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Home , Agonus cataphractus, Anarhichas minor, Anarhichas orientalis, Arctic alligatorfish, Arctic flounder, Arctic skate

List of Marine of the Region Annotated with Common Names and Zoogeographic Characterizations

November 2013 . Photo: Shawn Harper, University of Alaska Fairbanks of Alaska University Harper, Shawn . Photo: Boreogadus saida Boreogadus Polar , cod, Polar Acknowledgements

CAFF Designated Agencies:

• Directorate for Nature Management, Trondheim, • Environment Canada, Ottawa, Canada • Faroese Museum of Natural History, Tórshavn, Faroe Islands (Kingdom of Denmark) • Finnish Ministry of the Environment, Helsinki, Finland • Icelandic Institute of Natural History, Reykjavik, Iceland • The Ministry of Domestic Housing, Nature and Environment, Government of Greenland • Russian Federation Ministry of Natural Resources, Moscow, • Swedish Environmental Protection Agency, Stockholm, Sweden • United States Department of the Interior, and Wildlife Service, Anchorage, Alaska

CAFF Permanent Participant Organisations:

• Aleut International Association (AIA) • Arctic Athabaskan Council (AAC) • Gwich’in Council International (GCI) • Inuit Circumpolar Council (ICC) • Russian Indigenous Peoples of the North (RAIPON) • Saami Council

This publication should be cited as: Mecklenburg, C.W., I. Byrkjedal, J.S. Christiansen, O.V. Karamushko, A. Lynghammar and P. R. Møller. 2013. List of marine fishes of the arctic region annotated with common names and zoogeographic characterizations. Conservation of Arctic Flora and Fauna, Akureyri, Iceland. Front cover photo: Polar cod, Boreogadus saida: Shawn Harper, University of Alaska Fairbanks

Layout, photographs and editing: the authors

For more information please contact: CAFF International Secretariat Borgir, Nordurslod 600 Akureyri, Iceland Phone: +354 462-3350 Fax: +354 462-3390 Email: [email protected] Internet: www.caff.is

___ CAFF Designated Area Arctic Marine Fish List, 11 November 2013

List of Marine Fishes of the Arctic Region Annotated with Common Names and Zoogeographic Characterizations By: Catherine W. Mecklenburg, California Academy of Sciences, San Francisco, California, USA; and Point Stephens Research, Auke Bay, Alaska, USA; [email protected] Ingvar Byrkjedal, University Museum of Bergen, University of Bergen, Bergen, Norway Jørgen S. Christiansen, Department of Arctic and , UiT–The Arctic University of Norway, Tromsø, Norway Oleg V. Karamushko, Murmansk Marine Biological Institute, Kola Science Centre, Russian Academy of Sciences, Murmansk, Russia Arve Lynghammar, Department of Arctic and Marine Biology, UiT–The Arctic University of Norway, Tromsø, Norway Peter R. Møller, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark

Gymnocanthus tricuspis

Enophrys lucasi

Limanda proboscidea

November 11, 2013

Hypsagonus quadricornis

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Introduction A recent review of historical and new records of Assessment (Christiansen et al. 2013), resulting in presence produced an annotated list of the a smaller number of listed species. However, it marine and diadromous fishes occurring in the encompasses all the sentinel and desirable fish arctic region (Mecklenburg et al. 2010). Consider- monitoring stations identified in the CBMP’s able new taxonomic and distributional information Arctic Marine Biodiversity Monitoring Plan (Gill has already accumulated since that review went to et al. 2011:57). The listed marine species are, press (June 2010). In addition to bringing the therefore, the subjects of the CBMP’s Marine Fish scientific names list up to date with new or re- Expert Network (MFEN). moved species and various nomenclatural changes, Common names, or vernaculars, for all the in this edition we have provided the most common marine fishes in the region were previously not or officially recommended English, French, Nor- available in any one published list. In the present wegian, and Russian vernaculars. Zoogeographic list (Table 1), with few exceptions the English characterizations (e.g., arctic, arctic-boreal, boreal) names for North American fishes occurring at con- from the earlier review (Mecklenburg et al. 2010) tinental shelf depths are from the official list of the are also included, with a few additions following American Fisheries Society and American Society the same definitions. of Ichthyologists and Herpetologists (Page et al. The listed species have all been found, at least 2013). Although that work capitalizes the common once, in the area defined as the arctic region in the names, we follow prevailing usage in other coun- recent review (Figure 1). The arctic region was tries and only capitalize proper noun references defined faunistically. Its southern boundaries such as in Adolf’s , Norway pout, and reflect the southern limit of most arctic-adapted Atlantic . Individually authored family fish species’ distributions. In the Pacific , chapters in the classic treatise on northeastern for instance, they do not generally occur south of a Atlantic fishes (Whitehead et al. 1984–1986) line extending from the Russian coast just north of provided most of the English names for deepwater Cape Navarin and curving southward toward St. species; sources also included the recent Green- Matthew Island, and then northward to Alaska’s land checklist (Møller et al. 2010) and original Yukon Delta. The boundary generally reflects the descriptions of the species. The French names are position of the Polar Front, north of which en- from lists and guides to Canadian Arctic and vironmental conditions are optimal for arctic western Atlantic fishes (McAllister 1990, Coad species. In the Atlantic side of the arctic region the and Reist 2004, Nozéres et al. 2010), as well as the boundary is more complex and defined as well by AFS–ASIH list. The Norwegian names are mostly seafloor topography such as sills and ridges or from Pethon (2005) and the species nomenclature other conditions believed to restrict movement of database of the Norwegian Biodiversity Informa- species. Other arctic region definitions are in use tion Centre (Artsnavnebase 2013). A few new which cover a larger area, but they also encompass Norwegian and Russian names were invented for warmer waters and, consequently, large numbers the atlas of fishes (Wienerroither et al. of subarctic or boreal species that have not entered 2011). Russian names are mainly from the Fishes truly arctic waters. If those species are discovered of the Northern Seas of the USSR (Andriashev in the arctic region, they will become subjects of 1954), the five-language dictionary of great interest as possible reflections of climate names (Reshetnikov et al. 1989), and the catalog change. This faunistic definition circumscribes a of Kamchatka fishes (Sheiko and Fedorov 2000). somewhat smaller total area than, for instance, the The Russian names are given in cyrillic and trans- definition used in the recent Arctic Biodiversity literated to the Latin alphabet.

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Names in the non-English languages are pro- the arctic region; 14.5% (32) are arctic–boreal, vided if relevant and, preferably, the names are reproducing and common in both regions; 25.8% known and used locally. Some species do not have (57) occur solely or mainly in the arctic region; common names and these mostly are rare species, and 6.3% (14) are widely distributed, occurring in some known only from one record. the warm waters of two or more or the This list includes 221 marine species. The southern hemisphere and only rarely in the arctic number will change as more species enter the region. arctic region and as further study shows some Although improved and expanded since the species to be the same as other species (synony- earlier list (Mecklenburg et al. 2010), the present mous with them, in the taxonomic jargon) and yet list (Table 1) remains essentially a draft. The others to include more than one species (so-called continuing evolution of the list would perhaps species complexes). Of the current total, 92.8% pertain for any large-region faunal inventory but (205) are ray-finned fishes (actinopterygians) such for fishes is especially a consequence for the arctic as , , , and . The region, where the fish fauna is relatively poorly remainder, 7.2% (16), are cartilaginous fishes known and has only in recent years become a (chondrichthyans), namely sharks and skates, and focus of extensive sampling and analysis. The a hagfish (a jawless fish, or agnathan). Overall, present collaborators and the MFEN intend to 53.4% (118) have boreal or mainly boreal distri- revise the list from time to time as new discoveries bution patterns, with relatively small presence in and improved information accumulate.

