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Arthropoda Selecta 21(1): 2960 © ARTHROPODA SELECTA, 2012 Description of some Salticidae (Aranei) from the Malay Archipelago. II. Salticidae of Java and Sumatra, with comments on related species Îïèñàíèå íåêîòîðûõ Salticidae (Aranei) èç Ìàëàéñêîãî Àðõèïåëàãà. II. Salticidae îñòðîâîâ ßâà è Ñóìàòðà ñ êîììåíòàðèÿì î áëèçêèõ âèäàõ Jerzy Prószyñski*, Christa L. Deeleman-Reinhold** É. Ïðóøèíüñêèé*, Ê.Ë. Äeeëåìàí-Ðåéíõîëüä** * Museum and Institute of Zoology, Polish Academy of Sciences, ul. Wilcza 64, 00-679 Warszawa, Poland. E-mail: [email protected] ** 4619GA Ossendrecht, the Netherlands. E-mail: [email protected] KEY WORDS: Salticidae, new species, diagnostic characters, geographical distribution, Indonesia, Java, Sumatra. ÊËÞ×ÅÂÛÅ ÑËÎÂÀ: íîâûé âèä, äèàãíîñòè÷åñêèå ïðèçíàêè, ðàñïðîñòðàíåíèå, Èíäîíåçèÿ, ßâà, Ñó- ìàòðà. ABSTRACT. This paper describes two new genera íîâûõ âèäîâ: Bathippus petrae sp.n., Donoessus (Junxattus gen.n. and Leuserattus gen.n.) and 12 new kerinci sp.n., Emathis sumatranus sp.n., Heratemita species from Sumatra and Java: Bathippus petrae sp.n., tenenbaumi sp.n., Junxattus daiqini sp.n., Leuserattus Donoessus kerinci sp.n., Emathis sumatranus sp.n., gunung sp.n., Nannenus maughami sp.n., N. siedleckii Heratemita tenenbaumi sp.n., Junxattus daiqini sp.n., sp.n., Orcevia kuloni sp.n., Orsima thaleri sp.n., Leuserattus gunung sp.n., Nannenus maughami sp.n., Phintella conradi sp.n. è Pseudicius javanicus sp.n.. N. siedleckii sp.n., Orcevia kuloni sp.n., Orsima tha- Âïåðâûå îïèñàíà ñàìêà Thianitara spectrum Simon, leri sp.n., Phintella conradi sp.n., Pseudicius javani- 1903 è îäíîâðåìåííî îïèñàí áëèçêèé âèä èç Òàè- cus sp.n., as well as first description of unknown fe- ëàíäà Th. thailandica sp.n. Ðîä Orcevia Thorell, 1890 male of Thianitara spectrum Simon, 1903. A related âûâåäåí èç ñèíîíèìèè ñ Laufeia Simon, 1889 ñîîò- species Th. thailandica sp.n. is described from Thai- âåòñòâåííî âîññòàíîâëåíû èëè óñòàíîâëåíû ñëåäó- land. Genus Orcevia Thorell, 1890 is removed from þùèå êîìáèíàöèè: Laufeia eucola (Thorell, 1890) = synonymy with Laufeia Simon, 1889, and following O. eucola Thorell, 1890, L. keyserlingi (Thorell, 1890) new combinations of SE Asian species created: Laufe- = O. keyserlingi Thorell, 1890, L. perakensis (Simon, ia eucola (Thorell, 1890) = O. eucola Thorell, 1890, L. 1901) = O. perakensis Simon, 1901, L. proszynskii keyserlingi (Thorell, 1890) = O. keyserlingi Thorell, Song et al., 1988 = O. proszynskii (Song et al., 1988). 1890, L. perakensis (Simon, 1901) = O. perakensis comb.n. Îáñóæäàåòñÿ òàêñîíîìè÷åñêèé ñòàòóñ è Simon, 1901, L. proszynskii Song et al., 1988 = O. ïðèâåäåíû ðèñóíêè äëÿ Epeus flavobilineatus proszynskii (Song et al., 1988). comb.n. Taxonomic (Doleschall, 1859), Leikung porosa (Wanless, 1978), problems are discussed and new drawings added to the Mintonia tauricornis Wanless, 1984, Phaeacius fim- species Epeus flavobilineatus (Doleschall, 1859), Lei- briatus Simon, 1900 è Taraxella sumatrana Wanless, kung porosa (Wanless, 1978), Mintonia tauricornis 1987. Ïåðåñìîòðåíû ãðàíèöû ðîäà Emathis Simon, Wanless, 1984, Phaeacius fimbriatus Simon, 1900, 1899. Óñòàíîâëåíî, ÷òî îí âêëþ÷àåò âèäû òîëüêî and Taraxella sumatrana Wanless, 1987. The genus èç Þãî-Âîñòî÷íîé Àçèè. Ïÿòü âèäîâ èç Êàðèáñêî- Emathis Simon, 1899 is redefined and its name re- ãî áàññåéíà, ðàíåå îòíîñèìûõ ê Emathis, ïåðåâåäå- stricted to the Southeastern Asian species only. Five íû â íîâûé ðîä Petemathis gen.n.: P. luteopunctata Caribbean species earlier considered in Emathis are (Petrunkevitch, 1930) comb.n., P. minuta (Petrun- transferred into a new genus Petemathis gen.n.: P. kevitch, 1930) comb.n., P. tetuani (Petrunkevitch, 1930) luteopunctata (Petrunkevitch, 1930) comb.n, P. minu- comb.n. è P. unispina (Franganillo, 1930) comb.n. ta (Petrunkevitch, 1930) comb.n., P. portoricensis (Petrunkevitch, 1930) comb.n (type species), P. tetu- ani (Petrunkevitch, 1930) comb.n. and P. unispina Introduction (Franganillo, 1930) comb.n. This is the second part in a series of papers dealing ÐÅÇÞÌÅ. Èç Ñóìàòðû è ßâû îïèñàíû äâà íî- with Salticidae of the Malay Archipelago. Java and âûõ ðîäà Junxattus gen.n., Leuserattus gen.n. è 12 Sumatra together have records of 190 species of Salti- 30 J. Prószyñski, C.L. Deeleman-Reinhold cidae, of which 55 have some sort of diagnostic draw- preliminary drawings made in 19941997 of sample ings for both sexes, 62 species have only a single sex specimens. They were made available in the internet illustrated, and 54 have not been illustrated, although database, with the intention of a more detailed study of specimens are preserved in some collections so they specimens and more species at a later date. could be redescribed in the future; 16 species are