Checklist of South Korean Metopiinae Forster, 1869 (Hymenoptera, Ichneumonidae)

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55 HOSTED BY Contents lists available at ScienceDirect 56 57 Journal of Asia-Pacific Biodiversity 58 59 60 journal homepage: http://www.elsevier.com/locate/japb 61 62 63 Review Article 64 65 1 Checklist of South Korean Metopiinae Förster, 1869 (Hymenoptera, 66 2 67 3 Ichneumonidae) with new South Korean species and a note on 68 4 Seticornuta koreana 69 5 70 6 * 71 7 Q14 Jin-Kyung Choi, Jong-Wook Lee 72 8 Department of Life Sciences, Yeungnam University, Gyeongsan, South Korea 73 9 74 10 75 11 article info abstract 76 12 77 13 Article history: The subfamily Metopiinae (Hymenoptera: Ichneumonidae) was represented in South Korea as 96 species 78 14 Received 18 August 2016 belonging to 13 genera. In this study, six additional species belonging to the genus Triclistus are recorded 79 Received in revised form 15 for the first time from South Korea. A checklist of South Korean Metopiinae and a note on Seticornuta 80 7 September 2016 16 koreana are provided. Accepted 13 September 2016 81 Copyright Ó 2016, National Science Museum of Korea (NSMK) and Korea National Arboretum (KNA). 17 Available online xxx 82 Production and hosting by Elsevier. This is an open access article under the CC BY-NC-ND license (http:// 18 83 creativecommons.org/licenses/by-nc-nd/4.0/). 19 Keywords: 84 20 checklist 85 new record 21 86 22 Seticornuta koreana synonym 87 23 taxonomy 88 24 Triclistus 89 25 90 26 91 27 92 Introduction Materials and methods 28 93 29 94 The subfamily Metopiinae is one of the large subfamilies within Materials used in this study were collected by sweeping and 30 95 the family Ichneumonidae, consisting of approximately 900 species Malaise trapping, after which they were deposited in the animal 31 96 within 26 genera. systematic laboratory of Yeungnam University (YNU; Gyeongsan, 32 97 The South Korean species of Metopiinae have been studied in South Korea). Morphological terminology follows mostly that of 33 98 recent years mainly by Korean taxonomists (Choi et al 2015a, Gauld et al (2002). Specimens were examined using an AxioCam 34 99 2015b, 2016a, 2016b, 2016c; Kang et al 2016). Based an ongoing MRc5 camera attached to a stereo microscope (Zeiss SteREO Dis- 35 100 taxonomic study on South Korean Metopiinae, we report here six covery, V20; Carl Zeiss, Göttingen, Germany), processed using 36 101 unrecorded species of Triclistus for the first time from South Korea. AxioVision SE64 software (Carl Zeiss), and optimized with a Delta 37 102 Triclistus Förster, 1869 is a cosmopolitan genus comprising 82 imaging system (i-solution; IMT i-Solution Inc., Vancouver, BC, 38 103 described species, and is mainly known from the Palaearctic, Ori- Canada). Distributional data mainly follow that of Yu et al (2012). 39 104 ental, and Nearctic regions (Yu et al 2012). In the Eastern Palaearctic Diagnoses of newly recorded species are based on South Korean 40 105 region, 26 species are known with just five species occurring in specimens. 41 106 South Korea (Lee and Cha 2000). This genus attacks the larvae of Abbreviations used in this paper are as follows: CB, 42 107 microlepidopteran species such as Tortricidae (Gauld et al 2002). Chungcheongbuk-do; CN, Chungcheongnam-do; GB, 43 108 In this paper, we provide the first records of six species of Tri- Gyeongsangbuk-do; GN, Gyeongsangnam-do; GG, Gyeonggi-do; 44 109 clistus for the South Korean Metopiinae fauna and a note that is GW, Gangwon-do; JB, Jeollabuk-do; JN, Jeollanam-do; JJ, Jeju-do. 45 110 species valid on Seticornuta koreana. 46 111 AEI: American Entomological Institute, Gainesville, FL, USA 47 112 ANSP: Academy of Natural Sciences of Philadelphia, USA 48 113 BBM: Bernice P. Bishop Museum, Department of Entomology, 49 * Corresponding author. 114 Honolulu, Hawaii, USA 50 E-mail address: [email protected] (J.-W. Lee). 115 DEI: Deutsches Entomologisches Institut, Germany 51 Peer review under responsibility of National Science Museum of Korea (NSMK) and 116 Korea National Arboretum (KNA). 52 117 53 http://dx.doi.org/10.1016/j.japb.2016.09.006 118 54 pISSN2287-884X eISSN2287-9544/Copyright Ó 2016, National Science Museum of Korea (NSMK) and Korea National Arboretum (KNA). Production and hosting by Elsevier. 119 This is an open access article under the CC BY-NC-ND license (http://creativecommons.org/licenses/by-nc-nd/4.0/).

Please cite this article in press as: Choi J-K, Lee J-W, Checklist of South Korean Metopiinae Förster, 1869 (Hymenoptera, Ichneumonidae) with new South Korean species and a note on Seticornuta koreana, Journal of Asia-Paciﬁc Biodiversity (2016), http://dx.doi.org/10.1016/ j.japb.2016.09.006 JAPB185_proof ■ 5 October 2016 ■ 2/19

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1 ETH: Entomologisches Institut, Eidgenössische Technische si, Jangan-gu, Sanggwanggyo-dong, Mt. Gwanggyosan, 15e25 vii 66 2 Hochschule 2008 (JO Lim); 1_, GB, Cheongdo-gun, Unmun-myeon, Mt. 67 3 HOPE: Entomological Collection, University Museum, Oxford, En- Unmunsan, 3e20 vii 2009 (JW Lee); 1\, GB, Gyeonsan-si, Yeungnam 68 4 gland, United Kingdom Univ., 20 v 1990 (JS Lee); 1\, GB, Gyeonsan-si, Yeungnam Univ., 9 vi 69 5 HU: Hokkaido University, Faculty of Agriculture, Entomological 1987 (JY Cha); 1_, GN, Jinju-si, Ibanseong-myeon, Daecheon-ri, 70 6 Institute, Sapporo, Japan Gyeongsangnamdo Arb., 15 viie1 viii 2011 (JW Lee). 71 7 IZB: Institut za zastitu bilja, Fakultete poljoprivrednih znanosti, Distribution. South Korea, China, Russia. 72 8 Croatia 73 9 IZU: Instytut Zoologiczny Uniwersytetu, Sienkiewicza, Wroclaw, Genus Chorinaeus Holmgren, 1858 74 10 Poland Chorinaeus Holmgren, 1858: 305e394. Type species: Exochus 75 11 KU: Kyushu University, Entomological Laboratory, Fukuoka, funebris Holmgren. 76 12 Kyushu, Japan Polyrhabdus Walsh, 1873:65e166. Type species: Polyrhabdus car- 77 13 MACN: Museo Argentino de Ciencias Naturales, Avenida Angel iniger Walsh. 78 14 Gallardo, Argentina 79 2. Chorinaeus eniwanus Kusigemati, 1967 빛뭉툭맵시벌 15 MHN: Muséum d’Histoire Naturelle, Route de Malagnou, Genève, 80 Chorinaeus eniwanus Kusigemati, 1967a:17e27. Type: female; type 16 Switzerland 81 depository: HU. 17 MNHN: Muséum National d’Histoire Naturelle, Entomologie, Paris, 82 18 France 83 Material examined. South Korea: 1\, GW, Donghae-si, Samhwa- 19 MOMOI: Kobe University, Faculty of Agriculture, Entomological 84 dong, Mureunggyegok, 372704500N1260101700E, 31 ve15 vi 2005 20 Laboratory, Kobe, Japan 85 (JW Lee). 21 MZ: Musée Zoologique, Place Riponne, Lausanne, Switzerland 86 Distribution. South Korea, Japan, Russia. 22 NCM: Norwich Castle Museum, Norwich, United Kingdom 87 23 NHM: The Natural History Museum, Department of Entomology, 88 3. Chorinaeus flavipes Bridgman, 1881 연노랑다리뭉툭맵시벌 24 London, England, United Kingdom 89 Chorinaeus flavipes Bridgman, 1881:143e168. Type: female; type 25 NM: Naturhistorisches Museum, Burgring, Wien, Austria 90 depository: NCM. 26 NMI: National Museum of Ireland, Kildare Street, Dublin, Ireland 91 27 NNM: Nationaal Natuurhistorisch Museum, the Netherlands 92 Material examined. South Korea: 1\, CB, Danyang-gun, Cheong- 28 NR: Naturhistoriska Riksmuseet, Sektionen för Entomologi, Stock- 93 dong-ri, Sobaeksan National Park, 31 vii 1988 (HG Kim). 29 holm, Sweden 94 Distribution. South Korea, Czech Republic, Finland, France, Ger- 30 SIZ: Schmalhausen Institute of Zoology, Bogdan Khmelnitsky Street, 95 many, Ireland, Japan, Latvia, Lithuania, Moldova, Norway, Poland, 31 Ukraine 96 Russia, Tuvinskaya Respublika, Sweden, Turkey, Ukraine, United 32 TMA: Termeszettudomanyi Muzeum Allattara, Barossa-Utea, 97 Kingdom. 33 Budapest, Hungary 98 34 USNM: U.S. National Museum of Natural History, Smithsonian 99 4. Chorinaeus mandibulatus Choi and Lee, 2016 큰턱뭉툭맵시벌 35 Institute, Washington, DC, USA 100 Chorinaeus mandibulatus Choi and Lee, in Choi et al (2016, in press). 36 ZI: Zoologiska Institutionen, Sweden Q1 101 Type: female; type depository: YNU. 37 ZIN: Zoological Institute, Academy of Sciences, St. Petersburg, 102 38 Russia 103 Material examined. South Korea: Incheon-si, Seo-gu, Gajwa- 39 ZMHU: Zoologisches Museum, Humboldt Universität, Invalid- 104 dong, 1990; 1\, CB, Jincheon-gun, 11 viii 1991 (JW Lee); 1\, CN, 40 enstrasse, Berlin, Germany 105 Onyang-si, 19 vi 1983 (JW Lee); 1\, GN, Jinju-si, Gajwa-dong, 24e30 41 106 vi 1989; 1\, JB, Jeongeup-si, Naejang-dong, Naejangsan National 42 Taxonomy 107 Park Geumseonggyegok, 16e27 vii 2008 (JW Lee); 1\, Seoul-si, 43 108 Family Ichneumonidae Latreille, 1802 Songpa-gu, Oryun-dong, 1 x 1992 (YH Kim) 44 109 Subfamily Metopiinae Förster, 1869 Distribution. South Korea. 45 110 46 111 Diagnosis. Members of this subfamily have the following 5. Chorinaeus parvus Kusigemati, 1967 꼬마뭉툭맵시벌 47 112 morphological characters: size of species from small to large (fore- Chorinaeus parvus Kusigemati, 1967a:17e27. Type: female; Type 48 113 wing 2.2e16 mm). Clypeus not separated from face by groove; face depository: HU. 49 114 evenly convex except in Metopius (Metopius has a flat or concave 50 115 shield shaped). Sternaulus of mesopleuron absent or short. First Material examined. South Korea: 1\, GG, Gwangju-si, Docheong- 51 116 tergite with glymma and with spiracle before middle of tergite. myeon, Taehwasan, 26 vie7 vii 2007 (JO Lim); 2\\, Anyang-si, 52 117 Ovipositor short, usually not extending beyond metasomal apex. Manan-gu, Gwanaksan, 5e19 vii 2007 (JO Lim); 1\, Anyang-si, 53 118 M.T. 3, 26 vie4 vii 2007 (JO Lim); 1\, GW, Donghae-si, Samhwa- 54 119 Checklist of South Korean Metopiinae dong, Mureung valley, 31 ve15 vi 2005 (JW Lee); 1\, Inje-gun, 55 120 Girin-myeon, Jindong-ri, Jeombongsan, 26 vie28 vii 2012 (JY 56 Genus Acerataspis Uchida, 1934 121 Park); 1_, Hongcheon-gun, Bukbang-myeon, Gwangwon Prov. 57 Acerataspis Uchida, 1934a: 23. Type species: Cerataspis clavata 122 Environment Research Park, 15e30 v 2012 (SJ Jang); 1\, JB, 58 Uchida. 123 Namwon-si, Sannae-myeon, Jeonnyeongchi, 2 ixe15 x 2001 (JW 59 124 1. Acerataspis clavata (Uchida, 1934) 곤봉뭉툭맵시벌 Lee). 60 125 Cerataspis clavata Uchida, 1934b:275e277. Type: male; type de- Distribution. South Korea, Japan. 61 126 pository: KU. 62 127 6. Chorinaeus pectinatus Kusigemati, 1967 빗살뭉툭맵시벌 63 128 Material examined. South Korea: 1_, GW, Wonju-si, Heungeup- Chorinaeus pectinatus Kusigemati, 1967a:17e27. Type: female; type 64 129 myeon, Yeonse Univ., 29 vie15 vii 2011 (JW Lee); 15__, GG, Suwon- depository: HU. 65 130