Figure 1. Arctic marine ichthyofaunal region (Mecklenburg et al. 2010).

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Table 1. Marine fishes of the arctic region

Family numbers, placing the families in phylogenetic sequence, are from the classification of fishes of the world by Nelson (2006). Species are listed alphabetically by family or subfamily.

Order Myxiniformes (hagfishes) 001 MYXINIDAE (hagfishes) MYXININAE Myxine glutinosa Linnaeus, 1758 — Boreal Atlantic English name: Atlantic hagfish French name: myxine du nord Norwegian name: slimål Russian name: атлантическая миксина Russian name: (atlanticheskaya miksina)

Order Lamniformes ( sharks) 021 CETORHINIDAE (basking sharks) Cetorhinus maximus (Gunnerus, 1765) — Widely distributed English name: basking shark French name: pèlerin Norwegian name: brugde Russian name: гигантская акула Russian name: (gigantskaya akula)

022 LAMNIDAE (mackerel sharks) Lamna ditropis Hubbs & Follett, 1947 — Boreal Pacific English name: salmon shark Russian name: тихоокеанская сельдевая акула Russian name: (tikhookeanskaya seldevaya akula) Lamna nasus (Bonnaterre, 1788) — Boreal Atlantic English name: French name: maraîche – Nozéres et al. 2010 Norwegian name: håbrann Russian name: атлантическая сельдевая акула Lamna ditropis Russian name: (atlanticheskaya seldevaya akula)

Order Squaliformes (dogfish sharks) 034 SQUALIDAE (dogfish sharks) Squalus acanthias Linnaeus, 1758 — Widely distributed English name: spiny dogfish, piked dogfish French name: aiguillat commun Norwegian name: pigghå Russian name: катран

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Russian name: (katran) Squalus suckleyi (Girard, 1855) — Boreal Pacific English name: spotted spiny dogfish Russian name: короткопёрая колючая акула Russian name: (korotkopyoraya kolyuchaya akula)

037 SOMNIOSIDAE (sleeper sharks) Centroscymnus coelolepis Barbosa du Bocage & de Brito Capello, 1864 — Boreal Atlantic English name: Portuguese dogfish French name: pailona commun Norwegian name: dypvannshå Russian name: белоглазая акула Russian name: (beloglazaya akula) Somniosus microcephalus (Bloch & Schneider, 1801) — Arctic–boreal Atlantic English name: French name: laimargue Norwegian name: håkjerring Russian name: атлантическая полярная акула Russian name: (atlanticheskaya polyarnaya akula) Somniosus pacificus Bigelow & Schroeder, 1944 — Boreal Pacific English name: Pacific sleeper shark Russian name: тихоокеанская полярная акула Russian name: (tikhookeanskaya polyarnaya akula)

Order (skates) 048 RAJIDAE (skates) ARHYNCHOBATINAE Bathyraja parmifera (Bean, 1881) — Boreal Pacific English name: Alaska Russian name: щитоносный скат Russian name: (shchitonosniy skat) Bathyraja spinicauda (Jensen, 1914) — Mainly boreal Atlantic English name: spinytail skate French name: raie à queue épineuse Norwegian name: gråskate Russian name: шипохвостый скат Russian name: (shipokhovstiy skat) RAJINAE hyperborea (Collett, 1879) — Arctic English name: Arctic skate French name: raie boréale

Norwegian name: isskate Amblyraja hyperborea Russian name: северный скат

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Russian name: (severniy skat) Amblyraja radiata (Donovan, 1808) — Mainly boreal Atlantic English name: thorny skate, starry skate French name: raie épineuse Norwegian name: kloskate Russian name: звёздчатый скат Russian name: (zvezdchatiy skat) bathyphila (Holt & Byrne, 1908) — Boreal Atlantic English name: deepwater ray French name: raie bathyale Russian name: глубоководный скат Russian name: (glubokovodniy skat) Rajella fyllae (Lütken, 1887) — Boreal Atlantic English name: round ray French name: raie ronde Norwegian name: rundskate Russian name: круглый скат Russian name: (krugliy skat) Rajella lintea (Fries, 1838) — Mainly boreal Atlantic English name: sailray French name: raie linon Norwegian name: hvitskate Russian name: парусный скат Russian name: (parusniy skat)

Order Albuliformes () 073 (spiny ) chemnitzii Bloch, 1788 — Widely distributed English name: snubnosed spiny French name: tapir à grandes écailles Norwegian name: nordlig piggål Russian name: спиношип Хемница Russian name: (spinoship Khemnitsa)

080 SYNAPHOBRANCHIDAE (cutthroat eels) Diastobranchus capensis Barnard, 1923 — Widely distributed, primarily southern hemisphere English name: basketwork eel Norwegian name: kappål Russian name: капский диастобранх Russian name: (kapskiy diastobrankh)

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Order () 097 (herrings) CLUPEINAE harengus Linnaeus, 1758 — Mainly boreal Atlantic English name: Atlantic French name: hareng atlantique Norwegian name: sild Russian name: атлантическая сельдь Russian name: (atlanticheskaya seld) Clupea pallasii Valenciennes, 1847 — Arctic–boreal Pacific and eastern Atlantic; amphiboreal English name: Pacific herring French name: hareng du Pacifique Russian name: тихоокеанская сельдь Russian name: (tikhookeanskaya seld)

166 ARGENTINIDAE (argentines) Argentina silus (Ascanius, 1775) — Boreal Atlantic English name: Atlantic argentine, greater argentine French name: grande argentine Norwegian name: vassild Russian name: североатлантическая аргентина Russian name: (severoatlanticheskaya argentina)

Order Osmeriformes (smelts) 168 (pencilsmelts) BATHYLAGINAE Bathylagus euryops Goode & Bean, 1896 — Boreal Atlantic English name: goiter blacksmelt French name: garcette-goître Russian name: тёмный батилаг Russian name: (tyomniy batilag) Melanolagus bericoides (Borodin, 1929) — Widely distributed English name: smalleye blacksmelt French name: garcette à petits yeux Russian name: малоголовый меланолаг Russian name: (malogoloviy melanolag) MICROSTOMATINAE groenlandica (Reinhardt, 1840) — Widely distributed English name: French name: serp de Groenland Norwegian name: grønlandshavkrøkle Russian name: гренландская нансения Russian name: (grenlandskaya nansenia)

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169 PLATYTROCTIDAE (tubeshoulders) Maulisia microlepis Sazonov & Golovan, 1976 — Widely distributed English name: smallscale searsid

