AMERICAN MUSEUM NOVITATES Published by Number 1259 THE AMERICAN MUSEUM OF NATURAL HISTORY August 17, 1944 New York City

NOTES ON THE GOMPHINAE () WITH DESCRIPTIONS OF NEW BY ELSIE BROUGHTON KLOTS1

A study of gomphine wing venation ex- the narrow green collar; a narrow ante- tending over a period of 15 years or more humeral stripe bent forward at its upper has brought to my attention many speci- end and slightly widened at its lower end. mens of unusual interest. It seems ad- Mesepimeron obscured but apparently visable at this time to publish notes on with two irregular longitudinal pale stripes. some of these under one title, though they Metepimeron with a broad pale band bear no relationship other than that the covering most of its area. specimens are to be found in the American Legs pale with black spines; tibiae and Museum of Natural History. They are tarsi darker. Wings hyaline with black as follows: veins and tawny stigma; widest at the proximal end of the stigma. Antenodal 1. A possible new species of Diaphlebia cross veins of the fore wing 16-17, the first from Venezuela, with notes on the venation and the seventh thickened; of the hind of Diaphlebia and Desmogomphus. wing 12-13, the first and the sixth thick- 2. A new and species from Peru. ened. Postnodal cross veins of the fore 3. A new species of Phyllogomphus wing 15, of the hind wing 14. Cross veins from the Congo, with notes on Phyllo- under the stigma, in addition to the brace gomphus coloratus Kimmins. vein, six to seven in the fore wing, eight in 4. A species of Notogomphus from Ethi- the hind. Fore and hind wing discoidal opia which differs venationally from the triangle crossed; subtriangle and supra- genotype. triangle free. Costal side of the fore wing 5. A new species of Oxygomphus from triangle slightly longer than the proximal the Congo. side; costal side of the hind wing triangle more than twice as long as the proximal Diaphlebia species side. Distal side ef both triangles broken. Figures .5, 6, 18 Discoidal field of both wings of two rows LENGTH: Abdomen, including append- of cells, in the fore wing increasing to three ages, 36.8 mm.; appendages, 2 mm.; cells at the level of the nodus, and in the hind wing, 32 mm.; stigma, 5.5 mm. hind wing increasing halfway between the FEMALE: Lips and head pale brownish; median fork and the nodus. Cross veins labrum with lateral margins and a minute before the median fork seven in the fore median spot black; postclypeus with a wing, four in the hind; median fork narrow black mark on the anterolateral slightly asymmetrical. Sectors of the corners; ocelli ringed with yellow; vertex arculus rising just below its middle and dark brown, prominent, coming to a point separated from each other by a distance behind each rear ocellus, these two peaks greater than the width of either. connected by a concave ridge; border of Abdomen brown, marked with yellow occiput concave. as follows: segment 2 with a narrow Prothorax obscured. Synthorax pale trilobed spot on the middorsal line and a brown with a pair of green dorsal stripes ventral lateral band extending the length divergent forward and separated from of the segment; segment 3 with the ventral 1 Finch Junior College, New York City. lateral band its full length but reduced 2 AMERICAN MUSEUM NOVITA TES [No. 1259