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1 Material examined. South Korea: 1\, GB, Cheongdo-gun, Unmun- Univ., 29 v 1986 (MS Jang); 3__, GW, Wonju-si, Panbu-myeon, 66 2 myeon, Sinwon-ri, Mt. Unmunsan, Unmunsa, 3540025.7200N Seogok-ri, Paekunsan, 7 vii 1998 (DS Choi & DE Kim); 1_,GW, 67 3 12857037.5700E, 17 vii 1989 (JW Lee). Taebaek-si, Cheolam-dong, 22 vi 1991 (JY Cha); 1_, GG, Bokwangsa, 68 4 Distribution. South Korea, Japan. 9 vi 1986 (YJ Kim); 1_, Korea, GG, Cheonmasan, 11 vi 1982 (EY Lee); 69 5 1_, GG, ditto, 11 vi 1982 (HD Lim); 1_, GG, Myeongjisan, Kyimok, 70 6 7. Chorinaeus subcarinatus Holmgren, 1858 긴잿빛뭉툭맵시벌 Imsan, 29 vi 1999 (DH Hwang); 1_, CB, Wolagsan, 19e22 vi 2001 (S 71 7 Chorinaeus subcarinatus Holmgren, 1858: 305e394. Type: un- Cho, DP Lyu & J Lee); 2__, GB, Bonghwa-gun, Myeonho-myeon, 72 8 known; type depository: unknown. Bukgok-ri, Cheongnyangsan, 30 vi 2007 (JW Lee). 73 9 Distribution. South Korea, China, Japan, Russia. 74 10 Material examined. South Korea: 1_, CN, Gyeryeongsan, Gapsa, 75 11 24 iv 1983 (HC Park); 1\, GN, Jinju-si, Gajwa-dong, 7 iv 1987 (JS 10. Colpotrochia jozankeana Uchida, 1930 남방화장뭉툭맵시벌 76 12 Park); 1\, Korea, GN, Jirisan, Sunduryu, 3 vi 1989 (GS Kim). Colpotrochia jozankeana Uchida, 1930: 243e298. Lectotype: male; 77 13 Distribution. South Korea, Armenia, Austria, Bulgaria, Canada, type depository: HU. 78 14 Czech Republic, Finland, France, Germany, Hungary, Ireland, 79 15 Kazakhstan, Lithuania, the Netherlands, Norway, Poland, Romania, Material examined. South Korea: 1_, Seoul, Seocho-gu, 80 16 Russia, Slovakia, Sweden, Switzerland, Turkey, USA, Ukraine, United Umyeonsan, 19 ix 1991 (IS Kim); 1_, GW, Pyeongchang-gun, Hoer- 81 17 Kingdom, Uzbekistan. yeongbong, Deokgeori, 3 ix 1998 (YG Park); 3__, GW, Hongcheon- 82 18 gun, Hongcheon-eup, Jangjeongpyeong-ri, Geomdungae village, 7e 83 19 Genus Colpotrochia Holmgren, 1856 26 ix 2006 (JW Lee); 1_, GW, Hoengseong, Gapcheon-myeon, 84 20 Colpotrochia Holmgren, 1856:1e104. Type species: Ichneumon Junggeum-ri, Seomgang river side, 12 x 2002 (HW Byun); 1_, GG, 85 21 elegantulus Holmgren Kwacheon-si, Beolmang-dong, 3 ix 1994 (SJ Tang); 1_, GG, 86 22 Exochoides Cresson, 1868:1e38. Type species: Exochoides mexicana Kwangreung-si, 25 viii 1990 (EJ Ryu); 1_, GG, Anyang-si, Seoksu- 87 23 Cresson dong, 11 x 1994 (HW Seol); 1_, GB, Bohyeonsan, 19 ix 1998 (SJ Suh); 88 24 Alcocerus Förster, 1869:135e221. Type species: Tryphon trifasciatus 1_, GB, Bonghwa-gun, Daehyun-ri, 25 vii 1986 (KS Jang); 1_, JN, 89 25 Förster Baekyangsan, 5 viii 1974 (SH Nam). 90 26 Scallama Cameron, 1899:1e220. Type species: Scallama trilineata Distribution. South Korea, Japan, China. 91 27 Cameron 92 28 Aithris Cameron, 1900:1e114. Type species: Aithris coenutus 11. Colpotrochia kurisuei (Uchida, 1930) 어리화장뭉툭맵시벌 93 29 Cameron Colpotrochioides kurisuei Uchida, 1930: 243e298. Type: male; type 94 30 Ischyrocnemopsis Ashmead, 1900:1e220. Type species: Exochoides depository: HU. 95 31 texana Ashmead 96 32 Aethris Schulz, 1906:1e356. Material examined. South Korea: 1_, Onseiri, 24 vii 1924 (T 97 33 Inoresa Cameron, 1909:722e730. Type species: Inoresa pilosa Kurisue). 98 34 Cameron Distribution. South Korea, Japan. 99 35 Sychnoleteroides Brèthes, 1909: 225e256. Type species: Syc- 100 36 hnoleteroides ﬂavus Brèthes 12. Colpotrochia nigra Kusigemati, 1971 검은화장뭉툭맵시벌 101 37 Colpotrochioides Uchida, 1930: 243e298. Type species: Colpo- Colpotrochia nigra Kusigemati, 1971: 205e298. Type: female; type 102 38 trochioides orientalis Uchida depository: HU. 103 39 104 8. Colpotrochia cincta (Scopoli, 1763) 어리남방화장뭉툭맵시벌 40 Material examined. South Korea: no Korean specimens were 105 Sphex cincta Scopoli, 1763:1e420. Type: unknown; type de- 41 available for this study. 106 pository: unknown. 42 Distribution. South Korea, Japan. 107 Ichneumon elegantulus Schrank, 1781:1e548. Type: unknown; type 43 108 depository: unknown. 44 13. Colpotrochia nipponensis Uchida, 1930 네줄화장뭉툭맵시벌 109 Ichneumon mundator Thunberg, 1822: 249e281. Type: unknown; 45 Colpotrochia nipponensis Uchida, 1930: 243e298. Lectotype: fe- 110 type depository: unknown. 46 male; type depository: HU. 111 Colpotrochia afﬁnis Vollenhoven, 1875:1e16. Type: unknown; type 47 112 depository: unknown. 48 Material examined. South Korea: 1_, Seoul, Yongun-dong, 26 v 113 49 1991 (SH Bae); 1_, GW, Hongcheon-gun, Hongcheon-eup, 114 Material examined. South Korea: no Korean specimens were 50 Jangseonpyeong-ri, Geomdungae, 1e20 vi 2007 (JW Lee); 1_, GG, 115 available for this study. However, we have seen a voucher specimen 51 Cheonmasan, 12 vii 1983 (SH Kim); 1_, CB, Danyang-gun, Danyang- 116 from Hungarian Natural History Museum. 52 eup, Sobaeksan National Park, 25 vie9 vii 2007 (JW Lee); 1_, JN, 117 Distribution. South Korea, Austria, Belarus, Belgium, Bulgaria, 53 Kwangju-si, Kwangju Park, 22 vii 1985 (HJ Yun). 118 Croatia, Czech Republic, Finland, France, Georgia, Germany, 54 Distribution. South Korea, Japan. 119 Hungary, Italy, Japan, Kazakhstan, Lithuania, Macedonia, Moldova, 55 120 the Netherlands, Norway, Poland, Romania, Russia, Slovakia, Spain, 56 14. Colpotrochia orientalis (Uchida, 1930) 동양화장뭉툭맵시벌 121 Sweden, Switzerland, Turkey, Ukraine, United Kingdom, 57 Colpotrochioides orientalis Uchida, 1930:243e298. Lectotype: fe- 122 Yugoslavia. 58 male; type depository: HU. 123 59 124 9. Colpotrochia fusca Matsumura, 1931 화장뭉툭맵시벌 60 Material examined. South Korea: 1_, GG, Bukhansan, 5 viii 1989 125 Colpotrochia fusca Matsumura, 1931:1e1497. Lectotype: male; type 61 (MH Mok); 1_, JN, Haenam-gun, Durunsan,13 xi 2001 (SW Choi); 1_, 126 depository: HU. 62 CN, Buyeo-gun, Gyuam-myeon, Sumok-ri, 13 viie17 viii 2005 (JW 127 63 Lee); 1_, JN, Goheung-gun, Geunsan-myeon, 23 vi 2011 (JW Lee); 1_, 128 Material examined. South Korea: 1_, Seoul, Gangnam-gu, 64 GN, Tongyeong-si, Saleang-myeon, Eupeok-ri, 20 vii 2010 (EJ Hong); 129 Daeamsan, 1 x 1995 (SY Jeong); 1_, Seoul, Anam-dong, Korea 65 1_, JJ, Jeju-si, Donggye-myeon, Jeolmul, 25 viiie2 ix 2005 (CH Sin). 130

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1 Distribution. South Korea, China, Japan. Material examined. South Korea: 1\, GB, Gyeongsan-si, Daehak- 66 2 ro, Yeungnam Univ., 30 viiie25 x 2013 (JW Lee); 1\, GB, Kyeongju-si, 67 3 15. Colpotrochia osuzensis Kusigemati, 1971 점박이화장뭉툭맵 Hyeongok-myeon, Namsa-ri, 18e25 viii 2005 (JT Mun). 68 4 시벌 Distribution. South Korea, Finland, Germany, Hungary, 69 5 Colpotrochia osuzensis Kusigemati, 1971: 205e298. Type: male; Kazakhstan, the Netherlands, Norway, Poland, Russia, Slovakia, 70 6 type depository: HU. Sweden, Ukraine, United Kingdom. 71 7 72 8 Material examined. South Korea: 1_, Seoul, Seocho-gu, Cheong- 20. Exochus antis Tolkanitz, 2003 앞볼록뭉툭맵시벌 73 9 gyesan, 21 v 2002 (HJ Yun); 1_, GG, Yongmunsan, 27 v 1982 (UW Ko); Exochus antis Tolkanitz, 2003a: 1075e1085. Type: female; type 74 10 1_, GG, Suraksan, 29 v 1986 (SH Kim); 1_, GG, Cheonmasan, 6 vi 1990 depository: ZIN. 75 11 (JH Hwang); 1_, CB, Wolagsan,19 vi 2001 (S Cho); 1_, JN, Gurye-gun, 76 12 Sandong-myeon, Joaisa-ri, Banyabong, 13 vii 2002 (JS Sin). Material examined. South Korea: 1\, GB, Cheongdo-gun, Unmun- 77 13 Distribution. South Korea, Japan. myeon, Haksodaepokpo, 12 ve8 vi 2013 (JW Lee). 78 14 Distribution. South Korea, Russia. 79 15 16. Colpotrochia pilosa (Cameron, 1909) 붉은화장뭉툭맵시벌 80 16 Inoresa pilosa Cameron, 1909: 722-730. Type: male; type de- 21. Exochus areolaris Lee & Choi, 2016 넓적배볼배볼록뭉툭맵시벌 81 17 pository: NHM. Exochus areolaris Lee & Choi, 2016, in Choi et al (2016): 2332. Type: 82 18 female; type depository: YNU. 83 19 Material examined. South Korea: 1_, Seoul, Nowon-gu, Sulaksan, 84 20 25 v 1997 (IH Chae); 1_, Seoul, Gangnam-gu, Sanseong-dong, 11 viii Material examined. South Korea: 1\, Seoul, Noweon-gu, 85 21 1996 (SJ Lee); 1_, Seoul, Songpa-gu, Oryun-dong,1 x 1992 (YH Lim); Sangve4-dong, Mt. Surak. Alt. 230 m, 18 viie24 viii 2007 (JO Lim); 86 22 1_, GW, Wongju-si, Heungeup-myeon, Maeji-ri, Yonsei Univ., 26 v 1\, GG, Namyangju-si, Songchoan, Mt. Ungilsan (M.T.) S.N.U., Alt. 87 23 2004 (CY Kim & OB Kang); 1_, GG, Kimpo-gun, Haseong-myeon, 6 99 m, 5e18 ix 2009 (JO Lim); 1\, GG, Anyang-si, Manan-gu, Gwanak, 88 24 vi 2005 (BA Yun); 1_, CB, Danyang-gun, Yeongchun-myeon, Okdae- 5e19 vii 2007 (JO Lim); 1\, CN, Buyeo-gun, Gyuam-myeon, Sumok- 89 25 ri, 5 viii 1994 (SM Ryu); 1_, GB, Pohang-si, Jukjang-myeon, Haok-ri, ri, 27 vie12 vii 2005 (JW Lee); 1\, JN, Naju-si, Dado-myeon, 90 26 12 vi 2004 (IR Sin); 1_, ditto, 29 v 2004; 1_, GB, Pohang-si, Pungsan-ri, 9e30 ix 2005 (SB Yu). 91 27 Cheongha-myeon, Yugye-ri, 5 vi 2004 (IR Sin); 1_, ditto, 29 v Distribution. South Korea. 92 28 2004; 1_, ditto, 23 vi 2004; 1_, GN, Yangsan-si, Wondong-myeon, 93 29 Yeongpo-ri, Sinheungsa, 7 vi 2008 (JM Kwon); 1_, JN, Jeongeup-si, 22. Exochus argutus Tolkanitz, 1993 고운볼록뭉툭맵시벌 94 30 Yongsan-dong, 15 vi 2004 (GB Kim); 2__, JJ, Donggye-myeon, Exochus argutus Tolkanitz, 1993b:106e120. Type: male; type de- 95 31 Jeolmul, 14e28 vii 2005 (CH Sin). pository: SIZ. 96 32 Distribution. South Korea, China, India, Japan, Pakistan, the 97 33 Philippines. Material examined. South Korea: 1_, GN Gajisan, 28 vi 1989 (JG 98 34 Kim). 99 35 Genus Exochus Gravenhorst, 1829 Distribution. South Korea, Belarus, Mongolia, Russia, Ukraine. 100 36 Exochus Gravenhorst, 1829: 328. Type species: Ichneumon gravipes 101 37 Gravenhorst 23. Exochus britannicus Morley, 1911 브리타니볼록뭉툭맵시벌 102 38 Amesolytus Förster, 1869: 161. Type species: Amesolytus ferrugineus Exochus britannicus Morley, 1911:1e344. Type: male; type de- 103 39 Ashmead pository: NHM. 104 40 Mima Davis, 1897: 206. Type species: Mima washingtonensis Davis 105 41 Xanthexochus Morley, 1913: 292. Type species: Xanthexochus scu- Material examined. South Korea: 1_, GW Inje-gun, Bangdaesan, 106 42 tellatus Morley Maehwa-dong, 26 vi 1996 (JW Lee); 1 _, GG Eljeongbu-si, San- 107 43 ghowon, 11 vi 1998 (CY Yoon). 108 17. Exochus acostulatus Lee & Choi, 2016 민무민무늬볼록뭉툭맵시벌 44 Distribution. South Korea, Hungary, Kazakhstan, Moldova, 109 Exochus acostulatus Lee & Choi, 2016, in Choi et al (2016): 2329. 45 Poland, Romania, Russia, Turkey, Turkmenistan, Ukraine, United 110 Type: female; type depository: YNU. 46 Kingdom. 111 47 112 Material examined. South Korea: 1\, GW, Donghae-si, Samhwa- 48 24. Exochus carinalis Lee & Choi, 2016 줄무늬볼록뭉툭맵시벌 113 dong, Mureung valley (M.T.), 31 ve15 vi 2005 (JW Lee); 1\, GB, 49 Exochus carinalis Lee & Choi, 2016, in Choi et al (2016): 2334. Type: 114 Uljin-gun, Sangsa-ri, 24 v 1989 (JG Kim). 50 female; type depository: YNU. 115 Distribution. South Korea. 51 116 52 Material examined. South Korea: 1\, GB, Gyeongsan-si, Dae- 117 18. Exochus adentatus Lee & Choi, 2016 가시볼록뭉툭맵시벌 53 dong, Yeungnam Univ., 18 v 1989 (EA Hong); 1_, GB, Gyeongsan-si, 118 Exochus adentatus Lee & Choi, 2016, in Choi et al (2016): 2331. Type: 54 Dae-dong, Yeungnam Univ., 31 v 1988 (JH Kwon). 119 female; type depository: YNU. 55 Distribution. South Korea. 120 56 121 Material examined. South Korea: 1\, GW, Wonju-si, Baegun- 57 25. Exochus consimilis Holmgren, 1858 어리볼록뭉툭맵시벌 122 mycon, Baegunsan, 11 ve12 vi 2007 (JW Lee); 1\, JN, Gurye-gun, 58 Exochus consimilis Holmgren, 1858: 305e394. Type: unknown; type 123 Gurye-eup, Nogodan, Jirisan National park (M.T.), 2 vie10 x 2001 59 depository: unknown. 124 (JC Jeong). 60 Exochus parvispina Thomson, 1887:193e218. Type: unknown; type 125 Distribution. South Korea. 61 depository: ZI. 126 62 Exochus decoloratus Schmiedeknecht, 1924:103e112. Type: un- 127 19. Exochus alpinus (Zetterstedt, 1838) 알프스볼록뭉툭맵시벌 63 known; type depository: Germany. 128 Bassus alpinus Zetterstedt, 1838: 358e408. Type: female; type de- 64 Exochus subalpinus Schmiedeknecht, 1924:103e112. Type: un- 129 pository: ZI. 65 known; type depository: Germany. 130