171 ALEPOCEPHALIDAE (slickheads) Alepocephalus agassizii Goode & Bean, 1883 — Boreal Atlantic English name: dusky slickhead French name: alépocéphale obscur Russian name: гладкоголов Агассиза Russian name:(gladkogolov Agassiza)

172 OSMERIDAE (smelts) OSMERINAE Mallotus villosus (Müller, 1776) — Arctic–boreal; circumpolar English name: French name: capelan Norwegian name: lodde Russian name: мойва Mallotus villosus Russian name: (moyva)

Order (dragonfishes) 179 (bristlemouths) microdon (Günther, 1878) — Widely distributed English name: French name: Cyclothone à petites dents Norwegian name: svart lysfisk Russian name: малозубая циклотона Russian name: (malozubaya tsiklotona)

180 STERNOPTYCHIDAE (marine hatchetfishes) STERNOPTYCHINAE Argyropelecus hemigymnus Cocco, 1829 — Widely distributed, circumglobal English name: short silver hatchetfish, spurred hatchetfish French name: hache d’argent courte Norwegian name: flekket perlemorsfisk Russian name: малый топорик Russian name: (maliy toporik) MAUROLICINAE Maurolicus muelleri (Gmelin, 1789) — Boreal eastern Atlantic English name: pearlsides Norwegian name: laksesild Russian name: мавролик Мюллера

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Russian name: (mavrolik Myullera)

182 (barbeled dragonfishes) STOMIINAE Stomias boa (Risso, 1810) — Widely distributed except not in North Pacific English name: boa dragonfish French name: dragon-boa Norwegian name: storkjeft Russian name: стомия-боа Russian name: (stomiya-boa)

Order Aulopiformes (lizardfishes) 196 PARALEPIDIDAE (barracudinas) Arctozenus risso (Bonaparte, 1840) — Widely distributed English name: white barracudina French name: lussion blanc Norwegian name: liten laksetobis Russian name: северный веретенник Russian name: (severniy veretennik) Paralepis coregonoides Risso, 1820 — Widely distributed English name: sharpchin barracudina French name: lussion à menton Russian name: сиговидный веретенник Russian name: (sigovidniy veretennik)

Order () 200 MYCTOPHIDAE (lanternfishes) LAMPANYCTINAE Lampanyctus macdonaldi (Goode and Bean, 1896) — Widely distributed English name: rakery , rakery beaconlamp French name: lanterne-bouée râtelière Norwegian name: brun lysprikkfisk Russian name: лампаникт Макдональда Russian name: (lampanikt Macdonalda) Myctophum punctatum Rafinesque, 1810 — Boreal Atlantic English name: spotted lanternfish French name: lanterne ponctuée Norwegian name: liten lysprikkfisk Russian name: пятнистый миктоф Russian name: (pyatnistiy miktof) MYCTOPHINAE Benthosema glaciale (Reinhardt, 1837) — Arctic–boreal Atlantic English name: glacier lanternfish

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French name: lanterne glaciaire Norwegian name: nordlig lysprikkfisk Russian name: северная бентозема Russian name: (severnaya bentozema) arcticum (Lütken, 1892) — Arctic–boreal Atlantic English name: Arctic telescope French name: télescope arctique Norwegian name: laksestørje Russian name: арктический протомиктоф Russian name: (arkticheskiy protomiktof)

Order Lampriformes (opahs) 202 LAMPRIDAE (opahs) Lampris guttatus (Brünnich, 1788) — Widely distributed English name: opah Norwegian name: laksestørje Russian name: краснопёрый опах Russian name: (krasnopyoriy opakh)

Order (cods) 215 MACROURIDAE (grenadiers) MACROURINAE Coryphaenoides rupestris Gunnerus, 1765 — Boreal Atlantic English name: rock grenadier French name: grenadier de roche Norwegian name: skolest Russian name: тупорылый макрурус Russian name: (tuporyliy makrurus) Macrourus berglax Lacepède, 1801 — Mainly boreal Atlantic English name: roughhead grenadier French name: grenadier berglax Norwegian name: isgalt Russian name: северный макрурус Russian name: (severniy makrurus)

219 PHYCIDAE (phycid hakes) GAIDROPSARINAE Ciliata septentrionalis (Collett, 1875) — Boreal eastern Atlantic English name: northern rockling Norwegian name: nordlig tangbrosme Russian name: северный пятнистый налим Russian name: (severniy pyatiusiy nalim)

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Gaidropsarus argentatus (Reinhardt, 1837) — Arctic–boreal Atlantic English name: silver rockling, Arctic rockling French name: mustèle argentée Norwegian name: sølvtangbrosme Russian name: полярный налим Russian name: (polyarniy nalim) Gaidropsarus ensis (Reinhardt, 1837) — Arctic–boreal Atlantic English name: threebeard rockling French name: mustèle arctique à trois barbillons Russian name: северный трёхусый налим Russian name: (severniy tryokhusiy nalim)

220 (cods) Boreogadus saida GADINAE Arctogadus glacialis (Peters, 1872) — Arctic; circumpolar English name: ice cod, Arctic cod, polar cod French name: saïda imberbe Norwegian name: istorsk Russian name: ледовая (чёрная) треска Russian name: (ledovaya (chyornaya) treska) Boreogadus saida (Lepechin, 1774) — Arctic; circumpolar English name: polar cod, Arctic cod Norwegian name: polartorsk Russian name: сайка Russian name: (sayka) Eleginus gracilis (Tilesius, 1810) — Arctic–boreal Pacific English name: French name: jaune Russian name: тихоокеанская навага Russian name: (tikhookeanskaya navaga) Eleginus nawaga (Walbaum, 1792) — Arctic English name: navaga Norwegian name: nawagatorsk Russian name: северная навага Russian name: (severnaya navaga) Gadus chalcogrammus Pallas, 1814 — Mainly boreal Pacific and eastern Atlantic; amphiboreal English name: walleye pollock, Atlantic pollock Norwegian name: berlevågfisk Russian name: тихоокеанский минтай Russian name: (tikhookeanskiy mintay)

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Gadus macrocephalus Tilesius, 1810 — Arctic–boreal Pacific and Atlantic English name: , ogac French name: morue du Pacifique, ogac Russian name: тихоокеанская треска Russian name: (tikhookeanskaya treska) Gadus morhua Linnaeus, 1758 — Mainly boreal Atlantic English name: French name: morue franche Norwegian name: torsk Russian name: атлантическая треска Russian name: (atlanticheskaya treska) Melanogrammus aeglefinus (Linnaeus, 1758) — Mainly boreal Atlantic English name: Norwegian name: hyse Russian name: пикша Russian name: (piksha) Merlangius merlangus (Linnaeus, 1758) — Boreal eastern Atlantic English name: whiting Norwegian name: hvitting Russian name: мерланг Russian name: (merlang) Micromesistius poutassou (Risso, 1827) — Mainly boreal Atlantic English name: blue whiting Norwegian name: kolmule Russian name: путассу Russian name: (putassu) Pollachius virens (Linnaeus, 1758) — Boreal eastern Atlantic English name: pollock, saithe Norwegian name: sei Russian name: сайда Russian name: (sayda) esmarkii (Nilsson, 1855) — Boreal eastern Atlantic English name: Norway pout Norwegian name: øyepål Russian name: тресочка Эсмарка Russian name: (tresochka Esmarka) LOTINAE Brosme brosme (Ascanius, 1772) — Boreal Atlantic English name: cusk Norwegian name: brosme Russian name: менёк Russian name: (menyok)

