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Cretaceous: Albian) of Kansas Author(S): Rodney M Paleontological Society Huhatanka, a New Genus of Lobster (Decapoda: Mecochiridae) from the Kiowa Formation (Cretaceous: Albian) of Kansas Author(s): Rodney M. Feldmann and Ronald R. West Source: Journal of Paleontology, Vol. 52, No. 6 (Nov., 1978), pp. 1219-1226 Published by: SEPM Society for Sedimentary Geology Stable URL: http://www.jstor.org/stable/1303932 Accessed: 22/02/2009 21:33 Your use of the JSTOR archive indicates your acceptance of JSTOR's Terms and Conditions of Use, available at http://www.jstor.org/page/info/about/policies/terms.jsp. JSTOR's Terms and Conditions of Use provides, in part, that unless you have obtained prior permission, you may not download an entire issue of a journal or multiple copies of articles, and you may use content in the JSTOR archive only for your personal, non-commercial use. Please contact the publisher regarding any further use of this work. 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Paleontological Society and SEPM Society for Sedimentary Geology are collaborating with JSTOR to digitize, preserve and extend access to Journal of Paleontology. http://www.jstor.org JOURNAL OF PALEONTOLOGY,V. 52, NO. 6, P. 1219-1226, 1 PL., 3 TEXT-FIGS., NOVEMBER 1978 HUHATANKA, A NEW GENUS OF LOBSTER (DECAPODA: MECOCHIRIDAE) FROM THE KIOWA FORMATION (CRETACEOUS: ALBIAN) OF KANSAS RODNEY M. FELDMANN AND RONALDR. WEST Kent State University, Kent, Ohio 44242 and Kansas State University, Manhattan 66506 ABSTRACT-Reexamination of the type material of Squilla? kiowana Scott, 1970 and study of addi- tional specimens of the same taxon indicates that it is a mecochirid decapod referable to a new genus, Huhatanka. This assignment brings to five the number of North American species referable to the Mecochiridae. INTRODUCTION Formation but have yet to be verified and de- R. com- THE Kiowa Formation has been the subject scribed (Frederick Schram, pers. The of in the of a number of recent papers dealing with var- mun.). majority species family have been described from and the ious aspects of Cretaceous stratigraphy and Germany British Isles in addition to these lo- paleontology. In 1970, Scott described the or- although calities and North American ganisms collected from the unit, carefully de- occurrences, spe- cies have been described also from localities in fined their position in the stratigraphic se- New Zealand and Antarctica quence and detailed the paleoecological Africa, (Forster, this broad distribu- setting of the unit. One of the forms collected 1971). Notwithstanding mecochirids are rare fossils. was a small, poorly preserved arthropod rep- tion, resented two which Scott by specimens (1970, SYSTEMATIC PALEONTOLOGY p. 85) referred to the Squillidae, a family of 1803 stomatopods. Subsequently, the senior author Order DECAPODALatreille, received five specimens of equally poorly pre- Infraorder PALINURALatreille, 1803 1883 served specimens from Loren Avis, then a stu- Superfamily GLYPHEOIDEA Winckler, 1925 dent at Wichita State University. The speci- Family MECOCHIRIDAEvan Straelen, mens remained a curiosity until, in 1976, the Genus HUHATANKAn. gen. junior author brought together ten specimens Type species.--Squilla? kiowana Scott, which had been collected by students at Kan- 1970, by original designation. sas State University. This material, some of Description.-Cephalothorax with short, which was better preserved than the earlier smooth rostrum; median, subdorsal and sub- discoveries, permitted us to accurately identify orbital pustulose ridges; oblique, subparallel the organisms for the first time, refer them to cervical and postcervical grooves; branchio- the proper order and frame a more complete cardiac groove weak or absent. Abdomen gen- and detailed description of the animals. erally smooth; pleuron of first abdominal so- The placement of this species in the Meco- mite quadrate; that of remaining somites chiridae brings to five the total number of spe- elongate oval to nearly circular. Pereiopods cies in this family described from North Amer- long, slender, achelate throughout, generally ica. Two species of Pseudoglyphea, P. mulleri smooth. van Straelen, 1936 and P. bordenensis (Cope- Remarks.-Forster recently (1971) sum- land), 1960 have been described from the Ju- marized information on known representa- rassic of Nevada and Borden Island, Canada, tives of the Mecochiridae. He recognized only respectively and two species of Meyeria, M. Pseudoglyphea Oppel, 1861, and Mecochirus mexicanus (Rathbun) 1935 and M. ? harveyi German, 1827, as valid genera and considered Woodward, 1900 have been described from Meyeria M'Coy, 1849, synonymous with Me- the Cretaceous of Mexico and British Colum- cochirus; Selenisca von Meyer, 1847, synon- bia, respectively. The only other possible rep- ymous with Glyphea thereby placing it in the resentatives of the family are three specimens Glypheidea; and Praeatya Woodward, 1868, which have been collected from the Sundance synonymous with Pseudoglyphea. Selenisca Copyright ? 