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1 Material examined. South Korea: 2__, GW, Goseong-gun, Gan- 31. Exochus gratus Tolkanitz, 2003 작은작은볼록뭉툭맵시벌 66 2 seong-eup, Heyangrobong, 13 v 1992 (SM Ryu); 1_, CB, Danyang- Exochus gratus Tolkanitz, 2003a: 1075e1085. Type: female; Type 67 3 gun, Danyang-eup, Cheongdong-ri, Cheongdong valley, 19e30 iv depository: ZIN. 68 4 2007 (JW Lee). 69 5 Distribution. South Korea, Austria, Belarus, Belgium, Bulgaria, Material examined. South Korea: 1_, GB, Cheongdo-gun, Unmun- 70 6 Czech Republic, Finland, France, Georgia, Germany, Greenland, myeon, Unmunsan U2, 30 ve16 vi 2009 (JW Lee). 71 7 Hungary, Kazakhstan, Latvia, Lithuania, Moldova, Mongolia, the Distribution. South Korea, Russia. 72 8 Netherlands, Norway, Poland, Romania, Russia, Slovakia, Spain, 73 9 Sweden, Tunisia, Turkey, Ukraine, United Kingdom. 32. Exochus gravipes (Gravenhorst, 1820) 큰볼록뭉툭맵시벌 74 10 Ichneumon gravipes Gravenhorst, 1820: 384. 75 11 26. Exochus dentisternum Lee & Choi, 2016 배가시볼록뭉툭맵시 Exochus gravipes Gravenhorst, 1820: 351. Lectotype: female; type 76 12 벌 depository: IZU. 77 13 Exochus dentisternum Lee & Choi, 2016, in Choi et al (2016): 2335. 78 14 Type: male; type depository: YNU. Material examined. South Korea: 1\, GN Gajisan, 25 vi 1989 (JK 79 15 Kim); 1 \, JB Jirisan Sundeuryu, 10 vi 1989 (JK Kim). 80 16 Material examined. South Korea: 1_, GW, Hongcheon-gun, Buk- Distribution. South Korea, Austria, Belarus, Belgium, Bulgaria, 81 17 bang, Gwangwon Prov. Environment Research Park, 15e30 v 2012 Canada, China, Croatia, Czech Republic, Denmark, Estonia, Finland, 82 18 (SJ Jang). France, Germany, Hungary, Iran, Ireland, Italy, Latvia, Lithuania, the 83 19 Distribution. South Korea. Netherlands, Norway, Poland, Romania, Russia, Spain, Sweden, 84 20 Switzerland, Turkey, USA, Ukraine, United Kingdom. 85 21 27. Exochus depressus Lee & Choi, 2016 납작볼록뭉툭맵시벌 86 22 Exochus depressus Lee & Choi, 2016, in Choi et al (2016): 2336. Type: 33. Exochus guttatus Tolkanitz, 1999 얼룩볼록뭉툭맵시벌 87 23 female; type depository: YNU. Exochus guttatus Tolkanitz, 1999:191e201. Type: male; type de- 88 24 pository: ZIN. 89 25 Material examined. South Korea: 1\, GG, Namyangju-si, Choan- 90 26 myeon, Songchon-ri, Mt. Ungilsan (M.T.), Alt. 134 m, 1e26 v 2009 Material examined. South Korea: 1_, GB Youngyang-gun, Subi- 91 27 (JO Lim). myeon, Yeonhwasa (Eulryeonsan), 26 vii 2001 (JS Jeon). 92 28 Distribution. South Korea. Distribution. South Korea, Moldova, Russia. 93 29 94 30 28. Exochus destitutus Tolkanitz, 2003 강원볼록뭉툭맵시벌 34. Exochus kusigematii Tolkanitz, 2007 영남볼록뭉툭맵시벌 95 31 Exochus destitutus Tolkanitz, 2003b: 1211e1214. Type: female; type Exochus kusigematii Tolkanitz, 2007a: 348. 96 32 depository: ZIN. Exochus erythropus Kusigematii, 1971: 279. Type: female; type de- 97 33 pository: HU. 98 34 Material examined. South Korea: 1_, GW Inje-gun, Bangdaesan, 99 35 Maehwa-dong, 26 vi 1996 (JW Lee); 1_, GG Eljeongbu-si, Sangho- Material examined. South Korea: 1\, GB Gyongsan-si, Yeungnam 100 36 won, 11 vi 1998 (CY Yoon). Univ., 1 vii 1986 (JW Lee); 1\, GN Jinju-si, Gajoa-dong, 8 vii 1980 (JK 101 37 Distribution. South Korea, Russia. Kim); 1\, GN Ulju-gun, Sangbuk-myeon, 28 vi 1989 (JK Kim). 102 38 Distribution. South Korea, Japan, Russia. 103 39 29. Exochus ﬂavomarginatus Holmgren, 1856 노랑테볼록뭉툭 104 40 맵시벌 35. Exochus lictor Haliday, 1838 할리볼록뭉툭맵시벌 105 41 Exochus ﬂavomarginatus Holmgren, 1856: 75. Type: male, lost. Exochus lictor Haliday, 1838:112e121. Type: female; type de- 106 42 pository: NMI. 107 43 Material examined. South Korea: 1_, GB Gyongsan-si, Yeungnam Exochus decoratus Holmgren 1873:55e78. Type: lost. 108 44 Univ., 22 v 1990 (JR Lee); 1_, GN Mylyang-si, Cheonhwangsan Exochus pectoralis Haliday 1838:112e121. Type: male; type de- 109 45 Sajapyeong, 23 vi 1989 (JK Kim); 1_, GN Hapcheon-gun, Haeinsa, 15 pository: NMI. 110 46 v 1987. 111 47 Distribution. South Korea, Austria, Azerbaijan, Belgium, Czech Material examined. South Korea: 5__, GG, Cheonggyesan, 4 vii 112 48 Republic, Finland, France, Georgia, Germany, Hungary, Ireland, Italy, 1989 (JK Kim); 1_,ditto,4vi1989(JKKim);1_, GG Annyang-si, 113 49 Japan, Lithuania, Moldova, Mongolia, the Netherlands, Norway, Manan-gu, Gwanak, N 372501400E1275700100,5e19 vii 2007 (JO 114 50 Poland, Romania, Russia, Spain, Sweden, Turkey, Ukraine, United Lim); 1_, GG Anyang-si, Manan-gu, Gwanaksan, 26 vie4vii2007 115 51 Kingdom. (JO Lim); 1\, GW Wonju-si, Heungeom-myeon, Maeji-ri, Yeonse 116 52 Univ., Wonju campus, N 371605300E1275400200,21e30 2009 117 53 30. Exochus frontellus Holmgren, 1858 이마볼록뭉툭맵시벌 (HY Han); 1_, GW Taebaeksan, 13 viii 1989 (JK Kim); 1\,GWMt. 118 54 Exochus frontellus Holmgren, 1858: 305e394. Type: unknown; type Baegun, Wonju-si, Panbu-myeon, 9 vie5vii2011(JWLee);1_,GB 119 55 depository: unknown. Cheongsong-gun Kekwangsan, 25 v 1989 (JK Kim); 1\,GN 120 56 Exochus shimamatsensis Kusigemati, 1971: 205e298. Type: female; Yeongju-si, Punggi-eup, Jungnyeong 99site, 12 vie23 vii 2008 121 57 type depository: HU. (KB Kim); 1_,JBJirisan,Bamsagol,6vi1983;5__,JNGohung- 122 58 gun, Jeomam-myeon, Pallyeongsan (M.T.), 24 ve11 vii 2012 (JC 123 59 Material examined. South Korea: 1\, GG, Anyang-si, Naman-gu, Jeong). 124 60 Gwanak, 5e19 vii 2007 (JO Lim); 1\, GB, Andong-si, Imdong-myeon, Distribution. South Korea, Azerbaijan, Belarus, Belgium, 125 61 Gocheon-ri, Aegisan, 16 viii 1997 (JW Lee); 1\, CB, Cheongwon-si, Bulgaria, Canada, Czech Republic, Finland, France, Germany, 126 62 Miwon-myeon, Miwon-ri, 9e16 ix 2008 (JH Han). Hungary, Ireland, Italy, Japan, Latvia, Lithuania, the Netherlands, 127 63 Distribution. South Korea, Austria, Germany, Hungary, Japan, Poland, Romania, Russia, Spain, Turkey, USA, Ukraine, United 128 64 Romania, Russia, Sweden, Ukraine, United Kingdom. Kingdom. 129 65 130

