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Sereno 20060098.Vp
Basal abelisaurid and carcharodontosaurid theropods from the Lower Cretaceous Elrhaz Formation of Niger PAUL C. SERENO and STEPHEN L. BRUSATTE Sereno, P.C. and Brusatte, S.L. 2008. Basal abelisaurid and carcharodontosaurid theropods from the Lower Cretaceous Elrhaz Formation of Niger. Acta Palaeontologica Polonica 53 (1): 15–46. We report the discovery of basal abelisaurid and carcharodontosaurid theropods from the mid Cretaceous (Aptian– Albian, ca. 112 Ma) Elrhaz Formation of the Niger Republic. The abelisaurid, Kryptops palaios gen. et sp. nov., is repre− sented by a single individual preserving the maxilla, pelvic girdle, vertebrae and ribs. Several features, including a maxilla textured externally by impressed vascular grooves and a narrow antorbital fossa, clearly place Kryptops palaios within Abelisauridae as its oldest known member. The carcharodontosaurid, Eocarcharia dinops gen. et sp. nov., is repre− sented by several cranial bones and isolated teeth. Phylogenetic analysis places it as a basal carcharodontosaurid, similar to Acrocanthosaurus and less derived than Carcharodontosaurus and Giganotosaurus. The discovery of these taxa sug− gests that large body size and many of the derived cranial features of abelisaurids and carcharodontosaurids had already evolved by the mid Cretaceous. The presence of a close relative of the North American genus Acrocanthosaurus on Af− rica suggests that carcharodontosaurids had already achieved a trans−Tethyan distribution by the mid Cretaceous. Key words: Theropod, abelisaurid, allosauroid, carcharodontosaurid, Kryptops, Eocarcharia, Cretaceous, Africa. Paul C. Sereno [[email protected]], Department of Organismal Biology and Anatomy, University of Chicago, 1027 E. 57th Street, Chicago, Illinois, 60637, USA; Stephen L. Brusatte [[email protected]], Department of Earth Sciences, University of Bristol, Wills Memorial Building, Queen’s Road, Bristol BS8 1RJ, United Kingdom. -
Dr. N. G. Kotadiya Dinosaurs
Dinosaurs The word dinosaur was given by SIR RICHARD OWEN and it means 'terrible lizards'. From the middle of Triassic to the end of cretaceous the dinosaurs dominated the earth. At the end of the cretaceous period the dinosaurs became extinct. The dinosaurs include two orders, namely Saurischia (Reptile-like dinosaurs) and Ornithischia (Bird-like dinosaurs) Mesozoic Reptiles Thecodonts Dinosaurs pterosaurs Marine Therapsids (Terrible lizards) (Flying reptiles) reptiles (Mammal-like reptiles) Representative Types The Mesozoic reptiles include the following types: Dinosaurs: 1. Ornithomimus 2. Tyrannosaurus 3. Ornitholestes 4. Coelophysis 5. Allosaurus 6. Brontosaurus 7. Diplodocus 8. Camptosaurus 9. Trachodonts 10. Iguanodon 11. Stegosaurus 12. Ankylosaurus 13. Monoclonius 14. Triceratops Pterosaurs: 15. Rhamphorhynchus 16. Pteranodon Marine reptiles: 17. Ichthyosaurus (fish reptiles) 18. Pliosaurus (Swan dragons) Mammal-like reptiles : 19. Varanosaurus 20. Edaphosaurus 21. Dimetrodon 22. Cynognathus 23. Tritylodon 24. Diarthrognathus Dinosaurs Tyrannosaurus (Tyrant dinosaur) Dinosaurs 1. Tyrannosaurus (Tyrant dinosaur) Tyrannosaurus was a Saurischian dinosaur about 6.09 metre tall and 15.24 meter long. It was the largest bipedal terrestrial dinosaur of the cretaceous period. The hind limbs were very strong but fore limbs and hands greatly reduced. The jaws were long with large dagger- like teeth. It was a carnivore well adapted for hunting and killing other large reptiles including other dinosaur. Dinosaurs Brontosaurus Dinosaurs 2. Brontosaurus Brontosaurus was a giant sauropod Saurischiar of the jurassic times. It was about 22.86 m long and weighed about 30-60 tons. It was a quadrupedal dinosaur with strong fore and hind limb. But the fore limbs were slightly shorter. The feet were very broad with large foot pads. -
Estimating Mass Properties of Dinosaurs Using Laser Imaging and 3D Computer Modelling
