Dr. N. G. Kotadiya Dinosaurs
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The Fighting Pair
THE FIGHTING PAIR ALLOSAURUS VS STEGOSAURUS — Allosaurus “jimmadsoni” and Hesperosaurus (Stegosaurus) mjosi Upper Jurassic Period, Kimmeridgian Stage, 155 million years old Morrison Formation Dana Quarry, Ten Sleep, Washakie County, Wyoming, USA. THE ALLOSAURUS The Official State Fossil of Utah, the Allosaurus was a large theropod carnosaur of the “bird-hipped” Saurischia order that flourished primarily in North America during the Upper Jurassic Period, 155-145 million years In the spring of 2007, at the newly-investigated Dana Quarry in the Morrison Formation of Wyoming, the team from ago. Long recognized in popular culture, it bears the distinction of being Dinosauria International LLC made an exciting discovery: the beautifully preserved femur of the giant carnivorous one of the first dinosaurs to be depicted on the silver screen, the apex Allosaur. As they kept digging, their excitement grew greater; next came toe bones, leg bones, ribs, vertebrae and predator of the 1912 novel and 1925 cinema adaptation of Conan Doyle’s finally a skull: complete, undistorted and, remarkably, with full dentition. It was an incredible find; one of the most The Lost World. classic dinosaurs, virtually complete, articulated and in beautiful condition. But that was not all. When the team got The Allosaurus possessed a large head on a short neck, a broad rib-cage the field jackets back to the preparation lab, they discovered another leg bone beneath the Allosaurus skull… There creating a barrel chest, small three-fingered forelimbs, large powerful hind limbs with hoof-like feet, and a long heavy tail to act as a counter-balance. was another dinosaur in the 150 million year-old rock. -
Cranial Anatomy of Allosaurus Jimmadseni, a New Species from the Lower Part of the Morrison Formation (Upper Jurassic) of Western North America
Cranial anatomy of Allosaurus jimmadseni, a new species from the lower part of the Morrison Formation (Upper Jurassic) of Western North America Daniel J. Chure1,2,* and Mark A. Loewen3,4,* 1 Dinosaur National Monument (retired), Jensen, UT, USA 2 Independent Researcher, Jensen, UT, USA 3 Natural History Museum of Utah, University of Utah, Salt Lake City, UT, USA 4 Department of Geology and Geophysics, University of Utah, Salt Lake City, UT, USA * These authors contributed equally to this work. ABSTRACT Allosaurus is one of the best known theropod dinosaurs from the Jurassic and a crucial taxon in phylogenetic analyses. On the basis of an in-depth, firsthand study of the bulk of Allosaurus specimens housed in North American institutions, we describe here a new theropod dinosaur from the Upper Jurassic Morrison Formation of Western North America, Allosaurus jimmadseni sp. nov., based upon a remarkably complete articulated skeleton and skull and a second specimen with an articulated skull and associated skeleton. The present study also assigns several other specimens to this new species, Allosaurus jimmadseni, which is characterized by a number of autapomorphies present on the dermal skull roof and additional characters present in the postcrania. In particular, whereas the ventral margin of the jugal of Allosaurus fragilis has pronounced sigmoidal convexity, the ventral margin is virtually straight in Allosaurus jimmadseni. The paired nasals of Allosaurus jimmadseni possess bilateral, blade-like crests along the lateral margin, forming a pronounced nasolacrimal crest that is absent in Allosaurus fragilis. Submitted 20 July 2018 Accepted 31 August 2019 Subjects Paleontology, Taxonomy Published 24 January 2020 Keywords Allosaurus, Allosaurus jimmadseni, Dinosaur, Theropod, Morrison Formation, Jurassic, Corresponding author Cranial anatomy Mark A. -
Dinosaurs, by William Diller Matthew
The Project Gutenberg eBook of Dinosaurs, by William Diller Matthew http://www.gutenberg.org/files/19302/19302-h/19302-h.htm The[10][11][12][13][14][15][16][17][18][19][20][21][22][1][2][3][4][5][6][7][8][9] Project Gutenberg eBook, Dinosaurs, by T[100][103][104][105][106][108][109][110][111][112][113][115][117][118][119][120][121][123][124][125][126][127][128][129][130][131][132][133][134][135][136][137][138][139][140][141][142][143][144][145][146][147][148][149][150][151][152][153][154][155][156][157][158][159][161][162][10][11][12][13][14][15][17][18][19][20][21][22][23][24][26][27][28][29][30][31][34][35][36][38][39][40][41][42][43][44][45][46][47][48][49][50][51][52][53][54][55][56][57][58][61][63][64][65][66][67][68][69][70][71][72][73][74][76][77][78][79][80][81][83][84][85][86][87][88][90][91][93][94][95][96][97][98][99]OC