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Order Lophiiformes (anglerfishes) 227 LOPHIIDAE (goosefishes) Lophius piscatorius Linnaeus, 1758 — Boreal eastern Atlantic English name: angler Norwegian name: breiflabb Russian name: европейский удильщик Russian name: (evropeyskiy udilshchik)

295 SYNGNATHIDAE (pipefishes) SYNGNATHINAE Entelurus aequoreus (Linnaeus, 1758) — Boreal eastern Atlantic English name: snake pipefish Norwegian name: stor havnål Russian name: змеевидная рыба-игла Russian name: (zmeevidnaya ryba-igla)

Order (mail-cheeked fishes) 304 (rockfishes) SEBASTINAE Sebastes alutus (Gilbert, 1890) — Boreal Pacific English name: Pacific ocean perch Russian name: тихоокеанский клювач Russian name: (tikhookeanskiy klyuvach) Sebastes borealis Barsukov, 1970 — Boreal Pacific English name: shortraker rockfish Russian name: северный морской окунь Russian name: (severniy morskoy okun) Sebastes mentella Travin, 1951 — Mainly boreal Atlantic English name: deepwater redfish, beaked redfish French name: sébaste atlantique Norwegian name: snabeluer Russian name: глубоководный клювач Russian name: (glubokovodniy klyuvach) Sebastes norvegicus (Ascanius, 1772) — Mainly boreal Atlantic English name: golden redfish French name: sébaste orangé Norwegian name: vanlig uer Russian name: золотистый морской окунь Russian name: (zolotistiy morskoy okun)

316 HEXAGRAMMIDAE (greenlings) HEXAGRAMMINAE Hexagrammos octogrammus (Pallas, 1814) — Boreal Pacific

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English name: masked greenling Russian name: бурый терпуг Russian name: (buriy terpug) Hexagrammos stelleri Tilesius, 1810 — Mainly boreal Pacific English name: whitespotted greenling Russian name: пятнистый терпуг Russian name: (pyatnistiy terpug) PLEUROGRAMMINAE Pleurogrammus monopterygius (Pallas, 1810) — Boreal Pacific English name: Atka mackerel Russian name: северный однопёрый терпуг Russian name: (severniy odnopyoriy terpug)

320 (sculpins) atlanticus Jordan & Evermann, 1898 — Arctic–boreal English name: Atlantic hookear French name: hameçon atlantique Norwegian name: krokulke Russian name: европейский крючкорог Russian name: (evropeyskiy bychok-kryuchkorog) Artediellus camchaticus Gilbert & Burke, 1912 — Boreal Pacific English name: clownfin sculpin Russian name: тонкохвостый крючкорог Russian name: (tonkokhvostiy kryuchkorog) Artediellus gomojunovi Taranetz, 1933 — Boreal Pacific English name: spinyhook sculpin Russian name: высоколобый крючкорог Russian name: (vysokolobiy kryuchkorog) Artediellus ochotensis Gilbert & Burke, 1912 — Boreal Pacific English name: Okhotsk hookear sculpin Russian name: усатый крючкорог Russian name: (usatiy kryuchkorog) Artediellus pacificus Gilbert, 1896 — Boreal Pacific English name: hookhorn sculpin, paddled sculpin Russian name: лопастной крючкорог Russian name: (lopastnoy kryuchkorog) Artediellus scaber Knipowitsch, 1907 — Arctic English name: hamecon French name: hameçon rude Norwegian name: sibirkrokulke Artediellus scaber Russian name: шероховатый крючкорог Russian name: (sherokhovatiy kryuchkorog)

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Artediellus uncinatus (Reinhardt, 1834) — Mainly arctic Atlantic English name: Arctic hookear sculpin, snowflake hookear French name: hameçon neigeux Russian name: гренландский бычок-крючкорог Russian name: (grenlandskiy bychok-kryuchkorog) Enophrys diceraus (Pallas, 1788) — Mainly boreal Pacific English name: antlered sculpin Russian name: северный гребнерог Russian name: (severniy grebnerog) Enophrys lucasi (Jordan & Gilbert, 1898) — Boreal Pacific English name: leister sculpin Russian name: беринговоморский рогатый бычок Russian name: (beringovomorskiy rogatiy bychok) Gymnocanthus galeatus Bean, 1881 — Boreal Pacific English name: armorhead sculpin Russian name: узколобый шлемоносец Russian name: (uzkolobiy shlemonosets) Gymnocanthus pistilliger (Pallas, 1814) — Mainly boreal Pacific English name: threaded sculpin Russian name: нитчатый шлемоносец Russian name: (nitchatiy shlemonosets) Gymnocanthus tricuspis (Reinhardt, 1830) — Arctic; circumpolar English name: Arctic staghorn sculpin French name: tricorne arctique Norwegian name: glattulke Russian name: арктический шлемоносец Russian name: (arkticheskiy shlemonosets) Hemilepidotus jordani Bean, 1881 — Boreal Pacific English name: Russian name: белобрюхий получешуйник Russian name: (belobryukhiy polucheshuynik) Hemilepidotus papilio (Bean, 1880) — Mainly boreal Pacific English name: butterfly sculpin Russian name: Russian name: бычок-бабочка Russian name: (bychok-babochka) Icelus bicornis (Reinhardt, 1840) — Mainly arctic; circumpolar English name: twohorn sculpin French name: icèle à deux cornes Norwegian name: tornulke Russian name: атлантический двурогий ицел Russian name: (atlanticheskiy dvurogiy itsel) Icelus spatula Gilbert & Burke, 1912 — Arctic–boreal English name: spatulate sculpin

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French name: icèle spatulée Norwegian name: spateltornulke Russian name: восточный двурогий ицел Russian name: (vostochniy dvurogiy itsel) Icelus sp. — Arctic (?) Unidentified Icelus species (Mecklenburg et al. 2010) Icelus spiniger Gilbert, 1896 — Boreal Pacific English name: thorny sculpin Russian name: колючий ицел Russian name: (kolyuchiy itsel) Megalocottus platycephalus (Pallas, 1814) — Mainly boreal Pacific English name: belligerent sculpin Russian name: плоскоголовая широколобка Russian name: (ploskogolovaya shirokolobka) Microcottus sellaris (Gilbert, 1896) — Mainly boreal Pacific English name: brightbelly sculpin Russian name: седловидный бычок Russian name: (sedlovidniy bychok) jaok (Cuvier, 1829) — Mainly boreal Pacific English name: plain sculpin Russian name: керчак-яок Russian name: (kerchak-yaok) Myoxocephalus polyacanthocephalus (Pallas, 1814) — Boreal Pacific English name: great sculpin Russian name: многоиглый керчак Russian name: (mnogoigliy kerchak) Myoxocephalus quadricornis (Linnaeus, 1758) — Arctic; circumpolar English name: fourhorn sculpin French name: chaboisseau à dix-huit épines Norwegian name: hornulke Russian name: рогатка Russian name: (rogatka) Myoxocephalus scorpioides (Fabricius, 1780) — Arctic English name: Arctic sculpin French name: chaboisseau arctique Russian name: арктический керчак Russian name: (arkticheskiy kerchak) Myoxocephalus scorpius (Linnaeus, 1758) — Arctic–boreal; circumpolar English name: shorthorn sculpin French name: chaboisseau à épines courtes Norwegian name: vanlig ulke Russian name: европейский керчак

