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Sequence of Post-Moult Exoskeleton Hardening Preserved in a Trilobite Mass Moult Assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco
Editors' choice Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco HARRIET B. DRAGE, THIJS R.A. VANDENBROUCKE, PETER VAN ROY, and ALLISON C. DALEY Drage, H.B., Vandenbroucke, T.R.A., Van Roy, P., and Daley, A.C. 2019. Sequence of post-moult exoskeleton hardening preserved in a trilobite mass moult assemblage from the Lower Ordovician Fezouata Konservat-Lagerstätte, Morocco. Acta Palaeontologica Polonica 64 (2): 261–273. Euarthropods have a tough exoskeleton that provides crucial protection from predation and parasitism. However, this is restrictive to growth and must be periodically moulted. The moulting sequence is well-known from extant arthropods, consisting of: (i) the long inter-moult stage, in which no changes occur to the hardened exoskeleton; (ii) the pre-moult stage where the old exoskeleton is detached and the new one secreted; (iii) exuviation, when the old exoskeleton is moulted; and (iv) the post-moult stage during which the new exoskeleton starts as soft, thin, and partially compressed and gradually hardens to the robust exoskeleton of the inter-moult stage. Trilobite fossils typically consist of inter-moult carcasses or moulted exuviae, but specimens preserving the post-moult stage are rare. Here we describe nine specimens assigned to Symphysurus ebbestadi representing the first group of contemporaneous fossils collected that preserve all key stages of the moulting process in one taxon, including the post-moult stage. They were collected from a single lens in the Tremadocian part of the Fezouata Shale Formation, Morocco. Based on cephalic displacement and comparison to other trilobite moults, one specimen appears to represent a moulted exoskeleton. -
Databases in Vertebrate Paleontology Mark D
This article was downloaded by: [University of California, Berkeley] On: 08 January 2013, At: 09:29 Publisher: Taylor & Francis Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK Journal of Vertebrate Paleontology Publication details, including instructions for authors and subscription information: http://www.tandfonline.com/loi/ujvp20 From card catalogs to computers: databases in vertebrate paleontology Mark D. Uhen a , Anthony D. Barnosky b , Brian Bills c , Jessica Blois d , Matthew T. Carrano e , Marc A. Carrasco b , Gregory M. Erickson f , Jussi T. Eronen g h , Mikael Fortelius g , Russell W. Graham i , Eric C. Grimm j , Maureen A. O’Leary k , Austin Mast f l , William H. Piel m , P. David Polly n & Laura K. Säilä g a Department of Atmospheric, Oceanic and Earth Sciences, George Mason University, Fairfax, Virginia, 22030, U.S.A. b Department of Integrative Biology and Museum of Paleontology, University of California, Berkeley, California, 94720, U.S.A. c Center for Environmental Informatics, Pennsylvania State University, University Park, Pennsylvania, 16802, U.S.A. d Center for Climate Research, University of Wisconsin, Madison, Wisconsin, 53706, U.S.A. e Department of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, D.C., 20013, U.S.A. f Department of Biological Science, The Florida State University, Tallahassee, Florida, 32306, U.S.A. g Department of Geosciences and Geography, HY-00014, University of Helsinki, Finland h Senckenberg Research Institute und Nature Museum, Biodiversity and Climate Research Centre LOEWE BiK-F, Senckenberganlage 25, D-60325 Frankfurt am Main, Germany i Department of Geosciences, The Pennsylvania State University, University Park, Pennsylvania, 16802, U.S.A. -
The Case of the Diminutive Trilobite Flexicalymene Retrorsa Minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati Region
EVOLUTION & DEVELOPMENT 9:5, 483–498 (2007) Evaluating paedomorphic heterochrony in trilobites: the case of the diminutive trilobite Flexicalymene retrorsa minuens from the Cincinnatian Series (Upper Ordovician), Cincinnati region Brenda R. Hundaa,Ã and Nigel C. Hughesb aCincinnati Museum Center, 1301 Western Avenue, Cincinnati, OH 45203, USA bDepartment of Earth Sciences, University of California, Riverside, CA 92521, USA ÃAuthor for correspondence (email: [email protected]) SUMMARY Flexicalymene retrorsa minuens from the upper- rate of progress along a common ontogenetic trajectory with most 3 m of the Waynesville Formation of the Cincinnatian respect to size, coupled with growth cessation at a small size, Series (Upper Ordovician) of North