William Diller Matthew This eBook is for the use of anyone anywhere at no cost and with almost no restrictions whatsoever. You may copy it, give it away or re-use it under the terms of the Project Gutenberg License included with this eBook or online at www.gutenberg.org Title: Dinosaurs With Special Reference to the American Museum Collections Author: William Diller Matthew Release Date: September 16, 2006 [eBook #19302] Language: English Character set encoding: ISO-8859-1 ***START OF THE PROJECT GUTENBERG EBOOK DINOSAURS*** E-text prepared by Brian Janes, Suzanne Lybarger, Jeannie Howse, and the Project Gutenberg Online Distributed Proofreading Team http://www.pgdp.net) Transcriber's Note: Click the image to see a larger version. -
Analysis of Hindlimb Muscle Moment Arms in Tyrannosaurus Rex Using a Three-Dimensional Musculoskeletal Computer Model: Implications for Stance, Gait, and Speed
Paleobiology, 31(4), 2005, pp. 676±701 Analysis of hindlimb muscle moment arms in Tyrannosaurus rex using a three-dimensional musculoskeletal computer model: implications for stance, gait, and speed John R. Hutchinson, Frank C. Anderson, Silvia S. Blemker, and Scott L. Delp Abstract.ÐMuscle moment arms are important determinants of muscle function; however, it is chal- lenging to determine moment arms by inspecting bone specimens alone, as muscles have curvilin- ear paths that change as joints rotate. The goals of this study were to (1) develop a three-dimen- sional graphics-based model of the musculoskeletal system of the Cretaceous theropod dinosaur Tyrannosaurus rex that predicts muscle-tendon unit paths, lengths, and moment arms for a range of limb positions; (2) use the model to determine how the T. rex hindlimb muscle moment arms varied between crouched and upright poses; (3) compare the predicted moment arms with previous assessments of muscle function in dinosaurs; (4) evaluate how the magnitudes of these moment arms compare with those in other animals; and (5) integrate these ®ndings with previous biome- chanical studies to produce a revised appraisal of stance, gait, and speed in T. rex. The musculo- skeletal model includes ten degrees of joint freedom (¯exion/extension, ab/adduction, or medial/ lateral rotation) and 33 main muscle groups crossing the hip, knee, ankle, and toe joints of each hindlimb. The model was developed by acquiring and processing bone geometric data, de®ning joint rotation axes, justifying muscle attachment sites, and specifying muscle-tendon geometry and paths. Flexor and extensor muscle moment arms about all of the main limb joints were estimated, and limb orientation was statically varied to characterize how the muscle moment arms changed. -
An Evaluation of Flapping-Based Locomotory Hypotheses in Bird
The wings before the bird: an evaluation of flapping-based locomotory hypotheses in bird antecedents T. Alexander Dececchi1, Hans C.E. Larsson2 and Michael B. Habib3,4 1 Department of Geological Sciences, Queens University, Kingston, Ontario, Canada 2 Redpath Museum, McGill University, Montreal, Quebec, Canada 3 Keck School of Medicine of USC, Department of Cell and Neurobiology, University of Southern California, Los Angeles, California, United States 4 Dinosaur Institute, Natural History Museum of Los Angeles, Los Angeles, CA, United States ABSTRACT Background: Powered flight is implicated as a major driver for the success of birds. Here we examine the effectiveness of three hypothesized pathways for the evolution of the flight stroke, the forelimb motion that powers aerial locomotion, in a terrestrial setting across a range of stem and basal avians: flap running, Wing Assisted Incline Running (WAIR), and wing-assisted leaping. Methods: Using biomechanical mathematical models based on known aerodynamic principals and in vivo experiments and ground truthed using extant avians we seek to test if an incipient flight stroke may have contributed sufficient force to permit flap running, WAIR, or leaping takeoff along the phylogenetic lineage from Coelurosauria to birds. Results: None of these behaviours were found to meet the biomechanical threshold requirements before Paraves. Neither was there a continuous trend of refinement for any of these biomechanical performances across phylogeny nor a signal of universal applicability near the origin of birds. None of these flap-based locomotory models appear to have been a major influence on pre-flight character acquisition such as pennaceous feathers, suggesting non-locomotory behaviours, and less Submitted 23 January 2016 stringent locomotory behaviours such as balancing and braking, played a role in Accepted 27 May 2016 the evolution of the maniraptoran wing and nascent flight stroke. -