Russian name: (evropeyskiy kerchak) Myoxocephalus scorpius

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Porocottus mednius (Bean, 1898) — Boreal Pacific English name: Aleutian fringed sculpin Russian name: белопятнистый бахромчатый бычок Russian name: (belopyatnistiy bakhromchatiy bychok) Trichocottus brashnikovi Soldatov & Pavlenko, 1915 — Mainly boreal Pacific English name: hairhead sculpin Russian name: волосатоголовый бычок Бражникова Russian name: (volosatogoloviy bychok Brazhnikova) murrayi Günther, 1888 — Mainly boreal Atlantic English name: moustache sculpin French name: faux-trigle armé Norwegian name: nordlig knurrulke Russian name: атлантический триглопс Russian name: (atlanticheskiy triglops) Triglops nybelini Jensen, 1944 — Arctic; circumpolar English name: bigeye sculpin French name: faux-trigle aux grands yeux Norwegian name: grønlandsknurrulke Russian name: полярный триглопс Russian name: (polyarniy triglops) Triglops pingelii Reinhardt, 1837 — Arctic–boreal; circumpolar English name: ribbed sculpin French name: faux-trigle bardé Norwegian name: arktisk knurrulke Russian name: остроносый триглопс Russian name: (ostronosiy triglops)

323 HEMITRIPTERIDAE (sailfin sculpins) bilobus Cuvier, 1829 — Boreal Pacific English name: crested sculpin Russian name: двулопастной бычок Blepsias bilobus Russian name: (dvulopastnoy bychok) Nautichthys pribilovius (Jordan & Gilbert, 1898) — Mainly boreal Pacific English name: eyeshade sculpin Russian name: короткошипый бычок-кораблик Russian name: (korotkoshipiy bychok-korablik)

324 AGONIDAE (poachers) HYPSAGONINAE quadricornis (Valenciennes, 1829) — Boreal Pacific English name: Russian name: северный гипсагон Russian name: (severniy gipsagon)

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ANOPLAGONINAE monopterygius (Bloch, 1786) — Mainly boreal Pacific and western Atlantic; amphiboreal English name: French name: poisson-alligator atlantique Russian name: щитонос Russian name: (shchitonos) Aspidophoroides olrikii Lütken, 1877 — Mainly arctic; nearly circumpolar English name: French name: poisson-alligator arctique Norwegian name: arktisk panserulke Russian name: ульцина Aspidophoroides olrikii Russian name: (ultsina) AGONINAE (Linnaeus, 1758) — Boreal eastern Atlantic English name: hooknose – Andriashev 1986 Norwegian name: panserulke Russian name: европейская лисичка Russian name: (evropeyskaya lisichka) (Bloch & Schneider, 1801) — Arctic–boreal English name: Atlantic poacher French name: agone atlantique Norwegian name: tiskjegg Russian name: длинноусая лисичка Russian name: (dlinnousaya lisichka) accipenserinus (Tilesius, 1813) — Boreal Pacific English name: Russian name: многоусая лисичка Russian name: (mnogousaya lisichka) Podothecus veternus Jordan & Starks, 1895 — Arctic–boreal Pacific English name: Russian name: малоусая лисичка Russian name: (malousaya lisichka) frenatus (Gilbert, 1896) — Boreal Pacific English name: Russian name: тонкохвостая лисичка Russian name: (tonkokhvostaya lisichka) BRACHYOPSINAE dodecaedron (Tilesius, 1813) — Boreal Pacific English name: Russian name: двенадцатигранная лисичка Russian name: (dvenadtsatigrannaya lisichka)

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Pallasina barbata (Steindachner, 1876) — Mainly boreal Pacific English name: tubenose poacher Russian name: игловидная лисичка Russian name: (iglovidnaya lisichka)

325 (fathead sculpins) COTTUNCULINAE microps Collett, 1875 — Arctic–boreal Atlantic English name: polar sculpin French name: cotte polaire Norwegian name: paddeulke Russian name: малоглазый коттункул Russian name: (maloglaziy kottunkul) Eurymen gyrinus Gilbert & Burke, 1912 — Mainly boreal Pacific English name: smoothcheek sculpin Russian name: красногубый эвримен Russian name: (krasnogubiy evrimen) PSYCHROLUTINAE Psychrolutes paradoxus Günther, 1861 — Boreal Pacific English name: tadpole sculpin Russian name: мягкий психролют Russian name: (myagkiy psichrolyut) Psychrolutes subspinosus (Jensen, 1902) — Arctic Russian name: исландский психролют Russian name: (islandskiy psikhrolyut)

327 (lumpsuckers) Aptocyclus ventricosus (Pallas, 1769) — Boreal Pacific English name: smooth lumpsucker Russian name: мягкий круглопёр Russian name: (myagkiy kruglopyor) jordani Soldatov, 1929 — Arctic English name: smooth lumpfish French name: petite poule de mer douce Russian name: круглопёрик Джордана Russian name: (kruglopyorik Dzhordana) Cyclopteropsis mcalpini (Fowler, 1914) — Arctic English name: Arctic lumpsucker Norwegian name: dvergkjeks Russian name: круглопёрик Фаулера Russian name: (kruglopyorik Faulera) Linnaeus, 1758 — Mainly boreal Atlantic English name: lumpfish

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French name: grosse poule de mer Norwegian name: rognkjeks (female), rognkall (male) Russian name: пинагор Russian name: (pinagor) andriashevi Perminov, 1936 — Mainly boreal Pacific English name: pimpled lumpsucker Russian name: круглопёр Андрияшева Russian name: (kruglopyor Andriyasheva) Eumicrotremus derjugini Popov, 1926 — Arctic English name: leatherfin lumpsucker French name: petite poule de mer arctique Norwegian name: svartkjeks Russian name: круглопёр Дерюгина Russian name: (kruglopyor Deryugina) Eumicrotremus orbis Eumicrotremus orbis (Günther, 1861) — Boreal Pacific English name: Russian name: шаровидный круглопёр Russian name: (sharovidniy kruglopyor) Eumicrotremus spinosus (Fabricius, 1776) — Mainly arctic Atlantic English name: Atlantic spiny lumpsucker French name: petite poule de mer atlantique Norwegian name: vortekjeks Russian name: шиповатый круглопёр Russian name: (shipovatiy kruglopyor)