6 JK Choi, JW Lee / Journal of Asia-Paciﬁc Biodiversity xxx (2016) 1e19

1 36. Exochus lineifrons Thomson, 1887 이마줄무늬볼록뭉툭맵시 42. Exochus orbitalis Lee & Choi, 2016 내연볼록뭉툭맵시벌 66 2 벌 Exochus orbitalis Lee & Choi, 2016, in Choi et al (2016): 2341. Type: 67 3 Exochus lineifrons Thomson, 1887:193e218. Type: unknown; type female; type depository: YNU. 68 4 depository: ZI. 69 5 Material examined. South Korea: 1\, CB, Dangyang-gun, 70 6 Material examined. South Korea: 1\, CN, Seosan-si, Haemi- Danyang-eup, Cheondong-ri, Mt. Sobeak birobong, 22 v 1987 71 7 myeon, Daegok-ri, Hanseo Univ., 7 v 2013 (JG Kim); 1\, JN, (M.T.) (JW Lee). 72 8 Jeongeup-si, Jangseong-gun, Bukha-myeon, Namohanggol, 22 vi Distribution. South Korea. 73 9 2005 (JG Han). 74 10 Distribution. South Korea, Finland, Georgia, Germany, Hungary, 43. Exochus propodealis Lee & Choi, 2016 배고랑볼록뭉툭맵시 75 11 Moldova, Poland, Russia, Sweden, Turkey, Ukraine. 벌 76 12 Exochus propodealis Lee & Choi, 2016, in Choi et al (2016): 2342. 77 13 37. Exochus mitratus Gravenhorst, 1829 큰뿔볼록뭉툭맵시벌 Type: female; type depository: YNU. 78 14 Exochus mitratus Gravenhorst, 1829: 350. 79 15 Exochus affinis Holmgren 1858: 305e394. Material examined. South Korea: 1\, GW, Hongcheon-gun, Buk- 80 16 Exochus australis Thomson 1894: 2137. Lectotype: female; type bong, Environment Research Park, 15e30 v 2012 (SJ Jang). 81 17 depository: ZI. Distribution. South Korea. 82 18 Exochus pseudaffinis Strobl 1903:3e100. Type: male. 83 19 Exochus paradoxus Schmiedeknecht 1924:103e112. 44. Exochus selenanae Tolkanitz, 1999 잎말이볼록뭉툭맵시벌 84 20 Exochus punctifer Schmiedeknecht 1924:103e112. Type: female. Exochus selenanae Tolkanitz, 1999:191e201. Type: female; type 85 21 depository: SIZ. 86 22 Material examined. South Korea: 1_, GB Gyongsan-si, Yeungnam 87 23 Univ., 4 v 1988 (JH Kwon); 1_, ditto, 29 iv 1989 (KJ Hwang). Material examined. South Korea: 1\, GW, Donghae-si, Sanhwa- 88 24 Distribution. South Korea, Austria, Azerbaijan, Belgium, Bulgaria, dong, Mureung valley (M.T.), 21e30 v 2005 (JW Lee); 1\, GB, 89 25 Canada, Canary Islands, China, Croatia, Czech Republic, Finland, Cheongdo-gun, Unmun-myeon, Unmunsan U2, 23 vi 2008 (JW 90 26 France, Georgia, Germany, Hungary, Iran, Ireland, Italy, Japan, Lee); 1\, GB, Yeongju-si, Punggi-eup, Jungyeong 99 site, 12 vie23 vii 91 27 Kazakhstan, Latvia, Lithuania, Malta, Moldova, Mongolia, the 2008 (KB Kim). 92 28 Netherlands, Norway, Poland, Portugal, Romania, Russia, Slovenia, Distribution. South Korea, Moldova, Russia, Ukraine. 93 29 Spain, Sweden, Tunisia, Turkey, USA, Ukraine, United Kingdom, 94 30 Yugoslavia. 45. Exochus semilividus Vollenhoven, 1875 설악볼록뭉툭맵시벌 95 31 Exochus semilividus Vollenhoven, 1875:1e16. Type: unknown; type 96 32 38. Exochus momoii Tolkanitz, 2007 모모모모이볼록뭉툭맵시벌 depository: unknown. 97 33 Exochus momoii Tolkanitz, 2007a: 343. 98 34 Exochus affinis Momoi and Kusigemati, 1970: 414. Type: male; type Material examined. South Korea: 3\\, GG Cheonggyesan, 4 vii 99 35 depository: HU. 1989 (JK Kim); GW Soraksan Heungryeongsungsen, 18 viii 1987 (JK 100 36 Kim); 1\, Donghae-si, Samhwa-dong, Mureung valley (M.T.), N 101 37 Material examined. South Korea: 2__, Seoul, Korea Univ., 372705200E1290102600,27ixe5 x 2007; 1\, GB Ulreung, Naribunji, 102 38 Imyeupsi, SukwoanD, 1 viii 1987 (IC H); 3__, JB Namwon-si, San- 17 x 2001 (JC Jeong); 1\, JB Jeongeup-si, Yongsan-dong, N 35300E 103 39 nae-myeon, Baemsagol, N 352105500E1273405800, 31 vii 1992. 126510, 19 v 2004 (JW Lee). 104 40 Distribution. South Korea, Japan, Russia. Distribution. South Korea, Armenia, Austria, Azerbaijan, Belgium, 105 41 Bulgaria, France, Georgia, Germany, Hungary, Italy, Latvia, 106 42 39. Exochus nigrifaciatus Momoi, Kusigemati & Nakanishi, Lithuania, the Netherlands, Poland, Russia, Sweden, Switzerland, 107 43 1968 검은얼굴볼록뭉툭맵시벌 Turkey, Ukraine. 108 44 Exochus nigrifaciatus Momoi, Kusigemati & Nakanishi, 1968:211. 109 45 Type: female; type depository: KU. 46. Exochus suborbitalis Schmiedeknecht, 1924 금양볼록뭉툭 110 46 맵시벌 111 47 Material examined. South Korea: 1\1_, GB Gyongsan-si, Yeungnam Exochus suborbitalis Schmiedeknecht, 1924:103e112. Type: un- 112 48 Univ., 13 ix 1988 (JK Kim); 1_, Woljeongsa, 30 vii 1981 (YJ Cheon). known; type depository: unknown. 113 49 Distribution. South Korea, Japan, Russia. 114 50 Material examined. South Korea: 1_, GW, Chuncheon-si, Sang- 115 51 40. Exochus nigritulus Lee & Choi, 2016 검정볼록뭉툭맵시벌 dong, Gwangwon Prov. Arboretum, Alt. 81 m, 3755023.700N, 116 52 Exochus nigritulus Lee & Choi, 2016, in Choi et al (2016): 2338. Type: 12743031.700E, 14e27 vi 2012 (GY Lee); 2__, GB, Gunwi-gun, Goro- 117 53 female; type depository: YNU. myeon, Geumyang-ri, 14 v 2003; 1\, GB, Gyeongsan-si, Dae-dong, 118 54 Yeungnam Univ., 16e25 ix 2006 (JW Lee); 1\, ditto, 29 ive7 v 2008; 119 55 Material examined. South Korea: 1\, GN, Hadong-gun, Hwagae- 1\, ditto, 12 viie2 viii 2011; 1\, GB, Cheongdo-gun, Gakbuk-myeon, 120 56 myeon, Beumwang-ri, Jirisan, Chilbulam, 20 v 1989 (JG Kim). Namsan3-ri, 7 vie12 vii 2009, JW Lee; 1\, H., 1 ive9v1991. 121 57 Distribution. South Korea. Distribution. South Korea, Mongolia, Bulgaria, Czech Republic, 122 58 France, Germany, Hungary, Kazakhstan, Latvia, Lithuania, Moldova, 123 59 41. Exochus occipitalis Lee & Choi, 2016 뒷머리볼록뭉툭맵시벌 Poland, Romania, Russia, Turkey, Ukraine. 124 60 Exochus occipitalis Lee & Choi, 2016, in Choi et al (2016): 2340. Type: 125 61 female; type depository: YNU. 47. Exochus thomsoni Schmiedeknecht, 1924 톰슨볼록뭉툭맵시 126 62 벌 127 63 Material examined. South Korea: 1\, GW, Hongcheon-gun, Buk- Exochus crassicornis Thomson 1894: 2080e2137. Type: lost. 128 64 bong, Environment Research Park, 15e29 vi 2009 (SJ Jang). Exochus thomsoni Schmiedeknecht, 1924:103e112. Type: un- 129 65 Distribution. South Korea. known; type depository: unknown. 130

JK Choi, JW Lee / Journal of Asia-Paciﬁc Biodiversity xxx (2016) 1e19 7

1 Material examined. South Korea: 1\, GW, Hongcheon-gun, Buk- Material examined. South Korea: 1_, CN, Dangjin-gun, Sin- 66 2 bang, Gwangwon Prov. Environment Research Park, 15e29 vi 2009 pyeong-myeon, Geumcheon-ri, 10 ix 2006 (HS Lee); 1\, CN, Seosan- 67 3 (SJ Jang); 1\, GB, Cheongdo-gun, Gakbuk-myeon, Namsan3-ri, si, Haemi-myeon, Daegok-ri, Hanseo Univ., 11e23 vi 2009 (JW Lee); 68 4 35410N, 128350E, 29 vie6 vii 2008 (JW Lee); 1\, ditto, 28 ive27 v 1_, ditto, 9e21 x 2006 (JG Kim); 1\, GB, Cheongdo-gun, Unmun- 69 5 2013; 1_, ditto, 20 vi 2007; 1\, ditto, 29 vie6 vii 2008; 1\, GB, myeon, Mt. Unmunsan, 6e19 vi 2008 (JW Lee); 1_, ditto, 24e30 v 70 6 Gyeongsan-si, Dae-dong, Yeungnam Univ., 17e24 iv 2009 (JW Lee); 2009 (HJ Kwon); 1_, GB, Cheongdo-gun, Unmun-myeon, Unmunsa, 71 7 1\, ditto, 12 v 2010; 1\, GB, Gunwi-gun, Goro-myeon, Geumyang, 14 28 v 1989 (JG Kim); 1_, GB, Gyeongsan-si, Dae-dong, Yeongnam 72 8 v 2003; 1\, CB, Jecheon-si, Hansu-myeon, Songgye-ri, Woraksan Univ., 29 v 1989 (GJ Hwa); 1\, GG, Anyang-si, Manan-gu, Kwanag, 73 9 National Park, 20 ve3 vi 2006 (JC Jeong). 9e24 vi 2007 (JO Lim); 1\, GN, Jinju-si, Gajwa-dong, 26 ve2 vi 1987 74 10 Distribution. South Korea, China, Finland, Hungary, Kazakhstan, (JS Park); 1\, GW, Donghae-si, Samhwa-dong, Mureung valley, 21e 75 11 Mongolia, the Netherlands, Poland, Romania, Russia, Sweden, 30 v 2005 (JW Lee); 1\, JJ, Jeju-si, Ara1-dong, Jeju Univ., Alt. 333 m, 76 12 Turkey, Ukraine 1e10 vi 2009 (JW Lee). 77 13 Distribution. South Korea, Japan 78 14 48. Exochus velatus Tolkanitz, 2003 숨은볼록뭉툭맵시벌 79 15 Exochus velatus Tolkanitz, 2003a: 1075e1085. Type: female; type 53. Hypsicera brevicornis Momoi & Kusigemati, 1970 짧은뿔수 80 16 depository: ZIN. 중다리뭉툭맵시벌 81 17 Hypsicera brevicornis Momoi & Kusigemati, 1970: 413. Type: male; 82 18 Material examined. South Korea: 1_, GG Cheonggyesan, 4 vi 1989 type depository: HU. 83 19 (JK Kim); 1_, GB Jinryaong, Geumgok, 5 vii 1981 (JK Kim). 84 20 Distribution. South Korea, Russia. Material examined. South Korea: 1\, Seoul, Nowon-gu, Sanggye- 85 21 dong, Mt. Suraksan, 18 viie24 viii 2007 (JO Lim). 86 22 49. Exochus ventralis Holmgren, 1858 배볼배볼록뭉툭맵시벌 Distribution. South Korea, Japan, Russia. 87 23 Exochus ventralis Holmgren, 1858: 305e394. Lectotype: female; 88 24 type depository: NR. 54. Hypsicera carinata Momoi & Kusigemati, 1970 등줄수중다 89 25 리뭉툭맵시벌 90 26 Material examined. South Korea: 1_, GG, Ganghwa-do, Hwado, Hypsicera carinata Momoi & Kusigemati, 1970: 412. Type: male; 91 27 Manisan, Heungwang-ri, 24 ix 1994 (HS Won); 1_, GN Jinju-si, type depository: HU. 92 28 Gajoa-dong, 9 vi 1987 (JK Kim); 1\, ditto, 20 vi 1989 (JK Kim). 93 29 Distribution. South Korea, Austria, Belgium, Czech Republic, Material examined. South Korea: 1_, Daejeon, Dong-gu, Daejeon 94 30 France, Poland, Romania, Russia, Sweden, Ukraine. Univ., 16 ve5 vi 2006 (JW Lee); 2_, ditto, 8 xe30 xi 2007 (JW Lee); 95 31 2\3_, GB, Cheongdo-gun, Unmun-myeon, Mt. Unmunsan, Sim- 96 32 50. Exochus vexator Tolkanitz, 1993 악마볼록뭉툭맵시벌 simgyegok, 29 viiie19 ix 2014 (JW Lee); 1_, GB, Gunwi-gun, Bugye- 97 33 Exochus vexator Tolkanitz, 1993a:92e105. Type: female; type de- myeon, Dongsan-ri, Odoam, 15 viie29 viii 2014 (JW Lee); 1_, GB, 98 34 pository: SIZ. Yeongcheon-si, Cheogntong-myeon, Eunhaesa, 9e22 v 2015 (JW 99 35 Lee); 1\, GG, Anyang-si, Manan-gu, Kwanag, 26 viie7 viii 2008 (JO 100 36 Material examined. South Korea: 1\, JB Jirisan National Park, Lim); 1_, ditto, 26 vie4 vii 2007 (JO Lim); 1_, ditto, 15 viie25 vii 101 37 Nogodan, 15 v 1983 (KO H); 1\, ditto, 22 vii 1989 (JK Kim). 2008 (JO Lim); 1_, GG, Namyangju-si, Choan-myeon, Songcheon-ri, 102 38 Distribution. South Korea, Azerbaijan, Hungary, Moldova, Russia, Mt. Ungilsan, Alt. 99 m, 18 viiie14 ix 2009 (JO Lim); 1_, GN, 103 39 Ukraine. Kwangju-si, Buk-gu, Seumgok-dong, Mt. Mudeungsan, Wonhyosa, 104 40 26 vie27 vii 2013 (JK Choi); 1_, GN, Sancheong-gun, Samjang- 105 41 51. Exochus villosus Tolkanitz, 2003 털보볼보볼록뭉툭맵시벌 myeon, Jangdang valley, 24 viii 2011 (JC Jeong); 1\1_, GN, 106 42 Exochus villosus Tolkanitz, 2003a: 1075e1085. Type: female; type Sancheong-gun, Sicheon-myeon, Jungsan-ri, Mt. Jirisan, Sunduryu, 107 43 depository: ZIN. 9 viii 1989 (JG Kim); 1\1_, GW, Donghae-si, Samhwa-dong, Mur- 108 44 eung valley, 21e30 v 2005 (JW Lee); 1_, GW, Heungeop-myeon, 109 45 Material examined. South Korea: 1\, GW, Hongcheon-gun, Maeji-ri, Yeonsei Univ., 29 vie25 vii 2011 (JW Lee); 1_, GW, Inje- 110 46 Hongcheon-eup, Jangjeonpyeong1-ri, Geomdungae village (M.T.), gun, Mt. Bangdaesan, 24 vi 1996 (JW Lee); 1\, GW, Wonju-si, 111 47 E127520N37390,1e20 vi 2007 (JW Lee); 1\, GW, Hongcheon-gun, Heungeop-myeon, Maeji-ri, Yeonsei Univ., 1e31 vii 2010 (HY 112 48 Bukbang, Environment Research Park, E 1275101.700N3545015.600, Han); 1_, JB, Buan-gun, Byeonsanbando, Ganbeolii, 4 ive25 vii 2008 113 49 29 vie16 vii 2012 (SJ Jang). (JC Jeong); 3_, JB, Jeongeup-si, Naejang-dong, Mt. Naejangsan, 114 50 Distribution. South Korea, Russia. Wonjeokam, 22 viiie13 ix 2006 (JW Lee). 115 51 Distribution. South Korea, Japan. 116 52 Genus Hypsicera Latreille, 1829 117 53 Hypsicera Latreille, 1829: 288. Type species: Alomya sp. near femo- 55. Hypsicera femoralis (Geoffroy, 1785) 넓적수중다리뭉툭맵시 118 54 ralis Gravenhorst (¼Hypsicera femoralis) Monobasic. 벌 119 55 Q2 Metacoelus Förster, 1868: 161. Type species: Exochus femoralis Ichneumon femoralis Geoffroy, 1785: 396. Type: female; type de- 120 56 Gravenhorst (¼Hypsicera femoralis). pository: MHN. Lost. 121 57 Polyclistus Förster, 1868: 161. Type species: Ichneumon femoralis Exochus lævis Cresson, 1864: 286. Lectotype: male; type depository: 122 58 Fourcroy (¼Hypsicera femoralis). ANSP. 123 59 Plesioexochus Cameron, 1905: 202. Type species: Plesioexochus Crypturus bonaerensis Schrottky, 1902: 100. Type: male. Lost. 124 60 rufipes Cameron (¼Hypsicera femoralis) Monobasic. Plesioexochus rufipes Cameron, 1905: 203. Type: female; type de- 125 61 pository: NHM. 126 62 52. Hypsicera bicolor Momoi & Kusigemati, 1970 두색수중다리 Exochus platensis Brèthes, 1924: 10. Type: female; type depository: 127 63 뭉툭맵시벌 MACN. 128 64 Hypsicera bicolor Momoi & Kusigemati, 1970: 406. Type: female; Exochus amicus ruficoxis Seyrig, 1934: 45. Lectotype: male; type 129 65 type depository: HU. depository: MNHN. 130