on the apical fourth; segments 4 to 7 with of the fore wing are the thickened ones basal twin spots fused with the pale ventral instead of the first and fifth; the fore wing margins, on 4 to 6 the spots are small, but triangles are not so far removed from on 7 they extend one-third the segment the arculus as the length of the proximal length and almost meet on the middorsal side of the triangle; the venation is richer line. Segments 8 to 10 all brown. Apical. throughout. It agrees with only three carinae of 3 to 10 narrowly black and on of the characters in Needham's verifica- each of these segments a minute, black tion table (1940, p. 389): the absence lateroventral spot on the intersegmental of the basal subcostal cross veins (and membrane. Appendages pale brown, even this character may not hold, for there black tipped, longer than segment 10 and is a basal subcostal in one of the hind wings about as long as segment 9. Vulvar scale of this specimen), the possession of two elongate, tapering, slightly protuberant, cell rows in the discoidal field of the tri- and bifid to two-thirds its length; the two angle of the fore wing, and the asymmetry lobes approximated, reaching almost to the of the median fork. It thus invalidates end of segment 9. the presence or absence of cross veins in MATERIAL: Female, Mt. Duida, Vene- the fore wing triangle and the number of zuela, March 3, 1929 (G. H. H. Tate), in postanal cells as generic characters for dis- the American Museum of Natural History. tinguishing Diaphlebia from Desmogorn- Investigation has shown that there are phus, and raises the question as to whether very few specimens of Diaphlebia known. these two genera can be distinguished on Since Selys' descriptions, only D. nexans venational characters. The triangles of Calvert has been described, and no other D. angustipennis in the Cornell collection references have been made to the two are longer than those of the type of Des- Selysian species. In fact, when Selys mogomphus, but an analysis of the five described semilibera (1869) he suggested specimens of Desmogomphus which I it might prove to be another angustipennis, have been able to study shows some varia- its chief apparent difference being the tion in the character. If a value of 10 be absence of cross veins in all triangles ex- ascribed to the length of the proximal side cept the discoidal triangles of the hind wing. of the triangle, the ratios of the proximal, In this respect the American Museum speci- the costal, and the distal sides are as men is more like semilibera, but it is smaller, follows: Desmogomphus, fore wing 10: 10- has a richer wing venation, and differs in 12: 11-12, hind wing 10: 16-21: 16-24; the coloration of the abdomen. Diaphlebia sp., fore wing 10:13:15, hind It differs from angustipenni.s Selys in wing 10:23:26; Diaphlebia angustipennis, its richer wing venation, the presence of fore wing 10: 14: 15, hind wing 10:20:23. black marks on the labrum and postcly- Any estimate of the proportions of the peus, and the distinct trilobed spot on triangles of nexans taken from the figure abdominal segment 2. It differs from the would necessarily be very inexact, but the description of the male of nexans Calvert triangles do not seem to be so long as these in its larger size, the richer wing venation, other representatives of the genus. the slightly longer triangles, and the The relation of the anal crossing and the coloration of the abdomen. origin of A3 is also variable. In Diaphiebia, Although this specimen in the Ameri- sp. it rises virtually under ac, in angusti- can Museum does show these differences pennis it rises a little beyond ac, and in from the other known species, there is too Desmogomphus A3 rises before ac in eight little comparative material available to of the hind wings and almost underneath warrant its description as new. It does in two. add, however, to our knowledge of the Antenodals of the fore wing in the type venational range of the genus. It agrees of Desmogomphus number 12-13, but of the with Calvert's characterization of the 10 wings which I have studied only one genus (1903) differing only in minor re- has 12 and three have 13, while four have spects: the first and seventh antenodals 14 and two have 15. In Diaphlebia they 19441 NOTES ON THE GOMPHINAE (ODONATA) 3 range from the 13-14 of described species in the distal position of the arculus, the to the 16-17 in this specimen. narrow anal area of the fore wing, the slight The size of the stigma seems to remain hooking of the vein tips, the asymmetry of a good distinguishing character. It is one- the median fork, and the large thick stigma. eighth the wing length in Desmogomphus The unusual character of the genitalia of and one-sixth in Diaphlebia. the second abdominal segment, as described The hooking of the vein tips in Dia- for the species, is probably not generic: it phlebia also seems to be a good character. has been observed in thegenus Progomphus. This can best be described, though in- adequately, as a downward curve of the Peruviogomphus moyobambus, new sectors M2, R5, and R., beginning at the species level of