328 LIPARIDAE () Careproctus derjugini Chernova, 2005 — Arctic English name: Derjugin’s tadpole Norwegian name: vanlig snottfisk Russian name: карепрокт Дерюгина Russian name: (kareprokt Deryugina) Careproctus dubius Zugmayer, 1911 — Arctic English name: dubious Norwegian name: smal snottfisk Careproctus kidoi Knudsen & Møller, 2008 — Mainly arctic English name: Kido’s snailfish Careproctus knipowitschi Chernova, 2005 — Arctic English name: Knipowitsch’s tadpole Norwegian name: buttnesesnottfisk Russian name: карепрокт Книповича Russian name: (kareprokt Knipovicha) Careproctus longipinnis Burke, 1912 — Arctic–boreal Atlantic English name: longfin snailfish

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French name: limace à longues nageoires Russian name: длиннокрылый карепрокт Russian name: (dlinnokryliy kareprokt) Careproctus micropus (Günther, 1887) — Arctic English name: smalleye tadpole Norwegian name: liten snottfisk Russian name: исландский карепрокт Russian name: (islandskiy kareprokt) Careproctus reinhardti (Krøyer, 1862) — Arctic English name: sea tadpole French name: petite limace de mer Norwegian name: nordlig ringbuk Russian name: карепрокт Рейнхардта Russian name: (kareprokt Reynkhardta) Careproctus solidus Chernova, 1999 — Arctic. Known only from one specimen (the holotype) English name: thickset tadpole Careproctus sp. cf. rastrinus (Orr et al.) — Arctic–boreal Pacific? Unidentified Careproctus species (Rand and Logerwell 2010) atlanticus (Jordan & Evermann, 1898) — Mainly boreal western Atlantic English name: seasnail French name: limace atlantique Russian name: американский липарис Russian name: (amerikanskiy liparis) Liparis bathyarcticus Parr, 1931 — Mainly arctic; circumpolar English name: nebulous snailfish French name: limace nébuleuse Norwegian name: pukkelringbuk Russian name: глубинный липарис Russian name: (glubinniy liparis) Liparis bristolensis (Burke, 1912) — Mainly boreal Pacific. of L. tunicatus? English name: Bristol snailfish Liparis callyodon (Pallas, 1814) — Boreal Pacific English name: spotted snailfish Russian name: трёхзубый липарис Russian name: (tryokhzubiy liparis) Krøyer, 1847 — Arctic; circumpolar English name: gelatinous seasnail French name: limace gélatineuse Norwegian name: polarringbuk

Russian name: чернобрюхий липарис Liparis fabricii Russian name: (chernobryukhiy liparis) Liparis gibbus Bean, 1881 — Mainly arctic; circumpolar English name: variegated snailfish

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French name: limace marbrée Russian name: горбатый липарис Russian name: (gorbatiy liparis) Liparis marmoratus Schmidt, 1950 — Arctic-boreal Pacific. Synonym of L. tunicatus? English name: festive snailfish Russian name: мраморный липарис Russian name: (mramorniy liparis) Reinhardt, 1836 — Arctic; circumpolar English name: snailfish French name: limace des laminaires Norwegian name: tangringbuk Russian name: арктический липарис Russian name: (arkticheskiy liparis) Paraliparis bathybius (Collett, 1879) — Arctic English name: black seasnail French name: limace noir Norwegian name: svart ringbuk Russian name: полярный паралипарис Russian name: (polyarniy paraliparis) Paraliparis violaceus Chernova, 1991 — Arctic. Known only from one specimen (the holotype) Russian name: фиолетовый паралипарис Russian name: (fioletoviy paraliparis) Rhodichthys regina Collett, 1879 — Arctic English name: threadfin seasnail French name: limace à filaments Norwegian name: kongeringbuk Russian name: королевский родихт Russian name: (korolevskiy rodikht)

Order (perches) 422 ZOARCIDAE (eelpouts) GYMNELINAE Gymnelus andersoni Chernova, 1998 — Arctic Norwegian name: andersonålebrosme Russian name: гимнел Андерсона Russian name: (gimnel Andersona) Gymnelus esipovi Chernova, 1999 — Arctic. Synonym of G. retrodorsalis? Russian name: гимнел Есипова Russian name: (gimnel Esipova) Gymnelus hemifasciatus Andriashev, 1937 — Arctic–boreal Pacific English name: halfbarred pout Russian name: большеглазый гимнел Russian name: (bolsheglaziy gimnel)

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Gymnelus obscurus Chernova, 2000 — Boreal Pacific. Synonym of G. viridis? Russian name: тёмный гимнел Russian name: (tyomniy gimnel) Gymnelus retrodorsalis Le Danois, 1913 — Arctic English name: aurora pout French name: unernak aurore Norwegian name: spitsbergenålebrosme Russian name: тонкорукий гимнел Russian name: (tonkorukiy gimnel) Gymnelus taeniatus Chernova, 1999 — Arctic. Known only from one specimen (the holotype). Synonym of G. retrodorsalis? Russian name: ентотелый гимнел Russian name: (lentoteliy gimnel) Gymnelus viridis (Fabricius, 1780) — Mainly arctic English name: fish doctor French name: unernak caméléon Norwegian name: grønlandsålebrosme Russian name: гренландский гимнел Russian name: (grenlandskiy gimnel) LYCODINAE Lycenchelys kolthoffi Jensen, 1904 — Arctic English name: checkered wolf eel French name: lycode quadrillée Norwegian name: marmorert ålebrosme Russian name: пятнистый лиценхел Russian name: (pyatnistiy litsenkhel) Lycenchelys muraena (Collett, 1878) — Arctic English name: moray wolf eel French name: lycode murène Norwegian name: havålebrosme Russian name: муреновидный лиценхел Russian name: (murenovidniy litsenkhel) Lycenchelys paxillus (Goode & Bean, 1879) — Boreal western Atlantic English name: common wolf eel French name: lycode commune Lycenchelys platyrhina (Jensen, 1902) — Arctic Russian name: голый лиценхел Russian name: (goliy litsenkhel) Lycenchelys sarsii (Collett, 1871) — Boreal Atlantic English name: Sars’ wolf eel French name: lycode de Sars Norwegian name: sørlig ålebrosme Russian name: лиценхел Сарса

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Russian name: (litsenkhel Sarsa) adolfi Nielsen & Fosså, 1993 — Arctic English name: Adolf’s eelpout French name: lycode d’Adolf Norwegian name: silkeålebrosme Russian name: ликод Адольфа Russian name: (likod Adolfa) Lycodes brevipes Bean, 1890 — Boreal Pacific English name: shortfin eelpout Russian name: коротконогий ликод Russian name: (korotkonogiy likod) Lycodes esmarkii Collett, 1875 — Mainly boreal Atlantic English name: greater eelpout French name: grand lycode, lycode d’Esmark Norwegian name: ulvefisk Russian name: узорчатый ликод, ликод Эсмарка Russian name: (uzorchatiy likod, likod Esmarka) Lycodes eudipleurostictus Jensen, 1902 — Arctic English name: doubleline eelpout French name: lycode à deux lignes Norwegian name: båndålebrosme Russian name: двупёрый ликод Russian name: (dvupyoriy likod) Lycodes frigidus Collett, 1879 — Arctic English name: glacial eelpout French name: lycode glaciale Norwegian name: arktisk ålebrosme Russian name: абиссальный ликод Russian name: (abissalniy likod) Lycodes gracilis Sars, 1867 — Mainly boreal eastern Atlantic Norwegian name: vanlig ålebrosme Russian name: тонкий ликод Russian name: (tonkiy likod) Lycodes jugoricus Knipowitsch, 1906 — Arctic English name: shulupaoluk French name: lycode plume Russian name: югорский ликод Russian name: (yugorskiy likod) Lycodes lavalaei Vladykov & Tremblay, 1936 — Mainly boreal western Atlantic English name: Newfoundland eelpout French name: lycode du Labrador Norwegian name: labradorålebrosme