EVIDENCE of DINOSAURS in NEBRASKA Erwin Hinckley Barbour
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Bulletin of the University of Nebraska State Museum, University of Nebraska State Museum 1931 EVIDENCE OF DINOSAURS IN NEBRASKA Erwin Hinckley Barbour Follow this and additional works at: http://digitalcommons.unl.edu/museumbulletin Part of the Entomology Commons, Geology Commons, Geomorphology Commons, Other Ecology and Evolutionary Biology Commons, Paleobiology Commons, Paleontology Commons, and the Sedimentology Commons This Article is brought to you for free and open access by the Museum, University of Nebraska State at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Bulletin of the University of Nebraska State Museum by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. I) • ,~ I BULLETIN 21 VOLUME I _JULY, 1931 · f" -.' . UI ~ ; ; , . I, THE NEBRASKA STATE MU EUM ERWIN H. BARBOUR, Director t, .. • I"'! EVIDENCE OF DINOSAURS IN NEBRASKA 1 I' , i I' By ERWIN HINCKLEY BARBOUIt _____ , .. __________ ,,,.__ . Nebraska has long been a collecting ground famous for its fossil mammals, but as yet no dinosaurian bones have been reported, nor have they been expected. The distal end of a finely preserved femur, however, has recently been brought to light, supposedly occurring in position in the Dakota for mation of eastern Nebraska. It was discovered, collected, and donated by Mr. J. B. White, (University of Nebraska, Law, class of 1899) on his farm two miles south of Decatur, in northeastern Burt County, near the Missouri River. It was found in undoubted Dakota sand associated with many leaf impressions. This is taken as strong evidence, rather than proof positive, that the bone was actually in position. -
New Data on Small Theropod Dinosaurs from the Upper Jurassic Morrison Formation of Como Bluff, Wyoming, USA
Volumina Jurassica, 2014, Xii (2): 181–196 Doi: 10.5604/17313708 .1130142 New data on small theropod dinosaurs from the Upper Jurassic Morrison Formation of Como Bluff, Wyoming, USA Sebastian G. DALMAN1 Key words: dinosaurs, Theropoda, Upper Jurassic, Morrison Formation, Como Bluff, Wyoming, western USA. Abstract. In 1879, Othniel C. Marsh and Arthur Lakes collected in the Upper Jurassic Morrison Formation Quarry 12 at Como Bluff, Wyoming, USA, several isolated axial and appendicular skeletal elements of small theropod dinosaurs. Since the discovery the specimens remained unnoticed for over a century. The skeletal remains of small theropods are rare at Como Bluff and throughout the Morrison Forma- tion. Their bones are delicately constructed, so they are not as well-preserved as the bones of large-bodied theropods. The bones of small theropods described here were found mixed with isolated crocodile teeth and turtle shells. Comparison of the skeletal materials with other known theropods from the Morrison Formation reveals that some of the bones belong to a very small juvenile Allosaurus fragilis and Tor vosaurus tanneri and also to a new ceratosaur taxon, here named Fosterovenator churei, whereas the other bones represent previously unidentified juvenile taxa of basal tetanuran and coelurid theropods. The discovery and description of these fossil materials is significant because they provide important information about the Upper Jurassic terrestrial fauna of Quarry 12, Como Bluff, Wyoming. The presence of previously unidentified theropod taxa in the Morrison Formation indicates that the diversity of basal tetanuran and coelurid theropods may have been much greater than previously expected. Although the fossil material here described is largely fragmentary, it is tenable that theropods of different clades co-existed in the same ecosystems at the same time and most likely competed for the same food sources. -
A New Specimen of Acrocanthosaurus Atokensis (Theropoda, Dinosauria) from the Lower Cretaceous Antlers Formation (Lower Cretaceous, Aptian) of Oklahoma, USA