the proximal end of the stigma and Figures 11-16 continuing evenly to the wing margin. In the dried type specimen, the colors This curve accompanies a blunt-tipped are obscured and badly faded. The eyes wing which is usually widest just before show distinct traces of iridescent emerald the stigma. In the specimens that I green, and there is some indication that have seen of these two genera this serves these colors may also have been character- as a distinguishing character. istic of the thorax. LENGTH: Abdomen, including appen- PERUVIOGOMPHUS, NEW GENUS dages, 33 mm.; hind wing, 24.5 mm.; Basal subcostal cross vein present; stigma, 2.3 mm. arculus at, or slightly beyond, the second Face, frons, vertex, and occiput pale antenodal cross vein; median fork asym- green, the labrum bordered and possibly metrical; nine median cross veins in the crossed with yellow; the postclypeus with fore wing, six in the hind wing; fore wing anterior margin narrowly bordered with triangle directed outward, its costal side yellow and the outer corners broadly the shortest and its proximal and distal yellow. Frons low, its upper border deeply sides about equal to each other; one row concave on the median line. Vertex green, of cells beyond the fore wing triangle as the ocelli and base of antennae ringed with far out as the median fork; one paranal yellow; the prominent ridges around the row in the fore wing widening to two rows rear ocelli and across the rear border of the behind Cu2. Hind wing triangle slightly vertex yellow. Occiput green, its ele- broken on its distal side and with one cross vated border nearly straight and narrowly vein; A3 rising before the anal crossing; yellow. A2 rising from under the subtriangle. Thorax pale, the middorsal black stripe Stigma braced and surmounting four addi- but slightly wider than the pale green stripe tional cross veins. on either side and connected above with GENOTYPE: Peruviogomphus moyobam- the wide, black antehumeral; humeral bus, new species. suture with a narrow black stripe which is This genus resembles Archaegomphus connected above and below with the and Agriogomphus in having a single row antehumeral. Sides all pale green, yellow of cells beyond the triangle of the fore wing in the sutures and on the carinae and but otherwise seems to have no affinities yellowish ventrally and posteriorly. with them. It resembles the Progomphini Legs yellow, femora streaked with brown in having crossed triangles (in this speci- on the outside and with two double rows of men only the supratriangles and the tri- small irregular black spines; tibiae and angle of the hind wing are crossed), a tarsi darker. Wings hyaline; antenodal basal subcostal cross vein, a large number cross veins in the fore wing 17, postnodals of median cross veins, no extra cubito-anal 12; antenodals in the hind wing 13; cross veins, nodus beyond the middle of postnodals 12; cells under the stigma four. the wing, the stigma with a brace vein, and Abdomen with colors obscured. Seg- a tendency for vein A2 to converge with As ments 1 and 2 brown with pale markings distally. It approaches the Epigomphini on the sides, including the auricles which 4 AMERICAN MUSEUM NOVITATES [No. 1259 are prominent and black bristled on their an apical ring of yellow. Occiput brown, posterior surface. Segments 2 to 6 with a its elevated rear border sinuous. wide dorsal brown band forming a complete Prothorax brownish. Synthorax brown, ring at the apex and partially invaded at marked with green as follows: the in- the median carina with the yellow of the verted "7," which is broadly interrupted venter. This brown is finely but distinctly at the corners and the cross piece of which divided by a yellow line on the middorsal is'barely more than a hair line in width; carina. Dorsum of 2 with a broad bilobed a small spot on the basal half of the mid- yellow spot. Segments 3 to 7 have minute dorsal carina; two broad oblique bands blunt black spines scattered along on either on the sides, one under each wing, the first side of the middorsal line. Segment 7 with narrowly bordered with black and the the basal half diffusely yellow; 8 to 10 second with a broad black band along its brown with diffuse dorsal and lateral pale anterior margin. This black band is spots. Segments 7 to 9 with a distinct continuous across the dorsal end of the black spot on the apical carina near the pale band and fades to yellow in the suture ventral 'margin; segment 10 with apical preceding it. There is a notal green spot carina all black. Appendages brown, between each pair of wings. black-tipped; longer than segment 10. Femora brown, black at the apices; Penis with a pair of long, downward-curved tibiae and tarsi black, tibiae paler basally. bristles extending well beyond the end of Trochanters and femora with many small, the second segment. blunt, scattered spines. TYPE MATERIAL: Holotype, male, Wings uniformly enfumed with yellow- Moyobamba region, Peru, March, 1936 ish, deeper at the wing bases, with black (G. Klug), in the American Museum of veins and stigma. Antenodals of the fore Natural History. wing 17, the first and