Estimating Mass Properties of Dinosaurs Using Laser Imaging and 3D Computer Modelling Karl T. Bates1*, Phillip L. Manning2,3, David Hodgetts3, William I. Sellers1 1 Adaptive Organismal Biology Research Group, Faculty of Life Sciences, University of Manchester, Jackson’s Mill, Manchester, United Kingdom, 2 The Manchester Museum, University of Manchester, Manchester, United Kingdom, 3 School of Earth, Atmospheric and Environmental Science, University of Manchester, Manchester, United Kingdom Abstract Body mass reconstructions of extinct vertebrates are most robust when complete to near-complete skeletons allow the reconstruction of either physical or digital models. Digital models are most efficient in terms of time and cost, and provide the facility to infinitely modify model properties non-destructively, such that sensitivity analyses can be conducted to quantify the effect of the many unknown parameters involved in reconstructions of extinct animals. In this study we use laser scanning (LiDAR) and computer modelling methods to create a range of 3D mass models of five specimens of non- avian dinosaur; two near-complete specimens of Tyrannosaurus rex, the most complete specimens of Acrocanthosaurus atokensis and Strutiomimum sedens, and a near-complete skeleton of a sub-adult Edmontosaurus annectens. LiDAR scanning allows a full mounted skeleton to be imaged resulting in a detailed 3D model in which each bone retains its spatial position and articulation. This provides a high resolution skeletal framework around which the body cavity and internal organs such as lungs and air sacs can be reconstructed. This has allowed calculation of body segment masses, centres of mass and moments or inertia for each animal. However, any soft tissue reconstruction of an extinct taxon inevitably represents a best estimate model with an unknown level of accuracy. -
'Big Al'? Quantifying the Effect of Soft
Palaeontologia Electronica http://palaeo-electronica.org HOW BIG WAS ‘BIG AL’? QUANTIFYING THE EFFECT OF SOFT TISSUE AND OSTEOLOGICAL UNKNOWNS ON MASS PREDICTIONS FOR ALLOSAURUS (DINOSAURIA:THEROPODA) Karl T. Bates, Peter L. Falkingham, Brent H. Breithaupt, David Hodgetts, William I. Sellers, and Phillip L. Manning ABSTRACT MOR693, nicknamed ‘Big Al,’ is the most complete skeleton of the non-avian theropod Allosaurus and therefore provides the best opportunity to investigate the mass properties of this important Jurassic theropod through accurate physical or digital volumetric models. In this study, laser scanning and computer modelling software have been used to construct volumetric models of MOR693. A long-range laser scanner has been used to digitize the mounted cast of MOR693, allowing the reconstruction of body volumes and respiratory structures around and within the three-dimensional (3D) skel- etal model. The digital medium offered the facility to modify model properties non- destructively in a detailed sensitivity analysis to quantify the effects of the many unknown parameters involved in such reconstructions. In addition to varying the vol- umes of body segments and respiratory structures, we also extend the sensitivity anal- ysis to include uncertainties regarding osteological articulations in non-avian dinosaurs, including effects of inter-vertebral spacing and the orientation or ‘flare’ of the rib cage in MOR693. Results suggest body mass and inertial values are extremely uncertain and show a wide range in plausible values, whilst the CM (centre of mass) position is well constrained immediately in front and below the hip joint in MOR693, consistent with similar reconstructions of non-avian theropods. Karl T. -
The Fighting Pair
THE FIGHTING PAIR ALLOSAURUS VS STEGOSAURUS — Allosaurus “jimmadsoni” and Hesperosaurus (Stegosaurus) mjosi Upper Jurassic Period, Kimmeridgian Stage, 155 million years old Morrison Formation Dana Quarry, Ten Sleep, Washakie County, Wyoming, USA. THE ALLOSAURUS The Official State Fossil of Utah, the Allosaurus was a large theropod carnosaur of the “bird-hipped” Saurischia order that flourished primarily in North America during the Upper Jurassic Period, 155-145 million years In the spring of 2007, at the newly-investigated Dana Quarry in the Morrison Formation of Wyoming, the team from ago. Long recognized in popular culture, it bears the distinction of being Dinosauria International LLC made an exciting discovery: the beautifully preserved femur of the giant carnivorous one of the first dinosaurs to be depicted on the silver screen, the apex Allosaur. As they kept digging, their excitement grew greater; next came toe bones, leg bones, ribs, vertebrae and predator of the 1912 novel and 1925 cinema adaptation of Conan Doyle’s finally a skull: complete, undistorted and, remarkably, with full dentition. It was an incredible find; one of the most The Lost World. classic dinosaurs, virtually complete, articulated and in beautiful condition. But that was not all. When the team got The Allosaurus possessed a large head on a short neck, a broad rib-cage the field jackets back to the preparation lab, they discovered another leg bone beneath the Allosaurus skull… There creating a barrel chest, small three-fingered forelimbs, large powerful hind limbs with hoof-like feet, and a long heavy tail to act as a counter-balance. was another dinosaur in the 150 million year-old rock. -
