqa sJola"JBqolo suoneurquorsnoue^ ssessod slueld oseqJ'sqnsurued al^g pueo)tloi uloqlnosuo s.rrlrTerol ,{.\aJB rno{ paprocerer? sdnuD y pue oyfu1do1owa,.t' uoe,rleq spuq klelqeqord.spt.tqtH

'sdnuD 'y lepun ,{oI aql ur uo^r8eJe sanads o,rl eql uaonAleqsocuoJoJJrp eqJ .socBlderuos ur ozpuq,{q01 leedd? '^{oloq 'pue sercodso,r1 eql possnrsrpse sdnuD U 01perllE osle st tlltrtldolDruaumm)V 'Dll(tldoiowau t, JoJoleudoJddB paJoplsuoc $ rrJnodsoJoJoJeql puE (lousslel xe 'urq.-I 'Z IueJ l yauslau ruo{ qtueg olopdoldal ,slllllt .91ouototg x .v woJJDlptraopJ 'U tr 3'e) dnoJ8 cruouoxBt apr,r-?rT?llsnySIql ur sorcedsloqlo ouos uoo,rleqsoJuoleJJrp .ex?l sElu"crJruSls se eq ol J?eddeuoql ueeatoqsoruoleJJrp sqJ onateql Jo uorl€lBdosor{l roJ enEJ€salnqull? cqdelSoe8 pu8 opoll,{qd'aoueJsoJolJur eql uorl?urqum uI 'r3^o,{{oH uo{e} ueq,,[ lqBITsureos ^eur o11o!ruo1otVpueo tqdopwa, tz uoo,rloqsaruojoJJrp eqJ 'Dllo./iutolD) U Jo osoql ueql popluu/A^losreo, orolu ^rp ol puot ^eqt (e^oq? froSot?ctsru oql Jo sopoll,{qdqtl,r) slnccod?lre^o oloq,r }nq srql ueql ro}toqs,{l}uonbo{ oJesapoflfqd ar]t olltqdoiouau U uI teu pu? (uru g-Z x rur 9-9.€ ,{lpnsn)oleloacu?lqo 'teu .,{louJ?u,seuo8olec ,{I,,{oJJ?uJo o} aletol pue (uu g l I x uc 0l-E fll?nsn) fteurT o,rl Jo auo olur IleJ^aqt ,{ll?louo8tnq alq?U?^,{ro^ arBap0aDp).U Jo sepoll^qdoqJ 'saycunped qtl,r Suol ruc ? ol dn qceal saurlorrlos .o6.qdoputau.V ^eql lnq soJtunpedt Z qlur Suolturu t I ,{luouruocarB souecBl aqJ ur sE slx? opoll^qdul,{r?lrlos ale salrunpedoq1 q}rolt,rou eql uo pue sorurJeJasoql 'Ja^o^\oq'uelJo lno,rol8 orxos sauer?J.{Jellrxe ur po8uelJeala sslrunpadaLll D1ottut\lD, .V Jo'tno uI u.,r\oJ8s?q qcrql\ euec?l p Joocueftadd? oql a^sq lou seopelnpnJ]s eqt sepou Ie]oue8.tueudolo^op aseqllE pedolo^apte,{eu ojo,r sopolfqd Jr ue^g roqt ur ,{lleepeddop a^?q .olrunpod ,{?uIsopoll^qd oqi po^losqoaq louuec sl?osoloq,\t sos€J ,{4.0J ,{JeA eqt uI eql Jo oprslerx?q€ aql uo lolqouelqaql uo sJnmoJEOS ? oculs luaseld ocuo elo,r sopoJl^qdoql l?ql ftalo sr1r sos?o ]soru q (sopollfqdJo slxe ur selJunpod^Jplttos ssessod slelqou?Jq oseql uo $pou plsr.p)tuasqB sepoll^qd Supuolqns oql al? slolqrueJqeuros Jo es?q eql l?ou sopouaeJ ? le ^lerer -tuo^,iluo sepolfqd oqt Jo slx? aql ut ,{fflttos s,{B^Ue}sorulE oJe sopunpod oql 'V .V oll{qdopuau V ul puBlurDlto0w\ln pue seunpI8lseoc uo Burrrnl}o Dlltqdojmuau qlr,{ or4Bdoll€oJB soroods o,rl eql sopoll,{qdroql luelxeouos ol pu?smueJse:ogur rreql ,4q poqgn8uusrpoq uex tatqlnq DlptraotoTv elqeuB^Aqtrq oql ol petBIoJ^losolo flurB er sr '(Z/6t) ,(E/gt ,t9SI) Dll[qdonuau n)orU'(0g6t) fotqlq,|[ lrno] ueqluog oprJ,,{o1pur1 xi lee^^SDlloJlwDlDJ tr Jepun,4ul,{uou^sur pepnlcurueeq suq oueu $ql ,rou lrlun sa!J!u![!y '.{alqrq^A .U.g I' aoaulpollltlduollc.s Dlrrrl usrlertsnvqinos a t.(J t ulsphtr 32 NuyrsiaVo1.6. No. I (1987) Bushyspreading sft rub to 3 m tall, rarely taller.Branchlets slightly angled at extremities, dark red-brown,glabrous. Phyllodes nanowly ellipric to oblanceolateor narrowly oblanceolate, usually6'13 cm long and 8-25mm wide with l:w : 4-10,coriaceous, glabrous, glaucous to dull green;midrib and marginal nervesprominent, obscurely penninerved; pultinus prominentand 4-6 mm long. Racemesusually 2-6 cm long with 4-16 flower-heads,axes glabrollsPeduncles 2.5-6 mm long,rather stout, glabrous. F/ower-heads globlular, occasionally oblongoidin bud, bright golden,27-36-flowered. Bracteole laminae evident inbuds, dark brown to black,white fimbiolale. Flowers 5-merous.Calyx gamosepalous.Izgumes narrowly oblong, to 7 cm long, 8-13mm wide,firmly chartaceousto slightly coriaceous,convex on opposite sidesover alternateseeds, glabrous, stipitate. Seeds transverse, oblongoid ellipsoid, 4.5,5.5 mm long, 3-3.5mm wide, black,encircled by the filiform, red-brownfunicle.