fifth thickened; postnodals 15. Antenodals of the hind wing 14, postnodals 15. No basal sub- Phyllogomphus annulus, new species costal cross veins; arculus arising between Figures 8, 10 the first and second antenodal cross veins; LENGTH: Abdomen, including append- stigma covering five cells. Hind wing with ages, 61 mm.; hind wing, 48 mm.; stigma, anal loop of four cells; six postanal cells; 5.5 mm. Width of hind wing at level of A3 rising under the anal crossing. nodus, 12.8 mm. Abdominal segment 1 brown; segment Face and head mainly brown marked 2 brown with a pair of green, lateral, with green. Labium brown, its anterior basal spots, diffuse and tapering beyond half paler; mandibles 'pale green with the median carina, and with ventral mar- brown tips; labrum with a'pair of pale gins pale as far as the carina; segment 3 green spots and with the lateral margins brownish with a complete basal yellow ring bordered with darker brown; anteclypeus extending to the median carina, or one- with its lower third horizontal, except third the segment length, and continuous on the median line, and with its upper two- with the broad yellow ventral margins; thirds vertical and green; postclypeus segments 4, 5, and 6 all dark, brownish brown with a large green spot on each side black except for the pale ventral margins; and with nasal punctures' yellowish, the segment 7 brown with a yellow basal half pale yellowish continuing laterally along ring extending halfway down the sides and the anterior border; frons 'brown with a to one-fourth the segment length, and with pale green band across 'the top coverinmg a small rectangular projection on the mid- about one-third 'of the anterior surface dorsal line, ventral margins pale; seg- and one-half of the upper surface. Vertex ments 8 and 9 light yellowish brown on brown, black along the eyes; the transverse the side's, darker on the dorsum; leaves of ocellary ridge with e6ds bent backward to segment 8 pleated, black, extending the run' along the rear border iof the ocelli; length of segment 9 and 4 mm. wide at antennae brown, second-segment black with their widest 'point; segment 10''light red- 1944] NOTES ON THE GOMPHINAE (ODONATA) 5 dish brown. Segment 9 is one-half as long is the only one described. It has the forked as segment 8 and two-thirds as long as seg- tip reaching nearly to the end of segment ment 10. Segment 10, which bends down- 10, whereas this specimen has it reaching ward, has a conspicuous dorsal carina with a only one-third the length of segment 10. slight arch in its basal fourth, striking In the American Museum there are lateral carinae, and many minute trans- three males of P. coloratus Kimmins, col- verse encircling ridges. lected in the Cameroons, which agree with Vulvar scale extraordinarily long, broad, the original description except in the color scoop shaped to the end of segment 9 and pattern of abdominal segment 2. This then narrowed to a slender projection segment has a middorsal lanceolate pale which ends in a broader, black-tipped fork spot extending to the carina, instead of a which reaches to about one-third the length thin pale dorsal streak. A female collected of segment 10. The scoop stands out from at the same time is larger and heavier but the abdomen, but the slender projection differs in color only inasmuch as the poor bends back and almost touches segment 10. preservation of color conceals the original TYPE MATERIAL: Holotype, female, pattern of the face and the dorsum of the Medji, Congo, June 26, 1910 (Lang and thorax. It has the vulvar scale very long, Chapin), in the American Museum of reaching nearly to the end of segment Natural History. 10 (figs. 7, 9), thus resembling the scale This beautifully preserved specimen is of the MacLachlan female. clearly distinguishable from all described females. It differs from aethiops Selys in Notogomphus dorsalis Selys having but three pale stripes on each side Figure 19 of the middorsal carina of the thorax. A female collected in Addis Ababa, It differs from the description of coloratus Ethiopia, July 5, 1920, altitude 5000 feet, (Kimmins, 1931, did not describe the is so different venationally from my con- female because of its poor preservation) ception of this genus that I at first thought as follows: the frons is brown with a green to describe it as a new genus. It answers band across the top instead of green mar- the description of this species so closely, gined with brown; the legs have consider- however, that I shall not add further to able brown on the femora and tibiae; the complexities of the Noto-Podogomphus abdominal segment 3 has a distinct and group by describing it as new. But I do complete basal yellow ring instead of wish to note here the peculiarities of the lateral spots; segments 4, 5, and 6 are all venation in the hope that some one with brownish black with no lateral basal pale more material will make a detailed study of spots; and the lamellae do not seem to be the venational characters and define the "larger than in aethiops." It differs from limits of variation within this and closely the description of selysi Schouteden (1933) related genera. by the greater extent of brown on the face The basal subcostal cross vein is present; and head, by the yellow ring on segment 3, brace vein present and strongly aslant; and by the absence of orange yellow spots stigma surmounting only two and one-half on segments 4, 5, and 6. It differs from the cells; median fork symmetrical with two MacLachlan female, described as aethiops median cross veins between it and the by Selys (1878) and shown by Longfield arculus; triangle large, slightly broken (1936) not to be aethiops, in the greater on the outer side, that of the fore wing extent of color on the wings (a character but little larger than the subtriangle and which is, of course, variable), the greater with its proximal side but very slightly extent of brown on the face, the basal longer than its costal; sectors of the ring on segment 3, and the absence of basal arculus widely separated; two paranal spots on segments 4, 5, and 6. rows and three rows behind Cu2 of the The vulvar scale should be a good cri- fore wing. Hind wing with one cubito-anal terion for identification, but unfortunately cross vein; five double postanal rows be- the vulvar scale of the MacLachlan female tween A1 and A2; A3 well before the anal 6 AMERICAN MUSEUM NOVITATES [No. 1259 crossing; anal crossing distant from the tending from above the arms of the crest subtriangle by its own length; Cu1 and Cu2 down across the prothorax, gradually separated by four cells at the margin; anal widening to reach its greatest width before loop of two or three cells. the collar, each of these bands wider than I very much doubt if this and Notogom- the middorsal yellow; a narrow band be- phus ruppelyi Selys (as figured by Ris, 1909, hind the humeral suture; faint narrow fig. 5) are congeneric. However, wings brown lines along the alar carirae and in figured by Sjostedt (1909, pl. 2, figs. 3-5), the sutures. Schouteden (1934, fig. 69), and Campion Coxae yellow; femora yellow, lined with (1923, figs. 1, 3) show intermediate varia- black; tibiae and tarsi black. (Hind fe- tions. It does seem to differ from them mora missing.) all in the short bridge (a character which I Abdomen with a middorsal yellow band have not studied enough as yet to be able extending from segment 1 to segment 8, to define), the short stigma, and the double narrowly interrupted at the apices of all row of cells between A2 and A3. The wide segments and at the median carina of seg- area indicated by this character is due, ment 7; separated from the yellow of the partially of course, to the sex of the speci- sides and venter by a black band which men. runs full length of the abdomen on each I add a brief description of the specimeen side, the two approximating on the apical for the benefit of those who may be able fourth of segment 8 and fusing to cover to compare it with type material; there is the dorsum of segment 9. Segment 10 always the possibility that I have incor- brownish with a pair of black spots on the rectly determined it. base of the segment. Appendages brown. LENGTH: Abdomen, including appen- Vulvar scale broad, the width of the seg- dages, 29mm.; hind wing, 26mm.; stigma, ment notched at- the tip and extending not 2.6 mm. Width of hind wing at the level more than one-fifth the length of segment 9. of the nodus, 9 mm. Labium brown, median lobe paler bas- OXYGOMPHUS LACROIX1 ally; labrum yellow, entirely bordered Fraser, the first to describe this genus, with brown and crossed with a brown band says that it is distinguished from all other which is marked in the center by a yellow genera by segment 10 which is longer than depression; base of mandibles pale yel- segment 9 and abruptly constricted, cylin- low; genae dark brown; anteclypeus drical in the second half and telescoped into brown with a pair of round yellow spots 9; by the occiput which is elevated, re- in the middle, almost touching; postcly- curved forward and surmounted by two or peus and frons yellow with fine brown retic- more spines; and by the posterior hamules ulations, the brown more dense towards which project far out and forward toward the clypeal suture and in the depressions of the base of the abdomen. the nasal punctures; frons high and swol- A specimen in the American Museum len across the crest, which is slightly collection has genitalia so like those of notched in the middle and punctuated with Oxygomphus agilis (as figured by Fraser) six or seven sharp black elevated points on that I place it in that genus without ques- either side; vertex dark brown, slightly tion in my own mind. Its unique tenth lighter behind the postocellary ridge, which segment may well be that described by is curved to rearward on the midline-; occi- Fraser, but its simple occiput indicates put convex, very low, light brown, fringed 1 Martin (1908) described Notogomphlus agilis as with long black hair. Rear of eyes yellow-, having a recurved occiput and the last three segments of the abdomen fused. Ris (1909) expressed the ish brown with a wide black border along opinion that Notogomphus agilis Martin was not a the eye margin on either side of the occiput. true Notogomphus. Fraser (1936) ascertained that Martin's Notogomphus agilis and the specimen in Prothorax black with a wide middorsal Martin's collection labeled Oxygovaphus agilis were the same. When Lacroix described Oxygomphus yellow band. Thorax yellow, marked with martininus (1921) the genus had not been described, brown as follows: the V of the crest, a but since he compared his specimen with the type of 0. agilis this genus is attributed to him by Fpraser broad band on each side of the dorsum ex- (1936). 