A Census of Dinosaur Fossils Recovered from the Hell Creek and Lance Formations (Maastrichtian)
The Journal of Paleontological Sciences: JPS.C.2019.01 1 TAKING COUNT: A Census of Dinosaur Fossils Recovered From the Hell Creek and Lance Formations (Maastrichtian). ______________________________________________________________________________________ Walter W. Stein- President, PaleoAdventures 1432 Mill St.. Belle Fourche, SD 57717. [email protected] 605-210-1275 ABSTRACT: A census of Hell Creek and Lance Formation dinosaur remains was conducted from April, 2017 through February of 2018. Online databases were reviewed and curators and collections managers interviewed in an effort to determine how much material had been collected over the past 130+ years of exploration. The results of this new census has led to numerous observations regarding the quantity, quality, and locations of the total collection, as well as ancillary data on the faunal diversity and density of Late Cretaceous dinosaur populations. By reviewing the available data, it was also possible to make general observations regarding the current state of certain exploration programs, the nature of collection bias present in those collections and the availability of today's online databases. A total of 653 distinct, associated and/or articulated remains (skulls and partial skeletons) were located. Ceratopsid skulls and partial skeletons (mostly identified as Triceratops) were the most numerous, tallying over 335+ specimens. Hadrosaurids (Edmontosaurus) were second with at least 149 associated and/or articulated remains. Tyrannosaurids (Tyrannosaurus and Nanotyrannus) were third with a total of 71 associated and/or articulated specimens currently known to exist. Basal ornithopods (Thescelosaurus) were also well represented by at least 42 known associated and/or articulated remains. The remaining associated and/or articulated specimens, included pachycephalosaurids (18), ankylosaurids (6) nodosaurids (6), ornithomimids (13), oviraptorosaurids (9), dromaeosaurids (1) and troodontids (1). -
Limb Design, Function and Running Performance in Ostrich-Mimics and Tyrannosaurs
GAIA Nº 15, LISBOA/LISBON, DEZEMBRO/DECEMBER 1998, pp. 257-270 (ISSN: 0871-5424) LIMB DESIGN, FUNCTION AND RUNNING PERFORMANCE IN OSTRICH-MIMICS AND TYRANNOSAURS Gregory S. PAUL 3109 N Calvert St. Side Apt., BALTIMORE MD 21218. USA ABSTRACT: Examination of the limb morphology of small ornithomimids and large tyranno- saurids shows that they remained remarkably constant in design regardless of size. The changes that were present were consistent with maintaining limb strength and function constant with size. It is concluded that ornithomimid and tyrannosaurid legs functioned in a similar manner, and always exhibit the features normally associated with a fast running gait. This is in contrast to modern animals, in which elephants as gigantic as large tyranno- saurids have limbs that are modified for a slow walking gait. INTRODUCTION to run observed in elephants. The hypothesis can be challenged if it can be shown that at least some ex- Because they had long, gracile, bird-like legs, it tinct giants retained the skeletal adaptations for run- has long been accepted that the smaller predatory ning observed in smaller species, and in their own dinosaurs were swift runners (O [& GREG- SBORN offspring. In turn, the hypothesis that giants can run ], 1916; COLBERT, 1961; RUSSELL, 1972; ORY fast if they retain limbs similar to smaller runners can C , 1978; THULBORN, 1982; PAUL, 1988; OOMBS be challenged if it is shown that the skeleton is too H , 1994). Much more controversial has been OLTZ vulnerable to structural failure. the locomotory abilities of their giant relatives, which have been restored as no faster than elephants BODY MASSES (LAMBE, 1917; HALSTEAD &HALSTEAD, 1981; THUL- BORN, 1982; BARSBOLD, 1983), able to run at only Sources for mass data for extinct and extant ani- modest speeds (COOMBS, 1978; MOLNAR &FAR- mals include PAUL (1988, 1997), NOWAK (1991) and LOW, 1990; HORNER &LESSEM, 1993; FARLOW, MATTHEWS (1994). -
Cranial Anatomy of Allosaurus Jimmadseni, a New Species from the Lower Part of the Morrison Formation (Upper Jurassic) of Western North America