8 JK Choi, JW Lee / Journal of Asia-Paciﬁc Biodiversity xxx (2016) 1e19

1 Material examined. South Korea: 1_, GB, Cheongdo-gun, Unmun- Wonju-si, Panbu-myeon, Mt. Baegunsan, 30 viie28 viii 2013 (JW 66 2 myeon, Mt. Unmunsan, 1viie24 vii 2008 (JW Lee); 1_, GB, Gunwi- Lee); 2\\, GW, Wonju-si, Socho-myeon, Hakgong-ri, Mt. Chiaksan, 67 3 gun, Bugye-myeon, Dongsan-ri, Odoam, 15 viie29 viii 2014 (JW 20 vie19 vii 2013 (JW Lee). 68 4 Lee); 1\, GB, Gyeongsan-si, Dae-dong, Yeongnam Univ., 21 ive19 v Distribution. South Korea, Japan. 69 5 2004 (JW Lee); 1_, GB, Yeongyang-gun, Mt. Heungrimsan, 20 vii 1997 70 6 (JW Lee); 3\\2__, GN, Changnyeong-gun, Yueo-myeon, Daedae-ri, 59. Hypsicera makiharai Kusigemati, 1971 긴뺨수중다리뭉툭맵 71 7 Uponeup, 5e25 v 2015 (JW Lee); 2\\, GN, Jinju-si, Gajwa-dong, 시벌 72 8 18e24 v 1990 (JS Park); 1\,ditto,7e14 vii 1987; 1_, GN, Sancheong- Hypsicera makiharai Kusigemati, 1971: 257. Type: male; type de- 73 9 gun, Sicheon-myeon, Mt. Jirisan,10 viii 1989 (YJ Kwon); 1\, GN, Ulju- pository: HU. 74 10 gun, Sangbuk-myeon, Icheon-ri, 6e13 viii 1989 (JG Kim); 1\,GW, 75 11 Donghae-si, Samhwa-dong, Mureung valley, 2e15 v 2007 (JW Lee); Material examined. South Korea: 1_, GB, Cheongdo-gun, Unmun- 76 12 1\,ditto,5e25 x 2007 (JW Lee); 3\\, JN, Gurye-gun, Masan-myeon, myeon, Mt. Unmunsan, Haksodaepokpo, 22 vie28 vi 2014 (JW Lee); 77 13 Hwangjeon-ri, Hwaeom valley, 15 viie18 x 2011 (JC Jeong); 1_,ditto, 1_, GN, Millyang-si, Danjang-myeon, Sajapyeong, 25 vi 1987 (JG 78 14 9vie15 vii 2011 (JC Jeong); 1\1_, JN, Gurye-gun, Toji-myeon, Piagol, Kim); 1_, GN, Ulju-gun Sangbuk-myeon Icheon-ri, 25 vi 1987 (JG 79 15 Jirisan National Park, 10 viie15 vii 2011 (JC Jeong). Kim); 2__, GW, Wonju-si, Panbu-myeon, Mt. Baegunsan, 30 viie28 80 16 Distribution. South Korea, Argentina, Armenia, Australia, Austria, viii 2013 (JW Lee). 81 17 Azerbaijan, Azores, Belarus, Belgium, Brazil, Bulgaria, Canada, Ca- Distribution. South Korea, Japan. 82 18 nary, Chile, China, Croatia, Czech Republic, Finland, France, Georgia, 83 19 Germany, Greece, Hungary, Ireland, Italy, Japan, Kazakhstan, Latvia, 60. Hypsicera nigribasis Momoi & Kusigemati, 1970 꼬마수중다 84 20 Lithuania, Madagascar, Moldova, the Netherlands, New Zealand, 리뭉툭맵시벌 85 21 Norway, Poland, Portugal, Romania, Russia, South Africa, Spain, Hypsicera makiharai Momoi & Kusigemati, 1970: 411. Type: male; 86 22 Sweden, Switzerland, Turkey, USA, Ukraine, United Kingdom. type depository: MOMOI. 87 23 88 24 56. Hypsicera harrelli Momoi & Kusigemati, 1970 운문수중다리 Material examined. South Korea: 1_, GB, Cheongdo-gun, Unmun- 89 25 뭉툭맵시벌 myeon, Mt. Unmunsan, Haksodaepokpo, 29 viiie19 ix 2014 (JW Lee); 90 26 Hypsicera harrelli Momoi & Kusigemati, 1970: 410. Type: female; 1_, GB, Cheongdo-gun, Unmun-myeon, Mt. Unmunsan, Simsimgye- 91 27 type depository: BBM. gok, 29 viiie19 ix 2014 (JW Lee); 1_, GB, Cheongdo-gun, Unmun- 92 28 myeon, Unmunsa, 6 vie1 vii 2008 (JW Lee); GB, Choilgok-gun, 93 29 Material examined. South Korea: 1\, GB, Cheongdo-gun, Unmun- Dongmyeong-myeon, Hakmyeong-ri, Gasansanseong, 30 viiie22 ix 94 30 myeon, Mt. Unmunsan, 24 viie15 viii 2008 (JW Lee); 1\, GB, 2014 (JW Lee); 4__, GB, Gunwi-gun, Bugye-myeon, Dongsan-ri, 95 31 Cheongdo-gun, Unmun-myeon, Mt. Unmunsan, Haksodaepokpo,12 Odoam, 15 viie29 viii 2014 (JW Lee); 1_, GN, Sancheong-gun, 96 32 viiie22 ix 2012 (JW Lee); 1\, GB, Cheongdo-gun, Unmun-myeon, Mt. Sicheon-myeon, Jungsan-ri, Mt. Jirisan, Sunduryu, 24 vi 1989 (JG 97 33 Unmunsan, Simsimgyegok, 30 viie29 viii 2014 (JW Lee); 1\1_, GG, Kim); 1_, GN, Sancheong-gun, Sicheon-myeon, Mt. Jirisan,15 viiie21 98 34 Anyang-si Manan-gu Kwanag, 26 vie4 vii 2007 (JO Lim); 1\, GN, viii 1989 (JW Lee); 1\, GW, Donghae-si, Samhwa-dong, Mureung 99 35 Geochang-gun Wicheon-myeon Sangcheon-ri, 4 vi 2014 (JK Choi); valley, 2 ve15 v 2007 (JW Lee); 1_, JN, Gurye-gun, Toji-myeon, Piagol 100 36 2\\, GN, Hadong-gun, Hwagae-myeon, Mt. Jirisan, Daeseonggol, 29 Jirisan National Park,10 iiie15 vi 2010 (JW Lee); 1_, Seoul, Nowon-gu, 101 37 vii 1987 (JG Kim); 1\, GN, Sancheong-gun, Sicheon-myeon, Jungsan- Sanggye-dong, Mt. Seoraksan, 18 viie24 vii 2007 (JW Lee). 102 38 ri, Mt. Jirisan, Sunduryu, 28 vi 1989 (JG Kim); 1\, GW, Wonju-si, Distribution. South Korea, Japan. 103 39 Panbu-myeon, Mt. Baegunsan, 30 viie28 viii 2013 (JW Lee). 104 40 Distribution. South Korea, Japan, Russia 61. Hypsicera postfurcalis Kusigemati, 1971 두줄수중다리뭉툭 105 41 맵시벌 106 42 57. Hypsicera incarinata Momoi & Kusigemati, 1970 민무민무늬수 Hypsicera makiharai Kusigemati, 1971: 260. Type: female; type 107 43 중다리뭉툭맵시벌 depository: HU. 108 44 Hypsicera incarinata Momoi & Kusigemati, 1970: 411. Type: male; 109 45 type depository: BBM. Material examined. South Korea: 1\, Daegu, Dong-gu, Mt. Pal- 110 46 gongsan, 7 vii 1987 (JG Kim). 111 47 Material examined. South Korea: 1_, CB, Boeun-gun, Mt. Song- Distribution. South Korea, Japan. 112 48 nisan, Sanae-ri, Beomjusa, 5 ve31 viii 2011 (JC Jeong); 1_, Daejeon, 113 49 Dong-gu, Daejeon Univ., 12 ve27 v 2007 (JW Lee); 2__, GB, 62. Hypsicera rugosa Kusigemati, 1971 수중다리뭉툭맵시벌 114 50 Cheongdo-gun, Unmun-myeon, Mt. Unmunsan, Simsimgyegok, 29 Hypsicera makiharai Kusigemati, 1971: 255. Type: female; type 115 51 viiie19 ix 2014 (JW Lee); 1_, ditto, 30 viie29 viii 2014 (JW Lee); 1_, depository: HU. 116 52 GB, Gyeongsan-si Dae-dong Yeongnam Univ., 9 vi 1989 (JG Kim); 117 53 1_, GN, Hadong-gun, Hwagae-myeon, Mt. Jirisan, Daeseonggol, 29 Material examined. South Korea: 1_, GW, Panbu-myeon, Mt. 118 54 vii 1989 (JG Kim); 1_, GW, Donghae-si, Samhwa-dong, Mureung Baekunsan, 30 viie28 viii 2013 (JW Lee); 1\, JB, Jeongeup-si, 119 55 valley, 21e30 v 2005 (JW Lee); 1_, JN, Suncheon-si Seungju-eup Bokheung-myeon, Songgotbawi, 19 v 2004 (KB Kim). 120 56 Jukak-ri Seonamsa, 10 vii 1998. Distribution. South Korea, Japan. 121 57 Distribution. South Korea, Japan. 122 58 63. Hypsicera spiracularis Tolkanitz, 1995 구멍수중다리뭉툭맵 123 59 58. Hypsicera intermedia Momoi & Kusigemati, 1970 어리수중 시벌 124 60 다리뭉툭맵시벌 Hypsicera spiracularis Tolkanitz, 1995:40e43, 120e123. Type: fe- 125 61 Hypsicera intermedia Momoi & Kusigemati, 1970: 409. Type: male; male; type depository: ZIN. 126 62 type depository: BBM. 127 63 Material examined. South Korea: 1_, GB, Cheongdo-gun, Unmun- 128 64 Material examined. South Korea: 1\, GW, Wonju-si Heungeop- myeon, Mt. Unmunsan, 24 ve30 v 2009 (JW Lee). 129 65 myeon Maeji-ri Yeonsei Univ., 1 viie31 vii 2010 (HY Han); 1_,GW, Distribution. South Korea, Russia. 130