1944] NOTES ON THE GOMPHINAE (ODONATA) 7 either that his description of the genus will Wings hyaline, veins, including the costa have to be made more elastic or that a new dark brown; stigma light brown, paler in genus should be described for this species the apical third, surrounded by dark veins. and possibly martininus Lacroix. Antenodals 13-14 in the fore wing, 11 in the hind; postnodals 10. Oxygomphus chapini, new species Abdomen brownish with color pattern Figures 1-4, 17 obscured. Segment 1 yellowish; segment 2 LENGTH: Abdomen, including appen- with prominent auricles and with a narrow, dages, 38 mm.; hind wing, 29 mm.; stigma, irregular, middorsal, pale, lanceolate band 2.8 mm. four-fifths the segment length; segment Face and frons pale greenish, faintly 3 with an attenuated middorsal triangle margined with yellow; labrum with a me- three-fourths the segment length, very ob- dian, transversely oval depression. Frons scure, in some lights scarcely visible, with low and rounded over the region of the median and apical carinae darker; seg- crest, slightly bulbous below the antennae. ments 4, 5, and 6 reddish brown; segments Vertex brown, the area around the median 7, 8, and 9 black on the dorsum, 7 with a ocellus and across the front yellow. An- brown basal ring one-fourth the segment tennae brown except for the pale green length; segments 8 and 9 with expanded basal segment. Ocellary ridges completely lateral margins bright yellow and with a surrounding the rear ocelli, most con- middorsal yellow spot on the apical inter- spicuous on the outer side along the mar- segmental membrane; segment 10 brown- gin of the compound eyes. Rear of vertex ish with the basal half black and an irregu- sloping to the low ridge before the occiput. lar apical black margin; segment 10 a Occiput brown, almost straight, with only little longer than 9, abruptly constricted, a slight depression on the median line. slender and cylindrical at base as though it Rear of eyes brownish with a conspicuous might be telescoped into 9, encircled yellow spot on the lateral margins. basally with minute ridges. Prothorax reddish brown, the vertical HOLOTYPE MALE: Lukeolela, left bank sides of the median lobe and a median of the Congo River, 010 05' S., October 13, dorsal twin spot yellow. Synthorax yellow 1930 (J. P. Chapin), in the American on the dorsum, pale brown on the sides Museum of Natural History. marked with brownish black as follows: This species is named for Dr. James P. a pair of dorsal stripes, widened below and Chapin of the American Museum. confluent on the midline, forming a narrow Oxygomphus chapini is slightly smaller triangle which becomes restricted below than agilis (agilis abdomen, 43 mm., hind to form a small inverted cone interrupting wing, 31 mm. long) and resembles it the yellow collar and joining the black be- strongly as to color pattern and type of low; on each side a black antehumeral, appendages, though the superior appen- connected above and below with the mid- dages are more widely divaricate than those dorsal triangle, leaving an oblique elongate of agilis, as figured by Fraser (1936). yellow spot on the black of the dorsum It differs from agilis most strikingly in its each side; a wide black-humeral separated simple occiput. It differs from the de- from the antehumeral by a yellow stripe scription of martininus Lacroix in size one-third again as wide as the antehumeral (martininus abdomen, 51 mm., hind wing, but not quite so wide as the humeral. 43.3 mm. long) and in color pattern. First lateral suture with a reddish spot at Oxygomphus martininus is known only the top connected with the humeral stripe from a single female, the description of and a light brownish triangular spot above which makes no mention of the occiput or the spiracle; second lateral suture with the shape of the tenth segment. In the a narrow tapering stripe interrupted on the absence of any description we can only lower third; metepimeron with a triangular conclude that there is nothing unusual or dark spot on the posterior margin. striking about them. Granting that the Femora pale; tibiae and tarsi darker. peculiarities of the occiput of agilis and 8 AMERICAN MUSEUM NOVITATES [No. 1259 of the tenth segment of both agilis and the thinking martininus different from agilis male here described as chapini may be (Lacroix, 1921, p. 49). If it was too sexual characters, I still decline to accept large for agilis (agilis abdomen, 43 mm., martininus as the female of either of these hind wing, 31 mm. long) it is certainly too males. Martin agreed with Lacroix in large for chapini.