America lived ‘‘sequential’’ progenesis, or non-uniform changes in the rate of approximately 445 Ma and exhibited marked reduction in progress along a shared ontogenetic trajectory with respect to maximum size relative to its stratigraphically subjacent sister size, can also be rejected. Rather, differences between these subspecies, Flexicalymene retrorsa retrorsa. Phylogenetic subspecies are more consistent with localized changes in analysis is consistent with the notion that F. retrorsa retrorsa rates of character development than with a global hetero- was the ancestor of F. retrorsa minuens. F. retrorsa minuens chronic modification of the ancestral ontogeny. The evolution has been claimed to differ from F. retrorsa retrorsa ‘‘in size of F. retrorsa minuens from F. retrorsa retrorsa was largely alone,’’ and thus presents a plausible example of global dominated by modifications of the development of characters paedomorphic evolution in trilobites. Despite strong similarity already evident in the ancestral ontogeny, not by the origin of in the overall form of the two subspecies, F. -
71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas. -
001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More by Michelle M
Introduction to the Trilobites: Morphology, Ecology, Macroevolution and More By Michelle M. Casey1, Perry Kennard2, and Bruce S. Lieberman1, 3 1Biodiversity Institute, University of Kansas, Lawrence, KS, 66045, 2Earth Science Teacher, Southwest Middle School, USD497, and 3Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045 Middle level laboratory exercise for Earth or General Science; supported provided by National Science Foundation (NSF) grants DEB-1256993 and EF-1206757. Learning Goals and Pedagogy This lab is designed for middle level General Science or Earth Science classes. The learning goals for this lab are the following: 1) to familiarize students with the anatomy and terminology relating to trilobites; 2) to give students experience identifying morphologic structures on real fossil specimens 3) to highlight major events or trends in the evolutionary history and ecology of the Trilobita; and 4) to expose students to the study of macroevolution in the fossil record using trilobites as a case study. Introduction to the Trilobites The Trilobites are an extinct subphylum of the Arthropoda (the most diverse phylum on earth with nearly a million species described). Arthropoda also contains all fossil and living crustaceans, spiders, and insects as well as several other extinct groups. The trilobites were an extremely important and diverse type of marine invertebrates that lived during the Paleozoic Era. They only lived in the oceans but occurred in all types of marine environments, and ranged in size from less than a centimeter to almost a meter across. They were once one of the most successful of all animal groups and in certain fossil deposits, especially in the Cambrian, Ordovician, and Devonian periods, they are extremely abundant. -
Active Research Grants
Linda C. Ivany Professor Department of Earth and Environmental Sciences Heroy Geology Laboratory,Syracuse University, Syracuse, NY 13244 phone: (315) 443-3626 / fax: (315) 443-3363 / email: [email protected] http://thecollege.syr.edu/people/faculty/pages/ear/Ivany-Linda.html https://orcid.org/0000-0002-4692-3455 Education Ph.D. in Earth and Planetary Sciences, 1997, Harvard University Advisor: Stephen Jay Gould M.S. in Geology, minor in Zoology, 1990, University of Florida-Gainesville Advisor: Douglas S. Jones B.S. in Geology, minor in Zoology, 1988, Syracuse University Advisor: Cathryn R. Newton Academic Positions 2012-present Professor of Earth Sciences, Syracuse University 2005-2012 Associate Professor of Earth Sciences, Syracuse University 2001-2005 Assistant Professor of Earth Sciences, Syracuse University 2000-2001 Visiting Assistant Professor of Earth Sciences, Syracuse University 1997-2000 Michigan Society Fellow and Visiting Assistant Professor of Geological Sciences, University of Michigan General Research Interests Evolutionary Paleoecology, Paleoclimatology, Stable Isotopes in Paleobiology I am a marine paleoecologist and paleoclimatologist. My interests lie broadly in the evolution of the Earth-life system and how ecosystems and their component taxa evolve and respond to changes in the physical environment. Specific areas of interest include biotic and climatic change during the Paleogene (~65-24 million years ago); use of geochemical data, particularly stable isotopes, derived from accretionary biogenic materials for inference -