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Lycodes luetkenii Collett, 1880 — Arctic English name: Lütken’s eelpout French name: lycode de lütken Norwegian name: lutkenålebrosme Russian name: ликод Люткена Russian name: (likod Lyutkena) Lycodes marisalbi Knipowitsch, 1906 — Amphi-arctic English name: eelpout French name: lycode de la mer blanche Russian name: беломорский ликод Russian name: (belomorskiy likod) Lycodes mcallisteri Møller, 2001 — Arctic English name: McAllister’s eelpout French name: lycode de McAllister Lycodes mucosus Richardson, 1855 — Arctic English name: saddled eelpout French name: lycode à selles Russian name: слизистый ликод (slizistiy likod) Lycodes paamiuti Møller, 2001 — Arctic English name: Paamiut eelpout French name: lycode de Paamiut Norwegian name: paamiutålebrosme Lycodes palearis Lycodes palearis Gilbert, 1896 — Mainly boreal Pacific English name: wattled eelpout Russian name: гребенчатый ликод Russian name: (grebenchatiy likod) Lycodes pallidus Collett, 1879 — Arctic-boreal Atlantic English name: pale eelpout French name: lycode pâle Norwegian name: blek ålebrosme Russian name: бледный ликод Russian name: (bledniy likod) Lycodes polaris (Sabine, 1824) — Arctic; nearly circumpolar English name: polar eelpout, Canadian eelpout French name: lycode polaire Norwegian name: polarålebrosme Russian name: полярный ликод Russian name: (polyarniy likod) Lycodes raridens Taranetz & Andriashev, 1937 — Mainly boreal Pacific English name: marbled eelpout Russian name: редкозубый ликод Russian name: (redkozubiy likod)

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Lycodes reticulatus Reinhardt, 1835 — Arctic English name: Arctic eelpout French name: lycode arctique Norwegian name: nettålebrosme Russian name: сетчатый ликод Russian name: (setchatiy likod) Lycodes rossi Malmgren, 1865 — Arctic English name: threespot eelpout French name: lycode à trois taches Norwegian name: nordlig ålebrosme Russian name: ликод Росса Russian name: (likod Rossa) Lycodes sagittarius McAllister, 1976 — Arctic English name: archer eelpout French name: lycode à arc Lycodes seminudus Reinhardt, 1837 — Arctic English name: longear eelpout French name: lycode à oreilles Norwegian name: halvnaken ålebrosme Russian name: полуголый ликод Russian name: (polugoliy likod) Lycodes squamiventer Jensen, 1904 — Arctic English name: scalebelly eelpout Norwegian name: skjellålebrosme Russian name: чешуебрюхий ликод Russian name: (cheshuyebryukhiy likod) Lycodes turneri Bean, 1879 — Mainly arctic Pacific English name: estuarine eelpout Russian name: эстуарный ликод Russian name: (estuarniy likod) Reinhardt, 1831 — Mainly boreal western Atlantic English name: checker eelpout French name: lycode à carreaux Lycodonus flagellicauda (Jensen, 1902) — Arctic Norwegian name: spisshalet ålebrosme Russian name: тонкохвостый ликодон Russian name: (tonkokhvostiy likodon) Lycodonus mirabilis Goode & Bean, 1883 — Mainly boreal western Atlantic English name: chevron scutepout French name: lycaspine à chevrons Russian name: червонный ликодон Russian name: (chervonniy likodon)

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ZOARCINAE viviparus (Linnaeus, 1758) — Mainly boreal eastern Atlantic English name: Norwegian name: ålekvabbe Russian name: европейская бельдюга Russian name: (evropeyskaya beldyuga)

417 STICHAEIDAE () STICHAEINAE Eumesogrammus praecisus (Krøyer, 1836) — Arctic–boreal Pacific and western Atlantic English name: fourline snakeblenny French name: quatre-lignes atlantique Russian name: шипохвостый стихей Russian name: (shipokhvostiy stikhey) Stichaeus punctatus (Fabricius, 1780) — Arctic–boreal Pacific and western Atlantic English name: Arctic shanny French name: stichée arctique Russian name: пятнистый стихей Russian name: (pyatnistiy stikhey) LUMPENINAE Acantholumpenus mackayi (Gilbert, 1896) — Mainly boreal Pacific English name: blackline French name: terrassier à six lignes Russian name: колючий люмпен Russian name: (kolyuchiy lyumpen) Anisarchus medius (Reinhardt, 1837) — Arctic–boreal; circumpolar English name: stout eelblenny French name: lompénie naine Norwegian name: rundhalet langebarn Anisarchus medius Russian name: ильный люмпен Russian name: (ilniy lyumpen) maculatus (Fries, 1838) — Arctic–boreal Pacific and Atlantic English name: daubed shanny French name: lompénie tachetée Norwegian name: tverrhalet langebarn Russian name: пятнистый лептоклин Russian name: (pyatnistiy leptoklin) Lumpenus fabricii Reinhardt, 1836 — Arctic–boreal English name: slender eelblenny French name: lompénie élancée Norwegian name: arktisk langebarn Russian name: люмпен Фабриция Russian name: (lyumpen Fabritsiya)

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Lumpenus lampretaeformis (Walbaum, 1792) — Mainly boreal Atlantic English name: snakeblenny French name: lompénie-serpent Norwegian name: langhalet langebarn Russian name: миноговидный люмпен Russian name: (minogovidniy lyumpen) Lumpenus sagitta Wilimovsky, 1956 — Boreal Pacific English name: snake prickleback Russian name: стреловидный люмпен Russian name: (strelovidniy lyumpen) CHIROLOPHINAE ascanii (Walbaum, 1792) — Boreal Atlantic English name: Atlantic warbonnet French name: toupet marbré Norwegian name: hornkvabbe Russian name: европейская мохоголовая собачка Russian name: (evropeyskaya mokhogolovaya sobachka) Chirolophis decoratus (Jordan & Snyder, 1902) — Boreal eastern Pacific English name: Chirolophis snyderi (Taranetz, 1938) — Mainly boreal Pacific English name: bearded warbonnet Russian name: северная мохоголовая собачка Russian name: (severnaya mokhogolovaya sobachka) XIPHISTERINAE Alectrias alectrolophus (Pallas, 1814) — Boreal Pacific English name: stone cockscomb Russian name: бурый петушок Russian name: (buriy petushok)