BlBLIOGRAPHY CALVERT, PHILIP P. NEEDHAM, JAMEs G. 1903. On some American Gomphinae (Odo- 1940. Studies on neotropical Gomphinae. nata). Ent. News, vol. 14, pp. 183- Trans. Amer. Ent. Soc., vol. 65, pp. 192, pl. 8. 363-394. CAMPION, HERBERT Ris, F. 1923. Descriptions of three species of Noto- 1909. Abessinische Libellen, gesammelt von gomphus (Odonata) from East Africa. Dr. Eduard Ruipell. Ber. Sencken- Ann. Mag. Nat. Hist., ser. 9, vol. 12, bergischen Naturf. Gesellsch. Frank- pp. 659-669, figs. 1-4. furt-am-Main, vol. 40, pt. 2, Wiss. F6RSTER, F. Mitt., pp. 21-27, figs. 1-5. 1906. Forschungsreise durch Siudschoa, Galla 1921. The Odonata or of South und die Somalilinder von Carlos Frei- Africa. Ann. South African Mus., vol. herr von Erlanger. Libellen. Jahrb. 18, pp. 245-452, figs. 1-77, pls. 5-12. Nassauischer Ver. Naturk., Wies- SCROUTEDEN, HENRI baden, vol. 59, pp. 299-344, pl. A. 1933. Un nouveau "Phyllogomphus" congo- FRASER, F. C. lais (Odonata, Gomphinae). Rev. 1936. A note on the identifications of some Zool. Bot. Africaines, vol. 23, pp. 340- obscure genera and species of the 342. family (Odonata). Proc. 1934. Catalogues raisonn6s de la faune ento- Roy. Ent. Soc. London, ser. B, vol. 5, mologique du Congo Belge. Pseudo- pp. 137-143, figs. 1-2. n6vropteres, Odonates. Ann. Mus. KARSCH, FERD. Congo Belge, Zool., ser. 3, sect. 2, vol. 1890. Ueber Gomphiden. Ent. Nachr., vol. 3, fasc. 1, pp. 1-84, figs. 1-83, pl. 1. 16, pp. 370-382. SELYs-LONGCHAMPS, EDMOND DE 1899. Neue Odonaten aus Ost- und Siud- 1854. Synopsis des Gomphines. Bull. Acad. Afrika mit Einschluss des Seenge- Roy. Belgique, ser. 1, vol. 21, pt. 2, bietes. Ibid., vol. 25, pp. 369-382. pp. 23-112. KIMMINs, D. E. 1858. Monographie des Gomphines. M6m. 1931. Phyllogomphus coloratus, a new Afri- Soc. Roy. Sci. Li6ge, vol. 11, pp. 257- can . Ann. Mag. Nat. Hist., 720, pls. 1-23. ser. 10, vol. 7, pp. 217-219. 1859. Additions au synopsis des Gomphines. LACROIX, J. L. Bull. Acad. Roy. Belgique, ser. 2, vol. 1921. Notes sur quelques NMvropteres. Ann. 7, pp. 530-552. Soc. Linn. Lyons, vol. 67, pp. 45-59. 1869. Secondes additions au synopsis des LONGFIELD, CYNTHIA Gomphines. Ibid., ser. 2, vol. 28, pp. 1936. Studies on African Odonata. Trans. 168-208. Ent. Soc. London, vol. 85, pp. 467-498, 1878. Quatri6mes additions au synopsis des figs. 1-10. Gomphines. Ibid., ser. 2, vol. 46, MARTIN, RENE pp. 408-471. 1908. Voyage de feu Leonardo Fea dans l'- 1892. Causeries odonatologiques, No. 6. Les Afrique occidentale. Odonates. Ann. Gomphines d'Afrique. Ann. Soc. Ent. Mus. Civico Storia Nat., Genova, vol. Belgique, vol. 36, pp. 86-106. 43 (ser. 3, vol. 3), for 1907, pp. 649- SJ6sTEDT, YNGVE 667. 1909. Wissenschaftliche Ergebnisse der 1915. R6sultats scientifiques, Insectes, Schwedischen Zoologischen Expedition Pseudon6vropteres, II, Odonata. Voy- nach dem Kilimandiaro, dem Meru. age de Ch. Alluaud et R. Jeannel en Vol. 2, div. 14, Pseudoneuroptera, pt. Afrique orientale (1911-1912), pp. 19- 1, Odonata, pp. 1-52, pls. 1-2. 50, pls. 1-3. WILLIAMSON, E. B. NAVAS, LONGINOs 1920. A new gomphine genus from British 1936. N6vropt6res et insectes voisins Chine Guiana with a note on the classifica- et pays environnants. 9th ser. Notes tion of the subfamily (order Odonata). d'Ent. Chinoise, vol. 3, pp. 37-62, 117- Occas. Papers Mus. Zool., Univ. 132, figs. 66-90. Michigan, no. 80, pp. 1-12, pl. 1. 19441 NOTES ON THE GOMPHINAE (ODONATA) 9