Distribution.South Australia, Victoria and New SouthWales. In S.A.ranging from Tintinara (South-easternRegion) north through the Lofty Rangesand northern Yorke Peninsulato ArkabaStation (Flinders Range) and then east to GawlerRanges and near Cummins (southern Eyre Peninsula).Less common in N.S.W.and Victoria. In N.S.W.recorded for BrokenHill and near Menindeeand in Victoria from Nathalia and near Meringur.

A numberof SouthAustralian specimens previously included under L notabilishaye now beentransferred to either .4. alcockiior ? A. ancepsx notabilis(see above).

Afiinities. Allied to L ancepsfrom which it is distinguishedby its shorterpeduncles which ate arrangedin axillary racemes,smaller flower-heads and firmly chartaceousto slightly coriaceouslegumes (see key underA. anceps).Possible hybrids between these two species are discussedunder A. anceps.Also allied to A. beckleriTind. from which it is readily distinguishedby its glabrouspeduncles (not denselyminutely tomentellose)and transverse seeds(not longitudinal).The N.S.W. specimencited by Bentham(1864) under A. notabilis ts A. beckleri.

Acknowledgements

Prof.K.H. Rechingeris gratefullyacknowledged for providingthe Latin descriptions.John Briggsand FraserVickery are thankedfor providingvaluable field data and specimensof A. alcockii andA. cretacea.John is further thankedfor very constructivecdticism of the manuscript.Marion Simmons and the lateBruce Copley also provided very useful observations onA. cretacea.This specieswas first drawnto D.J.E.W.'sattention by the late MeIV Smith of Cowell.David Keaneis thankedfor illustratingFigures lE, lF and 38. SuzanneCurry, employedby the first author undera grantfrom the Bureauof Flora and Faunais thanked for competenttechnical assistance. The work was conductedat the WesternAustralian Herbarium,Perth (B.R.M.)and the StateHerbarium, Adelaide (D.J.E.W.).

References

"Flora Bentham,G. {1864). Austraiiensis."Vol. 2. (Reeve:London.) Bentham,G. (1875).Revision of the SuborderMimoseae. Trans. Linn. Soc.London 30: 335664. Court. A.B. (19?2).Notes on Austmlianacacias L Muelleria "Rare 2t 155163. Leigh,J., Briggs,J. and Hartley, W. (1981). or ThreatenedAustralian Planrs." (Australian National Parks and Wildlife Service: Canberra.l Maiden,J.H. and Blakely,W.F. (1927).Descriptions of fifteen new acaciasand noteson severalolher species.J. andProc. pl. Roy."TheSoc. N.S.W.60: 1?1 196. l3 18. Mueller,F. (1863). PlantsIndigenous to rhc Colooyof Victoria." Vol.2. 1.40 'Prodromu Stearn,W.T. (1960).An lntroductionto RobertBrown's s FloraeNovae Hollandiae' Facsimile edition. (Cramer:Weinhem.) "Acacias Whibley,D.J.E. (1980). of SouthAustralia." (covt. Printer:Adelaide.J