419 (gunnels) PHOLINAE fasciata (Bloch & Schneider, 1801) — Arctic–boreal English name: banded gunnel French name: sigouine rubanée Russian name: полосатый маслюк Russian name: (polosatiy maslyuk) Pholis gunnellus (Linnaeus, 1758) — Mainly boreal Atlantic English name: French name: sigouine de roche Norwegian name: tangsprell Russian name: атлантический маслюк Russian name: (atlanticheskiy maslyuk)

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APODICHTHYINAE Rhodymenichthys dolichogaster (Pallas, 1814) — Mainly boreal Pacific English name: stippled gunnel Russian name: северный одноцветный маслюк Russian name: (severniy odnotsvetniy maslyuk)

420 (wolffishes) denticulatus Krøyer, 1845 — Arctic–boreal Atlantic English name: French name: loup à tête large Norwegian name: blåsteinbit Russian name: синяя зубатка Russian name: (sinyaya zubatka) Anarhichas lupus Linnaeus, 1758 — Mainly boreal Atlantic English name: French name: loup atlantique Norwegian name: gråsteinbit Russian name: полосатая зубатка Russian name: (polosataya zubatka) Olafsen, 1772 — Mainly boreal Atlantic English name: spotted wolffish French name: loup tacheté Norwegian name: flekksteinbit Russian name: пятнистая зубатка, пёстрая зубатка Russian name: (pyatnistaya zubatka, pyostraya zubatka) Pallas, 1814 — Mainly boreal Pacific English name: Bering wolffish French name: loup de Béring Russian name: восточная зубатка Russian name: (vostochnaya zubatka) Anarrhichthys ocellatus Ayres, 1855 — Boreal eastern Pacific English name: wolf-eel Russian name: угревидная зубатка Russian name: (ugrevidnaya zubatka)

422 ZAPRORIDAE (prowfishes) Zaprora silenus Jordan, 1896 — Boreal Pacific English name: prowfish Russian name: запрора Russian name: (zaprora)

441 AMMODYTIDAE (sand lances) Ammodytes americanus DeKay, 1842 — Boreal western Atlantic

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English name: American Russian name: американская песчанка Russian name: (amerikanskaya peschanka) Ammodytes dubius Reinhardt, 1837 — Arctic–boreal western Atlantic English name: northern sand lance French name: lançon du nord Ammodytes hexapterus Pallas, 1814 — Arctic–boreal Pacific English name: Pacific sand lance, stout sand lance French name: lançon gourdeau Russian name: северная тихоокеанская песчанка Russian name: (severnaya tikhookeanskaya peschanka) Ammodytes marinus Raitt, 1934 — Mainly boreal eastern Atlantic English name: lesser sand-eel Norwegian name: havsil Russian name: европейская многопозвонковая песчанка Russian name: (evropeyskaya peschanka) Ammodytes tobianus Linnaeus, 1758 — Mainly boreal eastern Atlantic English name: small sand-eel French name: équille Norwegian name: sandgrevling Russian name: песчанка Russian name: (peschanka)

475 SCOMBRIDAE ( and tunas) Scomber scombrus Linnaeus, 1758 — Boreal Atlantic English name: Atlantic mackerel Norwegian name: makrell Russian name: атлантическая скумбрия Russian name: (atlanticheskaya skumbriya)

Order Pleuronectiformes (flatfishes) 493 (righteye flounders) HIPPOGLOSSINAE hippoglossus (Linnaeus, 1758) — Boreal Atlantic English name: Norwegian name: kveite Russian name: атлантический палтус Russian name: (atlanticheskiy paltus) Hippoglossus stenolepis Schmidt, 1904 — Boreal Pacific English name: Russian name: тихоокеанский белокорый палтус Russian name: (tikhookeanskiy belokoriy paltus)

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Reinhardtius (Walbaum, 1792) — Arctic–boreal English name: French name: flétan du Groenland Norwegian name: blåkveite Russian name: чёрный (синекорый) палтус Russian name: (chyorniy (sinekoriy) paltus)

HIPPOGLOSSOIDINAE Reinhardtius hippoglossoides Hippoglossoides platessoides (Fabricius, 1780) — Mainly boreal Atlantic English name: American , Canadian plaice French name: plie canadienne Norwegian name: gapeflyndre Russian name: камбала-ёрш Russian name: (kambala-yorsh) Hippoglossoides robustus Gill & Townsend, 1897 — Arctic–boreal Pacific English name: Bering flounder French name: plie de Béring Russian name: северная палтусовидная камбала Russian name: (severnaya paltusovidnaya kambala) polyxystra Orr & Matarese, 2000 — Boreal Pacific English name: northern rock sole Russian name: северная двухлинейная камбала Russian name: (severnaya dvukhlineynaya kambala) aspera (Pallas, 1814) — Mainly boreal Pacific English name: yellowfin sole Russian name: желтопёрая камбала Russian name: (zheltopyoraya kambala) Limanda limanda (Linnaeus, 1758) — Mainly boreal eastern Atlantic English name: dab Norwegian name: sandflyndre Russian name: лиманда, ершоватка Russian name: (limanda, ershovatka) Limanda proboscidea Gilbert, 1896 — Mainly boreal Pacific English name: longhead dab French name: limande carline Russian name: хоботная камбала Russian name: (khobotnaya kambala) Limanda sakhalinensis Hubbs, 1915 — Mainly boreal Pacific English name: Sakhalin sole Russian name: сахалинская камбала Russian name: (sakhalinskaya kambala)

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Liopsetta glacialis (Pallas, 1776) — Mainly arctic English name: French name: plie arctique Norwegian name: arktisk flyndre Russian name: полярная камбала Russian name: (polyarnaya kambala) kitt (Walbaum, 1792) — Mainly boreal eastern Atlantic English name: lemon sole Norwegian name: lomre Russian name: малоротая камбала Russian name: (malorotaya kambala) flesus (Linnaeus, 1758) — Boreal eastern Atlantic English name: flounder, Norwegian name: skrubbe Russian name: речная камбала Russian name: (rechnaya kambala) Platichthys stellatus (Pallas, 1787) — Arctic–boreal Pacific English name: starry flounder French name: flet étoilé Russian name: звёздчатая камбала Russian name: (zvyozdchataya kambala) platessa Linnaeus, 1758 — Boreal eastern Atlantic English name: plaice Norwegian name: rødspette Russian name: морская камбала Russian name: (morskaya kambala) Pleuronectes quadrituberculatus Pallas, 1814 — Mainly boreal Pacific English name: Alaska plaice Russian name: желтобрюхая камбала Russian name: (zheltobryukhaya kambala)

Acknowledgments This edition of the list is a contribution by the collaborators and the RUSALCA and TUNU programs to the CAFF, Circumpolar Biodiversity Monitoring Program. The RUSALCA (Russian-American Long- Term Census of the Arctic) is a multidisciplinary cooperative program of the U.S. National Oceanic and Atmospheric Administration and the Russian Academy of Sciences which focuses on the Pacific-Arctic region. The TUNU-Programme of the UiT–The Arctic University of Norway, Tromsø, Norway, focuses on Atlantic-Arctic fishes. The photographs (except for the cover) are by C.W.M.

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