5

-- '11 I 1 3,

Ud e 4

Fig. 1. Oxygomphus chapini, new species, terminal appendages, lateral aspect. Fig. 2. Idem, terminal appendages, dorsal aspect. Fig. 3. Idem, genitalia of the second abdominal segment, lateral aspect. Fig. 4. Idem, diagram of thoracic color pattern. Fig. 5. Diaphlebia species, vulvar scale, ventral aspect. Fig. 6. Idem, vulvar scale, lateral aspect. Fig. 7. Phyllogomphus coloratus Kimmins, vulvar scale and tenth segment, lateral aspect. Fig. 8. Phyllogomphu8 annulus, new species, vulvar scale and tenth segment, lateral aspect. Fig. 9. Phyllogomphus coloratus Ki4nmins, vulvar scale, ventral aspect. Fig. 10. Phyllogomphus annulus, new species, vulvar scale, ventral aspect. 10 AMERICAN MUSEUM NOVITATES [No. 1259

13

14

15

12

Fig. 11. Peruviogomphus moyobambus, new species, genitalia of second segment, ventral aspect. Fig. 12. Idem, genitalia of second segment, lateral aspect. Fig. 13. Idem, terminal appendages, lateral aspect. Fig. 14. Idem, terminal appendages, dorsal aspect. Fig. 15. Idem, face. CF

0

CI2

G0 *0u 00>

*0 c)