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Zeitschrift/Journal: Beiträge zur Paläontologie

Jahr/Year: 1999

Band/Volume: 24

Autor(en)/Author(s): Kennedy William James, Summesberger Herbert

Artikel/Article: New Late Campanian Ammonites from the Gschliefgraben near Gmunden (Ultrahelvetic, Austria) 23-39 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien Beitr. Paläont. 24:23-39, Wien 1999

New Late Campanian Ammonites from the Gschliefgraben near Gmunden (Ultrahelvetic, Austria)

by

William J. K ennedy 1 & Herbert Summesberger 1 2

Kennedy , WJ. & S ummesberger , H., 1999. New Late Campanian Ammonites from the Gschliefgraben near Gmunden (Ultrahelvetic, Austria). - Beitr. Palaont., 24:23-39, 1 Text-fig., 3 Plates, Wien.

Contents [in:] Tr Oger et al., 1999, this volume). The first note on the ammonite fauna of the Gschliefgraben was pub­ Abstract, Zusammenfassung...... 23 lished by M ojsisovics & Schl Onbach (1868, p. 213: Introduction...... 23 „Baculites, Hamites, Scaphites, Ammonites“). B rink ­ Systematic Palaeontology 24 mann (1935) noted five ammonite taxa. Twenty-five Sedimentology 31 Stratigraphic conclusions...... 31 taxa were described by K ennedy & S ummesberger Palaeobiogeography...... 31 (1984). The geologic situation of the Gschliefgraben References ...... 31 tectonic window is best described in Prey (1983) and Kennedy & S ummesberger (1984). The belemnites were published recently by Christensen (1998). The Abstract bivalves (Tröger , S ummesberger & S koumal 1999), Eleven taxa are described from the Late Campanian of echinoderms (Jagt 1999), crustaceans (Fraaye & Sum ­ the Gschliefgraben near Gmunden. Six of them are new mesberger 1999) and nannofossils (W agreich 1999) for the Gschliefgraben: Kitchinites sp., Yokoyamao- were investigated in an integrated study and are pub­ ceras ? sp., Hoplitoplacenticeraspreyi sp. nov., Didy- lished in this volume. The sponges will also be studied moceras binodosum (K ennedy & Cobban , 1993), and published in a subsequent volume. There is no Pseudoxybeloceras (Schlueterella) pseudoarmatum measurable section at the Gschliefgraben; a synthetic (Schlüter , 1872) and Polyptychoceratinae indet. Taken section (Kennedy & S ummesberger 1984, Text-fig. 3) together with the previously described forms, the am­ is based upon estimated thicknesses and stratigraphic monite fauna of the Gschliefgraben comprises 32 taxa. data in Prey (1983). All fossils were collected from the surface of an active mudflow, most of them by Zusammenfassung amateur collectors. About 250 ammonite specimens (32 taxa) comprising Aus dem Obercampan des Gschliefgrabens bei Gmun­ 18 heteromorph species, and 14 non-heteromorphs are den werden elf Taxa beschrieben, sechs davon neu für preserved in the collections of the Natural History die Fundstelle: Kitchinites sp., Yokoyamaoceras ? sp., Museum Vienna and the Austrian Geological Survey, Hoplitoplacenticeras preyi sp. nov., Didymoceras bino­ the private collections of Mag. Ulrich Roschger (Vien­ dosum (Kennedy & Cobban , 1993), Pseudoxybeloceras na), Dr. Peter S koumal (Vienna), Wolf-Peter M aherndl (Schlueterella) pseudoarmatum (S chlüter , 1872) und (Bad Ischl), and Ferdinand E stermann (Pinsdorf, Up­ Polyptychoceratinae indet. Zusammen mit den früher per Austria). The former S chauberger collection, the beschriebenen umfaßt die Ammonitenfauna des VA vra collection (inoceramids) and the Prey collec­ Gschliefgrabens 32 Taxa. tion (included in the legacy of Othmar K ühn ) are now integrated in the NHMW collection. The H ütter col­ Introduction lection (HÜ of K ennedy & S ummesberger 1984) was The Gschliefgraben is situated in Upper Austria, east donated to the Natural History Museum Vienna in 1998. of the Traunsee, some 55 km SW of Linz (Text-fig. 1 Conventions NHMW Naturhistorisches Museum Wien 1 Oxford University Museum of Natural History, Oxford, 0X1 3PW, United Kingdom RO Collection Roschger , Vienna 2 Naturhistorisches Museum, Burgring 7, A-1014 Wien, SK Collection S koumal , Vienna Austria ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