JK Choi, JW Lee / Journal of Asia-Paciﬁc Biodiversity xxx (2016) 1e19 9

1 64. Hypsicera yoshimotoi Momoi & Kusigemati, 1970 지리산수 Material examined. South Korea: 1_, GG, Anyang-si, Manan-gu, 66 2 중다리뭉툭맵시벌 Gwanak, 372501400N, 1275700100E, 5e19 vii 2007 (JO Lim); 1\, GG, 67 3 Hypsicera makiharai Momoi & Kusigemati, 1970: 407. Type: female; Bogwangsa, 12 vi (GG Sa); 1\, ditto, 9 vi 1978 (WB Lee); 1_, 68 4 type depository: BBM. Euijeongbu-si, Howon-dong, Wonhyosa, 6 vi 1985 (NH Jho); 1_, 69 5 ditto (YH Kim); 1\, Gapyeong-gun, Ha-myeon, Hyeon-ri, 15 ix 1984 70 6 Material examined. South Korea: 1\, Daejeon, Dong-gu, Daejeon (MR Chae); 1_, GG, Gwangju-si, Docheok-myeon, Chugok-ri, Taeh- 71 7 Univ., 8 xe30 xi 2007 (JW Lee); 1_, GB, Cheongdo-gun, Unmun- wasan, Gonziam, 17 viii 1997 (SY Kim). 72 8 myeon, Unmunsa, 1 viii 1989 (JG Kim); 1_, ditto, 28 vi 1989 (JG Distribution. South Korea, Austria, Belgium, China, Croatia, Czech 73 9 Kim); 1\, GG, Anyang-si, Manan-gu, Kwanag, 8 viiie9 viii 2008 (JO Republic, Finland, France, Germany, Greece, Hungary, India, Ireland, 74 10 Lim); 1_, GN, Jinju-si, Gajwa-dong, 18 viiie25 viii 1987; 2__, GN, Israel, Japan, Latvia, Nepal, the Netherlands, Norway, Poland, 75 11 Sancheong-gun, Sicheon-myeon, Mt. Jirisan, (JG Kim); 1_, GN, Ulju- Romania, Russia, Slovakia, Spain, Switzerland, Turkey, Ukraine, 76 12 gun, Sangbuk-myeon, Icheon-ri, 25 vie15 vii 1989 (JG Kim); 1\, JB, United Kingdom, Yugoslavia. 77 13 Jeongeup-si, Naejang-dong, Mt. Naejangsan, Wonjeokam, 22 viiie 78 14 13 ix 2006 (JW Lee); 1\, JN, Gurye-gun, Sandong-myeon, Nogodan, 67. Metopius (Ceratopius) citratus trifasciatus Uchida, 1930 세 79 15 22 viii 1997 (JW Lee). 줄뭉툭맵시벌 80 16 Distribution. South Korea, Japan. Metopius (Ceratopius) imperfectus Uchida, 1930: 251. Type: male; 81 17 type depository: HU. 82 18 Genus Metopius Panzer, 1806 Metopius (Ceratopius) dissectorius trifasciatus Uchida, 1930:251. 83 19 Metopius Panzer, 1806: 271. Type species: Sphex vespoides Scopoli. Type: female; type depository: HU. 84 20 [Metopius vespoides (Scopol)] 85 21 Peltastes Illiger, 1807: 189-199. Type species: Ichneumon necatorius Material examined. Japan: 1\, Sapporo, 25.vii.1925, Uchida. 86 22 Fabricius. [Metopius (Peltastes) necatorius (Fabricius)] Distribution. South Korea, Japan. 87 23 Peltocarus Thomson, 1887: 196. Type species: Metopius (Peltocarus) 88 24 croceicornis Thomson. [Metopius (Peltocarus) croceicornis Thomson] 68. Metopius (Ceratopius) hakiensis Matsumura, 1912 하키뭉툭 89 25 Cultrarius Davis, 1897: 197. Type species: Metopius (Cultrarius) rileyi 맵시벌 90 26 Marlatt. [Metopius (Cultrarius) rileyi Marlatt] Metopius hakiensis Matsumura, 1912:1e247. Type: male; type de- 91 27 Ceratopius Clément, 1927: 3461. Type species: Ichneumon dis- pository: HU. 92 28 sectorius Panzer. [Metopius (Ceratopius) dissectorius (Panzer)] 93 29 Peltopius Clément, 1927: 3461. Type species: Sphex vespoides Material examined. South Korea: 1\, GG, Bogwangsa, 6 vi 1973 (JI 94 30 (Clément). [Metopius vespoides (Scopol)] Kim); 1_, GG, Gwangju-si, Docheok-myeon, Taehwasan (Alt. 153 m), 95 31 Clemontia Michener, 1941: 2. Type species: Ichneumon micratorius 3718042.100N, 12718039.500E, M.T.VI, 28 vie14 vii 2008 (JO Lim); 96 32 Fabricius. [Metopius (Peltocarus) dentatus (Fabricius)] 2\\,1_, Namyangju-si, Pyeongnae-dong, Cheonmasan, 18 vi 1983 97 33 Peltales Townes & Townes, 1959: 91. Type species: Metopius (Pel- (JW Lee); 1_, Seoul, Dobong-gu, Dobong-dong, Dobongsan, 6 vi 98 34 tales) errantia Davis. [Metopius (Peltales) errantia Davis] 1983 (HY Choi); 1\, Seoul, Jungrang-gu, Sinnae-dong, 29 v 1998 (JH 99 35 Tylopius Townes & Townes, 1959: 97. Type species: Metopius (Pel- Kim). 100 36 tastes) pinatorius Brullé. [Metopius (Peltastes) pinatorius Brullé] Distribution. South Korea, Japan. 101 37 Ethimetopius Benoit, 1961:22e26. Type species: Metopius (Ethi- 102 38 metopius) nodiformis (Benoit). [Metopius (Ethimetopius) nodiformis 69. Metopius (Ceratopius) kiushuensis Uchida, 1932 큐슈뭉툭맵 103 39 (Benoit)] 시벌 104 40 Ichneumon dissectorius kiushuensis Uchida, 1932: 166. Type: female; 105 65. Metopius (Metopius) rufus browni Ashmead, 1905 누런줄뭉 41 툭맵시벌 type depository: KU. 106 42 107 Metopius browni Ashmead, 1905: 117. Type: male; type depository: 43 Material examined. South Korea: 1\,1_, GG, Bogwangsa, 24 vi 108 USNM. 44 1978 (WJ Kim); 1\, Seoul, Nowon-gu, Sanggye-dong, Suraksan, 109 Q3 Metopius kakogawanus Matsumura, 1912:100.Type: female; type 45 374004500N, 1270502800E, M.T.VIII, (Alt. 230 m), 18 vie24 viii 2007 110 depository: HU. 46 (JO Lim); 1\, GW, Donghae-si, Samhwa-dong, Murueng valley, M.T., 111 Metopius (Ceratopius) formosanus Clément, 1930: 410. Type: female; 47 372705200N, 1290102600E, 28 viie8 viii 2007; 1_, GW, Wonju-si, 112 type depository: TMA. 48 Heungeop-myeon, Maeji-ri, Yeonsei Univ., Wonju campus, 25 vi 113 49 2007 (HS Lee et al); 1_, ditto, 7 vii 2009 (HY Han et al). 114 Material examined. South Korea: 1\,1_, GW, Wonju-si Heun- 50 Distribution. South Korea, Japan, Russia. 115 geop-myeon, Maeji-ri, Yeonsei Univ., Wonju campus, 14 ix 1999 (TJ 51 116 Jin); 1 _, ditto, 13 ix 1999 (DE Kim); 1\, ditto, 371605300N, 52 70. Metopius (Ceratopius) maruyamensis Uchida, 1930 흰줄뭉 117 1275400200E, (M.T.) 16 viiie16 x 2009 (HY Han); 1_,GW, 53 툭맵시벌 118 Hongcheon-gun, Bukbang-myeon, Seongdong-ri, Jayeonhwan- 54 Metopius (Ceratopius) maruyamensis Uchida, 1930: 249. Type: male; 119 gyeongyeonsu park, 5e14 ix 2011; 1\, Hoengseong-gun, Cheongil- 55 type depository: HU. 120 myeon, Yupyeong-ri, Seomgang river side, the upstream, 8 ix 2002 56 121 (DS Choi et al). 57 Material examined. South Korea: 1_, GG, Cheonan-si, Dongnam- 122 Distribution. South Korea, China, India, Japan, Mongolia, the 58 gu, Gwangdeok-myeon, Gwangdeoksan, 16-18 vi 1994 (JI Kim); 1_, 123 Philippines, Thailand. 59 GG, Yangju-si, Bogwangsa, 29 vi 1975 (SI Lee); 1_, GW, Wonju-si, 124 _ 60 줄 Heungeop-myeon, Maeji-ri, Yeonsei Univ., 24 v 1998; 1 , ditto, 26 125 66. Metopius (Ceratopius) citratus citratus (Geoffroy, 1785) \ 61 뭉툭맵시벌 vi 2007 (HS Lee et al); 1 , GG, Yeongwol-gun, Bogyeongsa, 17 viii 126 62 1990 (JY Cha); 1\, CB, Jecheon-si, Hansu-myeon, Songgye-ri, 127 Ichneumon citratus Fourcroy, 1785: 403. Type: female; type de- 63 Wolaksan, 14 ix 1970 (EJ Jang). 128 pository: lost. 64 Distribution. South Korea, Japan. 129 Ichneumon dissectorius Panzer, 1805: 14. Type: lost. 65 130

10 JK Choi, JW Lee / Journal of Asia-Paciﬁc Biodiversity xxx (2016) 1e19

1 71. Metopius (Ceratopius) scrobiculatus Hartig, 1838 무늬줄뭉 76. Metopius (Peltastes) pocksungi Kim, 1958 복성뭉툭맵시벌 66 2 툭맵시벌 Metopius pocksungi Kim, 1958: 301. Type: female; type depository: 67 3 Metopius scrobiculatus Hartig, 1838: 246e274. Type: male; type YNU. 68 4 depository: ZMHU. 69 5 Material examined. South Korea: 1\, Seoul, Dobong-gu, Dobong- 70 6 Material examined. South Korea: 1_, GG, Gapyeong-gun, Buk- dong, Dobongsan, 10 ix 1957 (GD Lee). 71 7 myeon, Hwaaksan, 11e12 vii 1998 (YT Quan); 1\, GG, Yangpyeong- Distribution. South Korea. 72 8 gun, Seojong-myeon, Suip-ri, Godongsan, 9 ix 1980 (GS Jang); 1_, 73 9 CN, Gongju-si, Gongjujayeonghakseupwon, 11 viii 1995 (JY Cha); 77. Metopius (Peltastes) sapporensis Uchida, 1930 삿포로줄뭉 74 10 1_, GB, Yeongcheon-si, Hwabuk-myeon, Bohyeonsan, 10-11 vii 1998 툭맵시벌 75 11 (SM Ryu); 3__, Yeongju-si, Pupnggi-eup, Sucheol-ri, Sobaeksan, Metopius (Ceratopius) sapporensis Uchida, 1930: 254. Type: female; 76 12 Heuibang valley, 14 vii 1997 (JC Jeong). type depository: HU. 77 13 Distribution. South Korea, Austria, Czech Republic, Germany, 78 14 Kazakhstan, Lithuania, Poland, Russia, Slovakia, Ukraine. Material examined. South Korea: 1\, GB, Cheongdo-gun, Unmun- 79 15 myeon, Unmunsan, u1, 353804500N, 1285703300E, 25 vi-3 vii 2009 80 16 72. Metopius (Peltastes) arakawai Uchida, 1930 천마뭉툭맵시 (JW Lee); None, JB, Muju-gun, Anseong-myeon, Tongan-ri, Deo- 81 17 벌 kyusan, Chilyeon waterfall, 24 vii 1999 (ES Kang). 82 18 Metopius (Ceratopius) arakawai Uchida, 1930: 253. Type: male; type Distribution. South Korea, Japan. 83 19 depository: HU. 84 20 Metopius (Ceratopius) takabayashii Uchida, 1930: 253. Type: female; Genus Periope Haliday, 1838 85 21 type depository: HU. Monoplectron Holmgren, 1856:1e104. Type species: Monoplectron 86 22 Metopius (Ceratopius) temmanensis Uchida, 1955: 132. Type: fe- zygaenator Holmgren. 87 23 male; type depository: HU. Oligoplectron Förster, 1869:135e221. New name for primary 88 24 homonym. 89 25 Material examined. South Korea: 1\, JB, Jeongeup-si, Ssangam- Monoplectrochus Heinrich, 1949:101e127. Type species: Mono- 90 26 dong, Seoraebong maepyoso, 21 vii 2004 (JW Lee); 1\, JN, plectrochus hoerhammer Heinrich. 91 27 Jeongeup-si, Sunchang-gun, Bukheung-myeon, Daegajae, 27 vii 92 78. Periope hoerhammeri (Heinrich, 1949) 어리노랑다리뭉툭맵 28 2004 (JW Lee). 시벌 93 29 Distribution. South Korea, Japan. 94 Monoplectrochus hoerhammeri Heinrich, 1949:101e127. Type: 30 95 male; type depository: AEI. 31 73. Metopius (Peltastes) coreanus Uchida, 1930 참줄뭉툭맵시벌 96 32 Metopius (Ceratopius) coreanus Uchida, 1930: 248. Type: female; 97 Material examined. South Korea: 1_, GG, Bokwangsa, 1 v 1984 33 type depository: HU. 98 (JW Lee). 34 99 Distribution. South Korea, Japan, Germany, Lithuania, Poland, 35 Material examined. South Korea: 1\, Incheon, Ongjin-gun, 100 Russia, Ukraine. 36 Deokjeon-myeon Baekado, 2 viii 1982 (JI Kim). 101 37 Distribution. South Korea, China, Japan, Mongolia. 102 Genus Pseudometopius Davis, 1897 38 103 39 74. Metopius (Peltastes) fossulatus Uchida, 1933 둥근방패뭉툭 Pseudometopius Davis, 1897:193e348. Type species: Metopius 104 40 맵시벌 hagenii Davis. 105 41 Metopius (Metopius) fossulatus Uchida, 1933: 72. Type: male; type Odontotylocomnus Uchida, 1940:175e180. Type species: Odonto- 106 42 depository: HU. tylocomnus pilosus Uchida. 107 43 Tylocomnoides Uchida, 1940:175e180. Type species: Tylocomnoides 108 44 Material examined. South Korea: 1\, GN, Milyang-si, Sannae- egawai Uchida. 109 45 myeon, Samyang-ri, Gajisan, 353701600N, 12900011 00E, 19 v 2001 110 79. Pseudometopius egawai (Uchida, 1940) 어리줄뭉툭맵시벌 46 (JW Lee). 111 Tylocomnoides egawai Uchida, 1940:175e180. Type: female; type 47 Distribution. South Korea, Japan. 112 depository: HU. 48 113 49 75. Metopius (Peltastes) pinatorius Brullé,1846 검붉은줄뭉툭맵 114 Material examined. South Korea: 1\, Seoul, Gangnam-gu, Wonji- 50 시벌 115 dong, 4 vi 1985 (YS Kim); 1\, Seoul, Jeongryeung, 1 vi 1986 (JY Kim); 51 Metopius pinatorius Brullé, 1846: 120. Type: male; type depository: 116 1\, Seoul, Cheonggyesan, 7 vi 1986 (YT Jeong); 1\, Seoul, Bukhan- 52 MNHN. 117 san, 18 v 1985 (JY Kim); 1_, Seoul, Ui-dong, 27 v 1986 (SH Shin); 1\, 53 Metopius micratorius meridionalis Hensch, 1928: 104. Type: female; 118 GG, Soyosan, 28 v 1977 (SH Park); 1_, GG, Cheonmasan, 31 v 1996 54 type depository: IZB. 119 (EK Park); 1_, GG, Bokwangsa, 22 v 1985 (GN Kim); 1\, GB, 55 Metopius (Metopius) gracilis Clément, 1930: 397. Type: female; type 120 Cheongdo-gun, Namseonghyun, 20 v 2002 (JW Lee). 56 depository: NM. 121 Distribution. South Korea, Japan, Russia. 57 122 58 Material examined. South Korea: 1\, GN, Haman-gun, Jeisan, 22 v 123 Genus Seticornuta Morley, 1913 59 1996 (JY Cha); 1_, Wondang-ri, 4 vi 1996 (JI Kim). 124 Megatrema Cameron, 1907: 468. Type species: Megatrema albopi- 60 Distribution. South Korea, Austria, Belarus, Bulgaria, Croatia, 125 losa Cameron, by monotypy. Junior homonym of Megatrema Leach. 61 Czech Republic, Finland, France, Germany, Hungary, Ireland, the 126 Seticornuta Morley, 1913: 310. Type species: Seticornuta albicalcar 62 Netherlands, Poland, Romania, Russia, Slovakia, Spain, Turkey, 127 Morley (¼ albopilosa Cameron), by original designation. 63 Turkmenistan, Ukraine, Yugoslavia. 128 64 129 65 130