A new specimen of Acrocanthosaurus atokensis (Theropoda, Dinosauria) from the Lower Cretaceous Antlers Formation (Lower Cretaceous, Aptian) of Oklahoma, USA Philip J. CURRIE Royal Tyrrell Museum of Palaeontology, Box 7500, Drumheller, Alberta T0J 0Y0 (Canada) [email protected] Kenneth CARPENTER Denver Museum of Natural History, Department of Earth Sciences, City Park, Denver, Colorado 80205 (USA) [email protected] Currie P. J. & Carpenter K. 2000. — A new specimen of Acrocanthosaurus atokensis (Theropoda, Dinosauria) from the Lower Cretaceous Antlers Formation (Lower Cretaceous, Aptian) of Oklahoma, USA. Geodiversitas 22 (2) : 207-246. The data matrix is available at http://www.mnhn.fr/publication/matadd/g00n2a3.html ABSTRACT A new skeleton of Acrocanthosaurus atokensis is the most complete specimen collected and has the only known complete skull. Aspects of the new skeleton are described in detail, with special attention directed to the morphology of the skull and forelimb. Although unquestionably one of the largest theropods ever found, it is smaller than Carcharodontosaurus, Giganotosaurus and Tyrannosaurus. Comparison with other theropods suggests that Acrocanthosaurus bears a strong resemblance to these taxa because of charac- KEY WORDS ters that are size determinate, and the evidence suggests Acrocanthosaurus is Dinosaurs, more closely related to Allosauridae than to Carcharodontosauridae. Three theropods, Early Cretaceous, families (Allosauridae, Carcharodontosauridae, Sinraptoridae) are recognized USA. in the Allosauroidea. GEODIVERSITAS • 2000 • 22 (2) © Publications Scientifiques du Muséum national d’Histoire naturelle, Paris. www.mnhn.fr/publication/ 207 Currie P. J. & Carpenter K. RÉSUMÉ Un nouveau specimen d’Acrocanthosaurus atokensis (Theropoda, Dinosauria) du Crétacé inférieur de la Formation Antlers (Crétacé inférieur, Aptien) de l’Oklahoma, États-Unis. -
Tyrannosaurids (Dinosauria) of Asia and North America
Aspects of Nonmarine Cretaceous Geology Tyrannosaurids (Dinosauria) of Asia and North America KENNETH CARPENTER Oklahoma Museum of Natural History, University of Oklahoma, Norman, Oklahoma 73019, U.S. A. * ' ABSTRACT The theropod family Tyrannosauridae (Dinosauria) is composed of four genera and seven species. All taxa are known from nearly complete skeletons and/or skulls, thus making it one of the best documented large theropod families. The stratigraphic and palaeobiogeographic distribution of the Tyrannosauridae extends from the lower Campanian to upper Maastrichtian of North America,and to the Campanian-Maas- trichtian of Asia. INTRODUCTION Tyrannosaurid theropods are known only from the Upper Cretaceous of Asia and North America. Their earliest record is a fragmentary skeleton (genus unknown) from the lower Campanian Eagle Ford Sandstone of Montana (U.S.A. ) (Gilmore, 1920). By the upper Campanian, however, tyrannosaurids occur through out the western Interior and Gulf Coast of North America. They are known to have survived until the latest Maastrichtian in the Western Interior. In Asia, tyran nosaurids are known only from the Nemegt Formation estimated to be Campanian- Maastrichtian in age (Fox, 1978). Their apparent absence from upper Maastrichtian deposits in Asia is probably not due to extinction, but due to the lack of upper Maas trichtian deposits. The earliest tyrannosaurids described were the result of explorations by the geo logical surveys of Canada and the United States. The first specimen consisted of sev eral isolated and scattered teeth collected from the Judith River Formation of Mon tana. These were the first theropod teeth found in North America and were named Deinodon horridusby Leidy (1857). -
Upper Jurassic Dinosaur Bonebeds at Ten Sleep, Wyoming: Stratigraphy, Preliminary Results and Field Reports of 2016 and 2017
André de Sousa Saleiro Barros Licenciado em Biologia Upper Jurassic dinosaur bonebeds at Ten Sleep, Wyoming: stratigraphy, preliminary results and field reports of 2016 and 2017 Dissertação para obtenção do Grau de Mestre em Paleontologia Orientador: Octávio Mateus, Professor Associado, Faculdade de Ciências e Tecnologia da Universidade Nova de Lisboa Júri: Presidente: Prof. Doutor Paulo Alexandre Rodrigues Roque Legoinha Arguente: Prof. Doutor Mark Allen Norel Vogal: Prof. Doutor Octávio João Madeira Mateus Setembro 2018 André de Sousa Saleiro Barros Licenciado em Biologia Upper Jurassic dinosaur bonebeds at Ten Sleep, Wyoming: stratigraphy, preliminary results and field reports of 2016 and 2017 Sketch of of Bobcat Hill by Simão Mateus Dissertação para obtenção do Grau de Mestre em Paleontologia Orientador: Octávio Mateus, Professor Associado, Faculdade de Ciências e Tecnologia da Universidade Nova de Lisboa Acknowledgments I start by thanking my supervisor, Professor Octávio Mateus, not only for the help provided during my work, but also for the trust placed in me, for the free range given by him, allowing me to pursue my own objectives, both academically and professionally. I thank the American Museum of Natural History for funding the expeditions which were the focal point of this work, and the Museum of Lourinhã for providing me with a workspace, as well as allowing me to use the laboratorial facilities to prepare the specimens here described. Special thanks to the team members, and visitors, of the expedition, without whom this project would not exit. All fieldwork was carried out under Bureau of Land Manegement permits PA16-WY-245, PA16-WY-255, and PA16-WY-256. -