Th TRILO the Back to the Past Museum Guide to TRILO BITES
With regard to human interest in fossils, trilobites may rank second only to dinosaurs. Having studied trilobites most of my life, the English version of The Back to the Past Museum Guide to TRILOBITES by Enrico Bonino and Carlo Kier is a pleasant treat. I am captivated by the abundant color images of more than 600 diverse species of trilobites, mostly from the authors’ own collections. Carlo Kier The Back to the Past Museum Guide to Specimens amply represent famous trilobite localities around the world and typify forms from most of the Enrico Bonino Enrico 250-million-year history of trilobites. Numerous specimens are masterpieces of modern professional preparation. Richard A. Robison Professor Emeritus University of Kansas TRILOBITES Enrico Bonino was born in the Province of Bergamo in 1966 and received his degree in Geology from the Depart- ment of Earth Sciences at the University of Genoa. He currently lives in Belgium where he works as a cartographer specialized in the use of satellite imaging and geographic information systems (GIS). His proficiency in the use of digital-image processing, a healthy dose of artistic talent, and a good knowledge of desktop publishing software have provided him with the skills he needed to create graphics, including dozens of posters and illustrations, for all of the displays at the Back to the Past Museum in Cancún. In addition to his passion for trilobites, Enrico is particularly inter- TRILOBITES ested in the life forms that developed during the Precambrian. Carlo Kier was born in Milan in 1961. He holds a degree in law and is currently the director of the Azul Hotel chain. -
Curriculum Vitae
CURRICULUM VITAE John P. Hunter Department of Evolution, Ecology, and Organismal Biology (EEOB) The Ohio State University at Newark (OSU-N) Email: [email protected] Websites: http://newark.osu.edu/directory/hunter-john.html https://eeob.osu.edu/people/hunter.360 Founders Hall 2192 1179 University Drive Newark, Ohio 43055 Phone: 740-366-9102 FAX: 740-366-5047 PROFESSIONAL EXPERIENCE • Associate Professor of EEOB (2011–present) • Associate Professor of Earth Sciences (2011–present); courtesy appointment • Visiting Associate Researcher, University of California, Santa Cruz (2012–2013); sabbatical • Assistant Professor of EEOB (2005–2011) • Assistant Professor of Earth Sciences (2007–2011); courtesy appointment • Visiting Assistant Professor of EEOB (2004–2005) • Assistant Professor (1997–2004) and Postdoctoral Fellow/Instructor (1996–1997), New York College of Osteopathic Medicine (NYCOM) of New York Institute of Technology (NYIT) • Research Associate, American Museum of Natural History, New York (2000–present) HIGHER EDUCATION 1997 Ph.D. (Anatomy), State University of New York (SUNY) at Stony Brook Dissertation Title: Adaptive Radiation of Early Paleocene Ungulates Committee: David W. Krause (advisor), John G. Fleagle, Frederick E. Grine, and J. David Archibald (external) 1991 Geological Field Methods. Albion College, Michigan (summer course). 1989–90 Postgraduate Research at the University of Helsinki, Finland. Structure and function of mammalian teeth. Sponsoring Scientist: Mikael Fortelius. 1989 A.B. (Biology and Classics), Brown University, Providence, Rhode Island Honors Thesis Title: Tooth Wear in Juvenile and Adult Hadrosaurs Readers: Christine M. Janis (advisor) and J. Kelley RESEARCH GRANTS 2014 National Science Foundation. DISSERTATION RESEARCH: The Evolution of the Hypocone in Microbats (Microchiroptera). PI: John Hunter; CoPI: Sarah Gutzwiller. -
Introduction to the Trilobites: Morphology, Macroevolution and More by Michelle M
Introduction to the Trilobites: Morphology, Macroevolution and More By Michelle M. Casey1 and Bruce S. Lieberman1,2, 1Biodiversity Institute and 2Department of Ecology and Evolutionary Biology, University of Kansas, Lawrence, KS 66045 Undergraduate laboratory exercise for sophomore/junior level paleontology course Learning Goals and Pedagogy This lab is intended for an upper level paleontology course containing sophomores and juniors who have already taken historical geology or its equivalent; however it may be suitable for introductory biology or geology students familiar with geological time, phylogenies, and trace fossils. This lab will be particularly helpful to those institutions that lack a large teaching collection by providing color photographs of museum specimens. Students may find