JK Choi, JW Lee / Journal of Asia-Paciﬁc Biodiversity xxx (2016) 1e19 11

1 80. Seticornuta koreana Lee & Choi, 2015 (species valid) 검은날 82. Stethoncus sulcator Aubert, 1963 고랑뭉툭맵시벌 66 2 개뭉툭맵시벌 Stethoncus sulcator Aubert, 1963:82e86. Type: male; type de- 67 3 Seticornuta koreana Lee & Choi, 2015: 141. Type: female; type depository: MZ. 68 4 pository: YNU. 69 5 Material examined. South Korea: 1_, GN, Sancheong-gun 70 6 Material examined. South Korea: 1\, CN, Daejeon-si, Dong-gu, Sicheon-myeon Naedae-ri Jirisan National Park Saeseoksanjang, 71 7 Daejeon University, 16 ve5 vi 2006 (JW Lee); 1_, Seoul, Achasan, 24 12 x 2001 (JW Lee). 72 8 viii 1980 (KS Jang); 1\, GG, Yongmunsa, 1 ix 1980 (KS Jang); 1_, GG, Distribution. South Korea, Austria, Bulgaria, France, Germany, 73 9 Sudong, Chukryeongsan, 28 ix 1980 (JI Kim); 1\, CN, Buyeo-gun, Japan, Poland, Russia, Switzerland, Ukraine. 74 10 Gyuam-myeon, Sumok-ri, 1-15 vi 2005 (JW Lee); 1\, CN, Daejeon- 75 11 si, Dong-gu, Daejeon University, 16 v-5 vi 2006 (JW Lee); 1_, GB, Genus Triclistus Förster, 1869 76 12 Cheongdo-gun Unmun-myeon, Sinwon-ri, Unmunsan, Unmunsa, Triclistus Förster, 1869:135e221. Type species: Exochus podagricus 77 13 17 vii 1989 (JW Lee); 1_, GN, Jinju-si, Gajoa-dong, 19-23 vi 1989 Förster. 78 14 (JG Kim). 79 83. Triclistus aitkeni (Cameron, 1897) 이마판뭉툭맵시벌 15 Distribution. South Korea. 80 Exochus aitkini Cameron, 1897:1e144. Type: male; type depository: 16 Remarks. This species is distinguished from S. albopilosa by the 81 HOPE. 17 following characters: antenna entirely black (reddish brown in 82 Exochus aitkinii Dalla Torre, 1901:1e544. Type: unknown; type 18 basal half in S. albopilosa) and propodeum with areola separated 83 depository: unknown. 19 from area basalis (areola merged with area basalis in S. albopilosa). 84 Exochus curvicarinatus Cameron, 1902:419e449. Type: female; 20 Watanabe (2015) synonymized S. koreana with Seticornuta nigra. 85 type depository: NHM. 21 However, there has been no comparison of the holotype of 86 22 S. koreana with the holotype of S. nigra. Therefore, we have to 87 Material examined. South Korea: 1_, Daegu-si, Dalseo-gu, 23 compare type specimens of S. koreana and S. nigra. We provide 88 Daegok-dong, Daegu Arboretum, Alt. 88 m, 4-11 vii 2012 (SG Kang). 24 differences of S. koreana and S. nigra based on both original de- 89 Distribution. South Korea, China, India, Japan, the Philippines, 25 scriptions (Table 1). 90 Russia, Réunion. 26 91 27 Genus Spudaeus Gistel, 1848 92 84. Triclistus areolatus Thomson, 1887 작은작은무늬날개뭉툭맵시벌 28 Spudaeus Gistel, 1848:1e216. 93 (Figure 1) 29 Tylocomnus Holmgren, 1873:55e78. Type species: Trachyderma 94 Triclistus areolatus Thomson, 1887:193e218. Lectotype: female; 30 scabra Holmgren. 95 type depository: ZI. 31 96 81. Spudaeus orientalis (Meyer, 1930) 동양수중다리뭉툭맵시벌 32 97 Tylocomnus orientalis Meyer, 1930:165e180. Type: unknown; type Diagnosis. Color. Black. Palpi, ventral part of antenna, tegula, 33 98 depository: ZIN. and legs yellowish brown; hind coxa darkened basally 34 99 (Figure 1A). 35 100 Material examined. South Korea: 1\, Seoul, Hoagwi-dong, Face closely punctate (Figure 1B). Malar space shorter than 36 101 Kyeonghee Univ., 20 v 1994 (YN Lee); 1_, GW, Wongju-si, Gwirae- base of mandible. Clypeus not separated from face. Frons shiny, 37 102 myeon, from Cheoneunsa Temple, Sibjabong, 17 v 2003 (YH Kim, JH with longitudinal ridge. Occipital carina present. Antenna with 38 103 Lee & NR Lee); 1_, GW, Wongju-si, Heungeup-myeon, Maeji-ri, 29 ﬂagellomeres. Pronotum sunken and glabrous (Figure 1C). 39 104 Yonsei Univ., 8 v 2003 (JY Jo & SG Yang); 1\, GG, Soyosan, 3 vi Mesoscutum sparsely punctate, without notaulus (Figure 1D). 40 105 1997 (JS Ann); 1\, GG, Cheonmasan, 22 vi 1969; 1_, GG, Kwan- Scutellum without lateral carina. Mesopleuron sparsely punc- 41 106 gryeong, 14 v 1972 (JI Kim). tate and metapleuron glabrous. Propodeum without areola; 42 107 Distribution. South Korea, Russia. basal and areola areas not separated; costula incomplete 43 108 (Figure 1F). Median longitudinal carinae of propodeum reach 44 109 Genus Stethoncus Townes & Townes, 1959 over middle of propodeum. Propodeal spiracle reaches pleural 45 110 Stethoncus Townes & Townes, 1959:1e318. Type species: Stetho- carina. Forewing with areolet (Figure 1E). Hind wing with six 46 111 ncus arcticus Townes & Townes. distal hamuli; discoidella vestigial. Tergites sparsely punctate 47 112 and shiny. 48 113 Material examined. South Korea: 1\, GN, Sancheon-gun Gwan- 49 114 sulryeong, 6 ix 1998 (SM Ryu); 1\, GG, Yangpyeong-gun Seojong- 50 Table 1. Differences between Seticornuta koreana Lee & Choi, 2015 and Seticornuta 115 Q4 nigra Sheng, 2013. myeon Junggoksa, 23 v 1981 (JG Kim); 1\, GW, Hoengseong-gun 51 116 Anheung-myeon Gangrim-ri, 8 vi 1985 (KS Jeong). 52 Characters S. koreana S. nigra 117 Distribution. South Korea (new record), Austria, Belgium, Czech 53 118 Body length (mm) 11 10 Republic, Denmark, France, Germany, Lithuania, Poland, Russia, 54 Forewing length (mm) 9 7.5 119 Slovakia, Sweden, Turkey, Ukraine, United Kingdom. 55 Antenna length (mm) 9 6.5 120 Host. Earias clorana (Sedivy, 2001). 56 Antenna length/body 0.82 0.65 121 length 57 122 Malar space length 0.3 times as long as width 0.6 times as long as 85. Triclistus glabrosus Momoi & Kusigemati, 1970 등매끈뭉툭 58 of base of mandible width of base of 맵시벌 123 59 mandible 124 Triclistus glabrosus Momoi & Kusigemati, 1970:401e415. Type: fe- 60 Propodeal carina Basal area separated with Basal area combined with 125 central area by a low cross central area male; type depository: HU. 61 ridge 126 62 127 Propodeum size Propodeum 0.55 times as Propodeum 0.7 times as Material examined. No Korean specimens were available for this 63 long as wide long as wide 128 study. 64 Ovipositor Not reaching the end Reaching to end of 129 Distribution. South Korea, Japan. 65 of metasoma metasoma 130

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1 66 2 67 3 68 4 69 5 70 6 71 7 72 8 73 9 74 10 75 11 76 12 77 13 78 14 79 15 80 16 81 17 82 18 83 19 84 20 85 21 86 22 87 23 88 24 89 25 90 26 91 27 92 28 93 29 94 30 95 31 96 32 97 33 98 34 99 35 100 36 101 37 102 38 103 39 104 40 105 41 Figure 1. Triclistus areolatus (Thomson): A, habitus in lateral view; B, head in frontal view; C, head and mesosoma in lateral view; D, head and mesoscutum in dorsal view; E, wings; 106 42 F, propodeum. Scale bars: 1 mm (A, E); 0.5 mm (C); 0.2 mm (B, D, F). 107 43 108 44 86. Triclistus globulipes (Desvignes, 1856) 갈색다리뭉툭맵시벌 Distribution. South Korea, Japan, Mongolia, Russia. 109 45 Exochus globulipes Desvignes, 1856:1e120. Lectotype: male; type 110 46 depository: NHM. 88. Triclistus kamijoi Momoi & Kusigemati, 1970 무릉검은뭉툭 111 47 Exochus holmgreni Boheman, 1863:57e85. Type: unknown; type 맵시벌 112 48 depository: unknown. (Figure 2) 113 49 Triclistus fuscoapicalis Uchida, 1930: 243e298. Type: male; type Triclistus kamijoi Momoi & Kusigemati, 1970: 404. Type: female; 114 50 depository: HU. type depository: HU. 115 51 116 52 Material examined. South Korea: 1\, GW, Wonju-si, Socho-myeon, Diagnosis. 117 53 Hakgong-ri, Chiaksan National Park, 20 vie19 vii 2013 (JW Lee). Color. Black. Ventral part of antenna, palpi, and tegula yellowish 118 54 Distribution. South Korea, Austria, Azerbaijan, Belarus, Belgium, brown. Legs brown to dark brown (Figure 2A). 119 55 Bulgaria, China, Czech Republic, Denmark, Finland, France, Ger- Face coarsely and roughly punctate (Figure 2B). Frons with me- 120 56 many, Hungary, Ireland, Italy, Japan, Latvia, Lithuania, Moldova, dian longitudinal ridge. Malar space shorter than base of mandible. 121 57 Morocco, the Netherlands, Poland, Romania, Russia, Slovakia, Antenna with 26 flagellomeres. Pronotum sunken and glabrous, with 122 58 Sweden, Switzerland, Turkey, Ukraine, United Kingdom. epomia and wrinkles on ventroposterior area (Figure 2C). Meso- 123 59 scutum sparsely punctate and shiny; almost flat, without notaulus 124 60 87. Triclistus japonicus Kusigemati, 1971 왜뭉툭맵시벌 (Figure 2E). Mesopleuron sparsely punctate; speculum area glabrous. 125 61 Triclistus japonicus Kusigemati, 1971: 205e298. Type: female; type Scutellum without lateral carina. Metapleuron glabrous. Median of 126 62 depository: HU. propodeum glabrous, without areola (Figure 2F). Propodeal spiracle 127 63 reaches pleural carina. Forewing with areolet (Figure 2D). Hind wing 128 64 Material examined. South Korea: 1\, GN, Milyang-si, Danjang- with five distal hamuli. Median longitudinal carinae of first tergite 129 65 myeon, Gucheon-ri, Sajapyeong, 4-10 vi 1990 (JW Lee). narrow. Center of first and second tergites glabrous. 130