Forelimbs of the Theropod Dinosaur Dilophosaurus Wetherilli: Range of Motion, Influence of Paleopathology and Soft Tissues, and Description of a Distal Carpal Bone
Palaeontologia Electronica palaeo-electronica.org Forelimbs of the theropod dinosaur Dilophosaurus wetherilli: Range of motion, influence of paleopathology and soft tissues, and description of a distal carpal bone Philip J. Senter and Corwin Sullivan ABSTRACT Forelimb bones of the Early Jurassic theropod Dilophosaurus wetherilli were man- ually manipulated to determine the range of motion (ROM) of each forelimb joint and to test functional hypotheses of forelimb use. Using bony articular surface margins to delimit ROM, the humerus can be retracted to a position subparallel to the scapular blade, protracted no further than a subvertical position, and elevated only 65°. The elbow cannot be flexed beyond a right angle to the humerus. Supination and pronation are prevented by lack of rolling radio-ulnar joints, and the palms face medially. The dis- tal carpal block of Dilophosaurus, described here for the first time, is relatively flat and lacks a proximal trochlea, suggesting limited wrist mobility. The fingers diverge during flexion and are extremely hyperextensible. Experiments on American alligator cadav- ers show that ROM exhibits limited intraspecific variation and is greater in the fully- fleshed elbow than skeletal manipulation suggests. Here, we apply these findings and those of previous studies on extant archosaurs to reconstruct ROM in the forelimbs of live Dilophosaurus by adjusting the results of the bare-bones manipulations to account for the known influence of soft tissues. We also discuss the effects of skeletal deformi- ties on inferred ROM in Dilophosaurus. Forelimb morphology and ROM are consistent with two-handed prehension, clutching objects to the chest, manually hooking objects, and seizing prey beneath the predator’s chest or the base of its neck (but no further forward). -
Lower Rotational Inertia and Larger Leg Muscles Indicate More Rapid Turns in Tyrannosaurids Than in Other Large Theropods
Lower rotational inertia and larger leg muscles indicate more rapid turns in tyrannosaurids than in other large theropods Eric Snively1, Haley O’Brien2, Donald M. Henderson3, Heinrich Mallison4, Lara A. Surring3, Michael E. Burns5, Thomas R. Holtz Jr6,7, Anthony P. Russell8, Lawrence M. Witmer9, Philip J. Currie10, Scott A. Hartman11 and John R. Cotton12 1 Department of Biology, University of Wisconsin-La Crosse, La Crosse, WI, USA 2 Department of Anatomy and Cell Biology, Oklahoma State University Center for Health Sciences, Tulsa, OK, USA 3 Royal Tyrrell Museum of Palaeontology, Drumheller, AB, Canada 4 Museum fur Naturkunde, Berlin, Germany 5 Department of Biology, Jacksonville State University, Jacksonville, AL, USA 6 Department of Geology, University of Maryland, College Park, MD, USA 7 Department of Paleobiology, National Museum of Natural History, Washington, D.C., USA 8 Department of Biological Sciences, University of Calgary, Calgary, AL, Canada 9 Department of Biomedical Sciences, Ohio University, Athens, OH, USA 10 Department of Biological Sciences, University of Alberta, Edmonton, AL, Canada 11 Department of Geoscience, University of Wisconsin-Madison, Madison, WI, USA 12 Department of Mechanical Engineering, Ohio University, Athens, OH, USA ABSTRACT Synopsis: Tyrannosaurid dinosaurs had large preserved leg muscle attachments and low rotational inertia relative to their body mass, indicating that they could turn more quickly than other large theropods. Methods: To compare turning capability in theropods, we regressed agility estimates against body mass, incorporating superellipse-based modeled mass, centers of mass, and rotational inertia (mass moment of inertia). Muscle force relative to body Submitted 6 September 2016 mass is a direct correlate of agility in humans, and torque gives potential angular Accepted 10 January 2019 acceleration.