dubi- Unfortunately, the opportunity for such complementary ous. In addition to these rather poor statistics, the qual- studies never occurred, so we have decided to publish ity of the existing drawings is often insufficient for our drawings and notes of 1990s, as they are, in order identification, nor for the separation of related species, to serve as useful data for salticid taxonomists. while species without diagnostic drawings cannot be The main aim of this paper is to provide prelimi- identified. Identifications and definitions of species in nary reference diagnostic drawings to complement the Salticidae are based on comparison with existing diag- scanty literature data for certain South East Asian salti- nostic drawings of Salticidae in the literature, as sum- cid genera and species. The classification is provision- marized in the internet database Monograph of Salti- al in some cases, pending revisions of related genera, cidae (Araneae) of the World by Prószyñski [2011]. especially their insufficiently studied type specimens. By providing comparative drawings, basic for iden- An untapped source of taxonomic information are pho- tification, the present paper facilitates study of species tographs of Salticidae, available on the Internet, some not only from Java and Sumatra, but also for some of which are used in this paper to draw attention to species from a much broader area particularly Borneo, particularly interesting species. The authors realize the the Malay Peninsula and even Thailand. A broad com- limited sample sizes of the described material, but as- parison of species also permits the rearrangement of sume that they will promote future taxonomic research. several genera, especially Pseudicius Simon, 1885. The best method of describing species is compar- The methodology of taxonomic analysis in this pa- ing them directly, usually by placing them together per deserves some consideration. It is a long taxonomic under a stereomicroscope, and comparing simultaneous- tradition to describe characters of animals in words, ly particular characters, whilst at the same time docu- resulting in the tedious preparation of long texts, de- menting them by drawings or photographs. However, scribing morphology of a species, in a number of lan- some characters of the studied specimens had deterio- guages, and boring study of such descriptions for the rated so much during years of storage that old and fresh recognition of species. A refinement of that approach specimens were difficult to compare. This is well illus- is summarizing the data of numerous morphological trated by sets of photographs of both living and long characters in cladograms, usually arranged with the preserved specimens of the same species, made by help of specialized computer software. That results in Koomen [Prószyñski, 2011]. In addition, while females descriptions devoid of drawings, or with diagrammatic cannot be identified in many species without prepara- illustrations only, which do not permit recognition of tion of the epigyne, the preparation itself puts the or- the species. The other method, suitable for the majority gans in jeopardy: epigynes (and also palpus) stored in of Salticidae, is through the comparison of pictures of a microvials are getting lost during subsequent examina- species with other pictures, or with actual speci- tions, while those mounted in Canada Balsam on mi- mens. This method requires some artistic skills for the croscope slides become so transparent that they cannot preparation of drawings, or photographs, and the expe- be seen without restoration (dissolving the Canada Bal- rienced eye of a taxonomist. Hence, it is rather difficult sam, staining the preparation and mounting them again for beginners and people with poor picture perception. as a permanent or temporary slide). The safest and best Recently, Benjamin [2004] characterized the usage of documentation of internal structures of the epigyne is genitalic characters as single character systems be- perhaps the publication of good quality drawings, dia- lieved a priori to have greater importance than other grammatic illustrations are useless. characters in difference to the supposedly multi- There is no satisfactory standard terminology for character approach of cladistic analysis, which is ap- the description of the internal structures of Salticidae parently more scientific. Palpus, epigyne and internal epigynes, so the terminology I have used during the structures of the epigyne are single characters, in the last 50 years was rather intuitive. However, there is an same way as photographs of a human face are single excellent publication on these structures in Lycosidae, characters, although we recognize people by them. Pho- published recently by Useta et al. [2007], which we try tographs