previous exposure to phylogenetic methods and terminology helpful in completing this laboratory exercise. The learning goals for this lab are the following: 1) to familiarize students with the anatomy and terminology relating to trilobites; 2) to give students experience identifying morphologic structures on real fossil specimens, not just diagrammatic representations; 3) to highlight major events or trends in the evolutionary history and ecology of Trilobita; and 4) to expose students to the study of macroevolution in the fossil record using trilobites as a case study. Introduction to the Trilobites The Trilobites are an extinct subphylum of the Arthropoda (the most diverse phylum on earth with nearly a million species described). Arthropoda also contains all fossil and living crustaceans, spiders, and insects as well as several other solely extinct groups. The trilobites were an extremely important and diverse type of marine invertebrates that lived during the Paleozoic Era. They were exclusively marine but occurred in all types of marine environments, and ranged in size from less than a centimeter to almost a meter. -
Ordovician Stratigraphy and Benthic Community Replacements in the Eastern Anti-Atlas, Morocco J
Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco J. Javier Alvaro, Mohammed Benharref, Jacques Destombes, Juan Carlos Gutiérrez-Marco, Aaron Hunter, Bertrand Lefebvre, Peter van Roy, Samuel Zamora To cite this version: J. Javier Alvaro, Mohammed Benharref, Jacques Destombes, Juan Carlos Gutiérrez-Marco, Aaron Hunter, et al.. Ordovician stratigraphy and benthic community replacements in the eastern Anti- Atlas, Morocco. The Great Ordovician Biodiversification Event: Insights from the Tafilalt Biota, Morocco, 485, The Geological Society of London, pp.SP485.20, In press, Geological Society, London, Special Publication, 10.1144/SP485.20. hal-02405970 HAL Id: hal-02405970 https://hal.archives-ouvertes.fr/hal-02405970 Submitted on 13 Nov 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. The Geological Society Special Publications Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco --Manuscript Draft-- Manuscript Number: GSLSpecPub2019-17R1 Article Type: Research article Full Title: Ordovician stratigraphy and benthic community replacements in the eastern Anti-Atlas, Morocco Short Title: Ordovician stratigraphy of the Anti-Atlas Corresponding Author: Javier Alvaro Instituto de Geociencias SPAIN Corresponding Author E-Mail: [email protected] Other Authors: MOHAMMED BENHARREF JACQUES DESTOMBES JUAN CARLOS GUTIÉRREZ-MARCO AARON W. -
Of the Local Fossil Assemblages Us
Supplemental Data: References and Paleobiology Database collection numbers (where applicable) of the local fossil assemblages used to construct the ecological dominance dataset. Ordovician Bayer, T.N., 1967, Repetitive benthonic community in the Maquoketa Fm. (Ordovician) of Minnesota: Journal of Paleontology, v. 41, p. 417-422. (Collection 357 is pooled from many assemblages; 13 other collections from the paper are included) Berry, W.B.N., Lawson, D.A., and Yancey, E.S., 1979, Species diversity patterns in some middle Ordovician communities from California and Nevada: Palaeogeography, Palaeoclimatology, Palaeoecology, v. 26, p. 99-116. (Collections 296, 424, 425) Cisne, J.L., 1973, Beecher’s Trilobite Bed revisited: ecology of an Ordovician deepwater fauna: Postilla, v. 160, p. 1-25. (Collection 340) Frey, R.C., 1987, The paleoecology of a Late Ordovician shale unit from southwest Ohio and southeastern Indiana: Journal of Paleontology, v. 61, p. 242-267. (Collection 368, includes three pooled assemblages considered separately in the analysis). Frey, R.C., 1987, The occurrence of pelecypods in early Paleozoic epieric sea environments, late Ordovician of the Cincinnati, Ohio area: Palaios, v. 2, p. 3-23. (Collections 369, 371, 373, 377) Holland, S.M., Unpublished Data. (Collection 35351) Nilsson, R., 1977, A boring through Middle and Upper Ordovician strata at Koängen in western Scania, southern Sweden: Sveriges Geologiska Undersökning, Serie C, Avhandlingar och Uppsatser, v. 71, p. 1-58. (Collections 378, 379, 380) Parker, W.C., 1983, Fossil ecological successions in Paleozoic level bottom brachiopod- bryozoan communities: Unpublished Ph.D. Dissertation, University of Chicago, 217 p. (Collections 396, 398, 421) Patzkowsky, M.E., and Holland, S.M., 1999, Biofacies replacement in a sequence stratigraphic framework: Middle and Upper Ordovician of the Nashville Dome, Tennessee, USA: Palaios, v.