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1 66 2 67 3 68 4 69 5 70 6 71 7 72 8 73 9 74 10 75 11 76 12 77 13 78 14 79 15 80 16 81 17 82 18 83 19 84 20 85 21 86 22 87 23 88 24 89 25 90 26 91 27 92 28 93 29 94 30 95 31 96 32 97 33 98 34 99 35 100 36 101 37 102 38 103 39 104 40 105 41 Figure 2. Triclistus kamijoi Momoi & Kusigemati: A, habitus in lateral view; B, head in frontal view; C, head and mesosoma in lateral view; D, wings; E, head and mesoscutum in 106 dorsal view; F, propodeum. Scale bars: 1 mm (A, D); 0.5 mm (C); 0.2 mm (B, E, F). 42 107 43 108 44 109 45 Material examined. South Korea: 1\, GW, Donghae-si Mesopleuron with sparse hairs and impunctate; speculum area 110 46 Samhwa-dong Mureung valley, 21e30 v 2005 (JW Lee); 1\, weakly concave and glabrous (Figure 3C). Metapleuron glabrous. 111 47 Goseong-gun Gangseong-eup Heyangrobong, 13 vi 1992 (JW Propodeum slightly and roughly punctate; propodeal spiracle 112 48 Lee). reaching pleural carina. Propodeum without costula; basal area 113 49 Distribution. South Korea (new record), China, Japan, Russia. narrow. Forewing with areolet (Figure 3A); hind wing with five 114 50 Host. Unknown. distal hamuli; discoidella absent. First tergite impunctate, with 115 51 median longitudinal carinae in dorsal view; with lateral longitu- 116 52 89. Triclistus nigripes Momoi & Kusigemati, 1970 검정배뭉툭맵 dinal carina in lateral view. 117 53 시벌 Material examined. South Korea: 2\\, GW, Goseong-gun 118 54 (Figure 3) Gangseong-eup Heyangrobong, 13 vi 1992 (JW Lee); 2\\, ditto 119 55 Triclistus nigripes Momoi & Kusigemati, 1970: 405. Type: female; (SM Ryu). 120 56 type depository: HU. Distribution. South Korea (new record), Japan, Russia. 121 57 122 58 Diagnosis. 90. Triclistus semistriatus Kusigemati, 1971 지리뭉툭맵시벌 123 59 Color. Black. Palpi, ventral part of antenna, tegula, fore tibia, and Triclistus semistriatus Kusigemati, 1971: 205e298. Type: female; 124 60 fore tarsus yellowish brown. Mid and hind tibia and tarsus brown to type depository: HU. 125 61 dark brown (Figure 3A). 126 62 Face coarsely punctate (Figure 3B). Frons flat, with median Material examined. No Korean specimens were available for this 127 63 longitudinal ridge (Figure 3D). Antenna with 26 flagellomeres. study. This diagnosis is modified from the original description by 128 64 Pronotum sunken and glabrous. Mesoscutum sparsely punctate, Kusigemati (1971). 129 65 without notauli. Scutellum without lateral carinae (Figure 3E). Distribution. South Korea, Japan. 130

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1 66 2 67 3 68 4 69 5 70 6 71 7 72 8 73 9 74 10 75 11 76 12 77 13 78 14 79 15 80 16 81 17 82 18 83 19 84 20 85 21 86 22 87 23 88 24 89 25 90 26 91 27 92 28 93 29 94 30 95 31 96 32 97 33 98 34 99 35 100 36 101 37 102 38 103 39 104 40 105 41 106 Figure 3. Triclistus nigripes Momoi & Kusigemati: A, habitus in lateral view; B, head in frontal view; C, head and mesosoma in lateral view; D, head and mesoscutum in dorsal view; 42 E, propodeum. Scale bars: 1 mm (A); 0.5 mm (C); 0.2 mm (B, D, E). 107 43 108 44 109 45 91. Triclistus pallipes Holmgren, 1873 검둥뭉툭맵시벌 shiny. Epomia short (Figure 4D). Mesoscutum with sparse and weak 110 46 (Figure 4) punctures, notaulus absent. Scutellum convex, without lateral 111 47 Triclistus pallipes Holmgren, 1873: 59. Type: female; type de- carinae. Mesopleuron swollen, with sparse punctures in front half. 112 48 pository: NNM. Speculum area smooth and weakly concave. Propodeum shiny and 113 49 Triclistus nitifrons Thomson, 1887:193e218. Type: female; type impunctate, with median longitudinal and apical transverse 114 50 depository: ZI. carinae, with lateral longitudinal carinae (Figure 4E). Costula ab- 115 51 Triclistus pallidipes Dalla Torre, 1901:1e1141. sent. Propodeal spiracle round, joining pleural carina. Pleural carina 116 52 curved. Forewing with petiolated areolet (Figure 4F); nervulus 117 53 Diagnosis. distad to basal vein. Hind wing with six distal hamuli. Nervellus 118 54 Color. Black. Palpi, tegula and ventral parts of antenna yellow. inclivous, intercepted below the middle; discoidella absent. Legs 119 55 Dorsal part of antenna black. Mandible reddish yellow except for very stout. Hind femur 2.3 times as long as wide. Ratio between 120 56 basal part black. All legs yellowish brown (Figure 4A). lengths of hind tarsomeres 18:9:7:4:9. Spurs of mid tibia equal 121 57 Face convex, 1.1 times as long as wide in front view; face and length. Tarsal claws simple and curved. Petiole short, with lateral 122 58 clypeus densely punctate. Clypeus not separated from face carina. Median dorsal carinae extending to half of first tergite. 123 59 (Figure 4B). Mandible except teeth with moderate punctures. Lower Second tergite 0.7 times as long as longest wide. 124 60 tooth of mandible shorter than upper tooth. Malar space 0.5 times Material examined. South Korea: 2\\, Seoul, Nowon-gu Sang- 125 61 as long as basal width of mandible. Frons with median longitudinal gye-dong Suraksan, 18 viie24 viii 2007 (JO Lim); 1\, Daejeon-si 126 62 ridge. Diameter of lateral ocellus 0.8 times as long as shortest dis- Dong-gu Yongun-dong Daejeon Univ., 1e27 xi 2006 (JW Lee); 127 63 tance between ocellus and eye (Figure 4C). Occipital carina strong. 1\, CB, Cheongwon-gun Cheongjusan, 18 vi 1992 (JW Lee); 1\, CN, 128 64 Antenna with 27 flagellomeres. First flagellomere about 4 times as Seosan-si Haemi-myeon Daegok-ri Hanseo Univ., 7 v 2013 (JG 129 65 long as wide, next flagellomeres square. Pronotum impunctate and Kim); 1\, ditto, 21 vie27 vii 2011 (JK Kim); 1\, GN, Gajisan, 28 130

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1 66 2 67 3 68 4 69 5 70 6 71 7 72 8 73 9 74 10 75 11 76 12 77 13 78 14 79 15 80 16 81 17 82 18 83 19 84 20 85 21 86 22 87 23 88 24 89 25 90 26 91 27 92 28 93 29 94 30 95 31 96 32 97 33 98 34 99 35 100 36 101 37 102 38 103 39 104 40 105 41 106 42 107 43 108 44 109 45 110 46 Figure 4. Triclistus pallipes Holmgren: A, habitus in lateral view; B, head in frontal view; C, head in dorsal view; D, head and mesosoma in lateral view; E, propodeum; F, wings. Scale 111 bars: 1 mm (A); 0.5 mm (CeF); 0.2 mm (B). 47 112 48 113 49 viiie2 ix 1989 (JG Kim); 1\, GB, Gyeongsan-si Daehak-ro 280 92. Triclistus podagricus (Gravenhorst, 1829) 계룡검은뭉툭맵시 114 50 Yeungnam Univ., 8e15 vii 2013 (JW Lee); 1\, Yeongju-si Punggi- 벌 115 51 eup Sobaeksan Haebang valley, 13 v 1997 (JW Lee); 1\,GW, (Figure 5) 116 52 Hongcheon-gun Bukbang-myeon Gangwon prov. Environment Exochus podagricus Gravenhorst, 1829: 336. Neotype: female; type 117 53 Research Park, 31 vie31 viii 2012 (SJ Jang); 1\, Pyeongchang-gun depository: ETH. 118 54 Yongpyeong-myeon Gyebangsan, 14 vii 1995 (HC Park); 1\, ditto, Triclistus nigritellus Holmgren: 55e78. 119 55 28 vie12 viii 2012 (JY Park); 1\, Inje-gun Girin-myeon Jindong-ri 120 56 Jeombongsan, 26 vie28 vii 2012 (JY Park); 1\, Jeongseon-gun Diagnosis. 121 57 Sabuk-eup Duwibong, 10 iv 2000 (JW Lee); 1\, Jeongseon-gun Color. Black. Ventral part of antenna, tegula, inner part of fore and 122 58 Hambaeksan, 11 vii 1999 (JW Lee); 1\, Goseong-gun Ganseong- mid tibia, and tarsi and basal of all tibiae yellowish brown (Figure 5A). 123 59 eup Heyangrobong, 13 v 1992 (SM Ryu); 1\, JB, Jeongeup-si Nae- Face closely punctate (Figure 5B). Frons sparsely punctate, with 124 60 jang-dong Naejangsan Wonjeog valley, 19 iv 2008 (JW Lee); 1\, strong median longitudinal ridge. Antenna with 33 ﬂagellomeres. 125 61 Jeongeup-si Naejang-dong Naejangsan Wonjeogam, 25 vie5 vii Pronotum sunken and glabrous (Figure 5C). Mesoscutum with 126 62 2007 (JW Lee); 1\, Wangju-gun Unju-myeon Sanbung-ri Dae- short notauli (Figure 5E). Scutellum without lateral carinae. Mes- 127 63 dunsan, 28 ix 2000 (JW Lee). opleuron sparsely punctate. Lower part of metapleuron glabrous; 128 64 Distribution. South Korea (new record), worldwide. upper part of metapleuron sparsely punctate. Propodeal carinae 129 65 Host. Unknown. fused and glabrous in the center; without costula (Figure 5F). 130

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1 66 2 67 3 68 4 69 5 70 6 71 7 72 8 73 9 74 10 75 11 76 12 77 13 78 14 79 15 80 16 81 17 82 18 83 19 84 20 85 21 86 22 87 23 88 24 89 25 90 26 91 27 92 28 93 29 94 30 95 31 96 32 97 33 98 34 99 35 100 36 101 37 102 38 Figure 5. Triclistus podagricus (Gravenhorst): A, habitus in lateral view; B, head in frontal view; C, head and mesosoma in lateral view; D, wings; E, head and mesoscutum in dorsal 103 view; F, propodeum. Scale bars: 1 mm (A, D); 0.5 mm (C, E, F); 0.2 mm (B). 39 104 40 105 41 Propodeal spiracle reaching pleural carina. Forewing with petio- Color. Black. Ventral part of antenna, palpi, tegula and legs yel- 106 42 lated areolet (Figure 5D). Hind wing with eight distal hamuli; dis- low. Mandible reddish brown. 107 43 coidella vestigial. All femora stout and swollen. First tergite with Face granulated (Figure 6B); frons sparsely punctate, with 108 44 short median longitudinal carinae and shiny. median longitudinal ridge. Antenna with 23 flagellomeres. Mes- 109 45 Material examined. South Korea: 1\, CN, Gyeryong-si Sindoan- osoma depressed (Figure 6A). Pronotum sunken and glabrous, 110 46 myeon Bunam-ri Gyeryongsan National Park (M.T.), 14e28 vii 2012 with wrinkles on lower corner (Figure 6C). Mesoscutum flat, shiny 111 47 (JC Jeong); 1\, GN, Sancheong-gun Samjang-myeon Daepo-ri and sparsely punctate; without notaulus (Figure 6E). Scutellum 112 0 00 0 00 48 Jangdanggol Bakkatjangdang, 35 32 07.59 N12779 83.19 E, 12 sparsely punctate and shiny, without lateral carina. Mesopleuron 113 49 vie11 viii 2001 (JW Lee). sparsely punctate; speculum area glabrous and flat. Metapleuron 114 50 Distribution. South Korea (new record), Austria, Belarus, glabrous. Propodeum without costula and areola (Figure 6F). 115 51 Belgium, Bulgaria, Canada, Czech Republic, Denmark, Finland, Forewing with areolet (Figure 6D). Hind wing with six distal 116 52 France, Georgia, Germany, Hungary, Ireland, Italy, Japan, hamulil; discoidella absent. Legs stout. Metasoma shiny and 117 53 Kazakhstan, Latvia, Lithuania, Luxembourg, the Netherlands, Nor- impunctate. 118 54 way, Poland, Romania, Russia, Slovakia, Spain, Sweden, Material examined. South Korea: 1\, CB, Danyang-gun 119 55 Switzerland, Turkey, USA, Ukraine, United Kingdom. Danyang-eup Chendong-ri Sobaeksan National Park Birobong, 120 56 Host. Phycita roborella (Sedivy, 2001). 22 v 1997 (JW Lee); 1\, Yeongdong-gun Sangchon-myeon Mul- 121 57 han valley, 23 v 2002 (JW Lee); 3\\, JN, Gurye-gun Sandong- 122 58 93. Triclistus pygmaeus (Cresson, 1864) 노고검고검은뭉툭맵시벌 myeon Nogodan Jirisan National Park, (light trap), 22 viii 1997 123 59 (Figure 6) (JW Lee). 124 60 Exochus pygmæus Cresson, 1864:257e321. Lectotype: female; type Distribution. South Korea (new record), Austria, Belgium, 125 61 depository: ANSP. Canada, China, Czech Republic, Denmark, Finland, France, 126 62 Triclistus taiwanensis Uchida, 1932:133e222. Type: female; type Georgia, Germany, Italy, Lithuania, Moldova, Norway, Poland, 127 63 depository: DEI. Russia, Sweden, Switzerland, Taiwan, USA, Ukraine, United 128 64 Kingdom. 129 65 Diagnosis. Host. Unknown. 130

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18 JK Choi, JW Lee / Journal of Asia-Paciﬁc Biodiversity xxx (2016) 1e19

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