Cranial anatomy of Allosaurus jimmadseni, a new species from the lower part of the Morrison Formation (Upper Jurassic) of Western North America Daniel J. Chure1,2,* and Mark A. Loewen3,4,* 1 Dinosaur National Monument (retired), Jensen, UT, USA 2 Independent Researcher, Jensen, UT, USA 3 Natural History Museum of Utah, University of Utah, Salt Lake City, UT, USA 4 Department of Geology and Geophysics, University of Utah, Salt Lake City, UT, USA * These authors contributed equally to this work. ABSTRACT Allosaurus is one of the best known theropod dinosaurs from the Jurassic and a crucial taxon in phylogenetic analyses. On the basis of an in-depth, firsthand study of the bulk of Allosaurus specimens housed in North American institutions, we describe here a new theropod dinosaur from the Upper Jurassic Morrison Formation of Western North America, Allosaurus jimmadseni sp. nov., based upon a remarkably complete articulated skeleton and skull and a second specimen with an articulated skull and associated skeleton. The present study also assigns several other specimens to this new species, Allosaurus jimmadseni, which is characterized by a number of autapomorphies present on the dermal skull roof and additional characters present in the postcrania. In particular, whereas the ventral margin of the jugal of Allosaurus fragilis has pronounced sigmoidal convexity, the ventral margin is virtually straight in Allosaurus jimmadseni. The paired nasals of Allosaurus jimmadseni possess bilateral, blade-like crests along the lateral margin, forming a pronounced nasolacrimal crest that is absent in Allosaurus fragilis. Submitted 20 July 2018 Accepted 31 August 2019 Subjects Paleontology, Taxonomy Published 24 January 2020 Keywords Allosaurus, Allosaurus jimmadseni, Dinosaur, Theropod, Morrison Formation, Jurassic, Corresponding author Cranial anatomy Mark A. -
Dinosaurs, by William Diller Matthew
The Project Gutenberg eBook of Dinosaurs, by William Diller Matthew http://www.gutenberg.org/files/19302/19302-h/19302-h.htm The[10][11][12][13][14][15][16][17][18][19][20][21][22][1][2][3][4][5][6][7][8][9] Project Gutenberg eBook, Dinosaurs, by T[100][103][104][105][106][108][109][110][111][112][113][115][117][118][119][120][121][123][124][125][126][127][128][129][130][131][132][133][134][135][136][137][138][139][140][141][142][143][144][145][146][147][148][149][150][151][152][153][154][155][156][157][158][159][161][162][10][11][12][13][14][15][17][18][19][20][21][22][23][24][26][27][28][29][30][31][34][35][36][38][39][40][41][42][43][44][45][46][47][48][49][50][51][52][53][54][55][56][57][58][61][63][64][65][66][67][68][69][70][71][72][73][74][76][77][78][79][80][81][83][84][85][86][87][88][90][91][93][94][95][96][97][98][99]OC William Diller Matthew This eBook is for the use of anyone anywhere at no cost and with almost no restrictions whatsoever. You may copy it, give it away or re-use it under the terms of the Project Gutenberg License included with this eBook or online at www.gutenberg.org Title: Dinosaurs With Special Reference to the American Museum Collections Author: William Diller Matthew Release Date: September 16, 2006 [eBook #19302] Language: English Character set encoding: ISO-8859-1 ***START OF THE PROJECT GUTENBERG EBOOK DINOSAURS*** E-text prepared by Brian Janes, Suzanne Lybarger, Jeannie Howse, and the Project Gutenberg Online Distributed Proofreading Team http://www.pgdp.net) Transcriber's Note: Click the image to see a larger version. -
New Tyrannosaur from the Mid-Cretaceous of Uzbekistan Clarifies Evolution of Giant Body Sizes and Advanced Senses in Tyrant Dinosaurs
New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of giant body sizes and advanced senses in tyrant dinosaurs Stephen L. Brusattea,1, Alexander Averianovb,c, Hans-Dieter Suesd, Amy Muira, and Ian B. Butlera aSchool of GeoSciences, University of Edinburgh, Edinburgh EH9 3FE, United Kingdom; bZoological Institute, Russian Academy of Sciences, St. Petersburg 199034, Russia; cDepartment of Sedimentary Geology, Saint Petersburg State University, St. Petersburg 199178, Russia; and dDepartment of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560 Edited by Neil H. Shubin, The University of Chicago, Chicago, IL, and approved January 29, 2016 (received for review January 5, 2016) Tyrannosaurids—the familiar group of carnivorous dinosaurs in- We here report the first diagnostic tyrannosauroid from the mid- cluding Tyrannosaurus and Albertosaurus—were the apex predators Cretaceous, a new species from the Turonian (ca. 90–92 million in continental ecosystems in Asia and North America during the years ago) Bissekty Formation