1978, The Society of Economic Paleontologists and Mineralogists 1219 1220 RODNEY M. FELDMANN AND RONALD R. WEST The degree of difference shown between I forms typically assigned to the two genera ap- I pears to us to be great enough that the genus I Meyeria Meyeria should be retained to include those I 0 (0m species with thicker, spinose or pustulose, in- -kc U_ c ornamented abdomina and relative- co I tegument, .- c LUJ -v Il ly shorter first pereiopods. These types of dif- OuU ferences are comparable to those used as Huhatanka points of generic distinction in better known, I recent decapods, such as genera of the Ne- I phropidae (Holthuis, 1974). I Using generic discriminators similar to those I i used by Forster (1971), but retaining Meyeria I as a distinct genus, the specimens referred to I Huhatanka n. cannot be I gen. clearly assigned to taxa is I existing (Text-fig. 1). Pseudoglyphea u Mecochirus characterized of well Li by possession developed, I broad postcervical and branchiocardiac (1 5 I grooves, neither of which parallels the cervical a I groove; spinose rostrum; subquadrate pleura Q, on the second abdominal somite; and a flat- tened, spiney propodus on the first pereiopods. 0 A point of similarity between this genus and 7O I 3 Psud l Huhatanka is that both have propoda that do _|)*"- Pseudo lyphea not exceed the length of the cephalothorax. Both Mecochirus and Meyeria have extremely Li I long first pereiopods and in both cases, the --_ I propoda exceed the length of the cephalotho- L I rax. The pleura of the second abdominal so- mite in Mecochirus are triangular whereas TEXT-FIG.1 -Stratigraphic distribution of genera those of are reduced and referableto the Mecochiridae. data and Meyeria subquad- Range rate. The of both these morphologicsketches, with the exceptionof those groove patterns genera for Huhatanka, were taken from Forster (1971) resemble that of Huhatanka in that they are and Glaessner(1969). reduced and much more subtly expressed than in Pseudoglyphea. This character has led to quite variable interpretations of groove pat- and Praeatya had been questionably referred terns (compare, for example, Forster, 1971, to the Mecochiridae by Glaessner (1969, p. fig. 5 and Glaessner, 1969, fig. 271, 1). The R466-R467). Meyeria was placed, without branchiocardiac groove in Huhatanka is re- question, in the Mecochiridae by Glaessner duced to the point that it can no longer be (1969, p. R465) and was considered distin- identified with certainty. guishable from Mecochirus based on the or- Another aspect of the morphology of this namentation of the cephalothorax and abdo- group that has been subject to several different men as well as the length of the first pereiopod. interpretations is that of the nature of the ter- Forster (1971, p. 398) argued that the distinc- mination of the pereiopods. Forster (1971, p. tion between the two genera was primarily 404), for example, described the first two pe- stratigraphic and that because the species as- reiopods of Mecochirus as subchelate although signed to the two genera were typically pre- the widely accepted definition of subchelate served in different styles, it was difficult to (Moore & McCormick, 1969, p. R102) would make detailed morphological comparisons. not embrace them. Subchelae must be grasp- Where it is possible, he suggested that the ing structures in which the dactylus rotates characters grade from one genus into another back onto some part of the propodus to pro- and the genera are, therefore, probably syn- duce a grasping surface. In Mecochirus and, onymous. for that matter, Pseudoglyphea the backward CRETACEOUS LOBSTER FROM KANSAS 1221 rotation of the dactylus is restricted by the presence of a spine on the distal termination of the propodus but no functional occlusal sur- face results (Text-fig. 1). Rather, the termi- nations of these appendages seem to be better modified for probing and, perhaps, raking the substrate in search of food. They could better be defined as achelate to distinguish them from the truly prehensile subchelae observed on TEXT-FIG.2-Reconstruction of Huhatanka kio- some glypheids, some palinurids and some sto- wana (Scott).Details of sixth abdominalsomite, telson and are matopods, for example. If the definition was uropods conjectural. thus restricted, none of the Mecochiridae would possess subchelae. Etymology.-The generic name is derived vical groove; marginal groove distinct on pos- from two Plains Indian words; Huha = limbs, terior and posteroventral margin of carapace, and tanka = large (Riggs, 1893). unobservable on anteroventral surface.
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