of Uzbekistan. This formation has latest Cretaceous (ca. 80–66 million years ago). Their colossal sizes recently emerged as one of the most important records of mid- and keen senses are considered key to their evolutionary and eco- Cretaceous dinosaurs globally (9–11). Possible tyrannosauroid logical success, but little is known about how these features devel- specimens from the Bissekty Formation were reported more than oped as tyrannosaurids evolved from smaller basal tyrannosauroids a half century ago (12), and, more recently, several isolated fossils that first appeared in the fossil record in the Middle Jurassic (ca. 170 were assigned to the group (9, 13), but none of these has been million years ago). -
Body-Size Evolution in the Dinosauria
8 Body-Size Evolution in the Dinosauria Matthew T. Carrano Introduction The evolution of body size and its influence on organismal biology have received scientific attention since the earliest decades of evolutionary study (e.g., Cope, 1887, 1896; Thompson, 1917). Both paleontologists and neontologists have attempted to determine correlations between body size and numerous aspects of life history, with the ultimate goal of docu- menting both the predictive and causal connections involved (LaBarbera, 1986, 1989). These studies have generated an appreciation for the thor- oughgoing interrelationships between body size and nearly every sig- nificant facet of organismal biology, including metabolism (Lindstedt & Calder, 1981; Schmidt-Nielsen, 1984; McNab, 1989), population ecology (Damuth, 1981; Juanes, 1986; Gittleman & Purvis, 1998), locomotion (Mc- Mahon, 1975; Biewener, 1989; Alexander, 1996), and reproduction (Alex- ander, 1996). An enduring focus of these studies has been Cope’s Rule, the notion that body size tends to increase over time within lineages (Kurtén, 1953; Stanley, 1973; Polly, 1998). Such an observation has been made regarding many different clades but has been examined specifically in only a few (MacFadden, 1986; Arnold et al., 1995; Jablonski, 1996, 1997; Trammer & Kaim, 1997, 1999; Alroy, 1998). The discordant results of such analyses have underscored two points: (1) Cope’s Rule does not apply universally to all groups; and (2) even when present, size increases in different clades may reflect very different underlying processes. Thus, the question, “does Cope’s Rule exist?” is better parsed into two questions: “to which groups does Cope’s Rule apply?” and “what process is responsible for it in each?” Several recent works (McShea, 1994, 2000; Jablonski, 1997; Alroy, 1998, 2000a, 2000b) have begun to address these more specific questions, attempting to quantify patterns of body-size evolution in a phylogenetic (rather than strictly temporal) context, as well as developing methods for interpreting the resultant patterns. -
Tyrannosaurs in Science
Media Inquiries: Kristin Elise Phillips, Department of Communications September 2010 212-496-3419; [email protected] www.amnh.org ________________________________________________________________________ SCIENTIFIC UNDERSTANDING OF T. REX REVISED BY A DECADE OF NEW RESEARCH AND DISCOVERY TYRANNOSAURS ARE MORE THAN LARGE CARNIVORES AT THE TOP OF THE FOOD CHAIN We’ve all heard this story: the Late Cretaceous of Asia and North America— about 65 million years ago—was dominated by several large-headed, bipedal predatory dinosaurs like Tyrannosaurus rex and Tarbosaurus that had tiny arms. But a decade of new fossil discoveries that have more than doubled the number of known tyrannosaur species has changed this tale. Older and smaller tyrannosaurs have made the evolutionary tree of this group richer and more complex. Furthermore, a series of innovative research projects on topics like bone growth and biomechanics have added an enormous amount of information about tyrannosaurs, so much so that the group could now be considered an exemplar for studying many themes in paleontology research. A new paper describing recent research and a new evolutionary tree is published in Science this week. “T. rex is the most iconic of all dinosaurs,” says Mark Norell, curator in the Division of Paleontology at the American Museum of Natural History. “Its star power has allowed a research focus into questions not normally undertaken with fossils, questions like bone growth, biomechanics, and neurology.” “We know more about tyrannosaurs than any other group of dinosaurs—even more than some groups of living organisms,” says Stephen Brusatte, a graduate student affiliated with the Museum and first author of the paper.