24 Beitr. Palàont. 24, Wien 1999

Systematic Palaeontology mesberger , 1984, p. 156, is the original of Seunes , 1892, pi. 13(4), fig. 2; paralectotype is the original of S eunes , Order Ammonoidea Zittel, 1884 1892, pi. 12(3), fig. 2; pi. 13(4), fig. 3, from the Late Suborder Phylloceratina A rkell , 1950 Maastrichtian, Assize à Pachydiscus Fresvillensis, P Superfamily Phyllocerataceae Zittel, 1884 Jacquoti et Stegaster, Gan, Route de Gan à Rébénacq, Family Phylloceratidae Z ittel, 1884 Pyrénées-Atlantiques, France, in the Sorbonne Collec­ Subfamily Phylloceratinae Zittel, 1884 tions, Paris. Genus Phylloceras S uess , 1866 Material: NHMW/1998z/29/2, a single specimen. Type species: Ammonitesheterophyllus J.Sow- Discussion: The adult individual of 90 mm in erby , 1820, p. 119, pi. 226, by monotypy. diameter compares well with the lectotype, showing the biconcave constrictions and delicate associated Phylloceras sp. ? collar ribs on the body chamber. (Pi. 1, Figs. 1-3) Occurrence: Late Campanian to Late Maastricht­ ian, Pyrénées-Atlantiques and Landes in France, the ? 1984 Phylloceras (Hypophylloceras ) sp. Kennedy and SUMMESBERGER, p. 153, pi. 1, fig. 5. coastal sections of the Biscay region of France and NW Spain, the Gschliefgraben, Austria, and possibly Mada­ Discussion: What appears to be a second speci­ gascar. men of a phylloceratid from the Gschliefgraben is rep­ resented by NHMW/1998z/29/l, a very crushed indi­ Subfamily Puzosiinae Spath , 1922 vidual with a maximum preserved diameter of 60 mm. Genus Kitchinites Spath , 1922 Delicate lirae, typical of the group, are preserved on Type species Holcodiscus pondicherryanus the outer flank and venter of the initial 240° of the outer Kossmat , 1897, p. 40(147), pi. 6(17), fig. 6, by origi­ whorl; the last 90° sector, possibly the adapertural part nal designation. of the body chamber, is characterized by growth lines only. The specimen is specifically indeterminate, and Kitchinites sp. in the absence of sutural details, cannot be placed in a (PI. 1, Figs. 4, 5) subgenus. Description NHMW/1998z/29/3 (pi. 1, fig. 4) Occurrence: Late Campanian of the Gschlief­ is a crushed composite mould 46 mm in diameter. Coil­ graben. ing appears to have been fairly involute, the umbilicus Suborder Ammonitina Hyatt, 1889 comprising 30% of the diameter. The original whorl Superfamily Desmocerataceae Zittel, 1895 proportions cannot be established, but the umbilicus Family Desmoceratidae Zittel, 1895 appears to have been shallow with a low wall, the Genus Desmophyllites S path , 1929 umbilical shoulder narrowly rounded, whorl sides

Type species Desmoceras larteti S eunes , subparallel and the venter and ventrolateral shoulders 1891, p. 19, pi. 12(3), fig. 2; pi. 13(4), figs. 2, 3, by sub­ broadly rounded: a planulate shell form. There are sequent designation by S path , 1921, p. 46, as type spe­ traces of four constrictions on the outer whorl; promi­ cies of Schlueteria, of which Desmophyllites is the re­ nent, straight and prorsiradiate on the flanks, they placement name. sweep forward and are concave across the ventrola­ teral shoulder, forming a marked chevron on the ven­ Desmophyllites larteti (S eunes , 1892) ter. The constrictions are flanked by low, incipiently (PI. 1, Figs. 6-9) bullate collar ribs. Delicate striae arise on the inner 1892 Desmoceras larteti Seunes , p. 19, pi. 12(3), fig. 2; flank between constrictions, and strengthen into pi. 13(4), figs. 2, 3. flexuous riblets, convex across the middle of the flank 1984 Desmophyllites larteti (Seunes , 1891); Kennedy and Summesberger , p. 156, pi. 1, fig. 6; pi. 7, figs. 1-3, and feebly convex on the outer flank, where they are 5, 6 (with synonymy) at their maximum development, and are relatively 1992 Desmophyllites larteti (Seunes , 1891); Kennedy and coarse. NHMW/1998z/29/4 (PI. 1, fig. 5) is a very Henderson , p. 407, pi. 18, figs. 1-6. crushed individual with a maximum preserved diam­ 1993 Desmophyllites larteti (Seunes , 1891); Ward & eter of 65 mm. Ornament is as in the previous speci­ Kennedy , p. 23, figs. 23.1, 23.2, 23.3, 23.5-23.7, 23.9-23.12, 24, 25.15. men to a diameter of 50 mm, beyond which the speci­ 1993 Desmophyllites larteti (Seunes , 1892); Hancock & men appears to be body chamber, lacking constrictions, Kennedy , p. 154, pi. 2, figs. 1-3, 10, 11, 14; pi. 3, with smooth inner flanks, the outer flanks ornamented figs. 1, 4, 5. by relatively distant coarse ribs of variable strength. 1993 Desmophyllites larteti (Seunes , 1892); Kennedy & Discussion: These specimens are interpreted as Hancock , p. 578, pi. 1, figs. 1-3, 8-10. poorly preserved examples of Kitchinites on the basis Types Lectotype, designated by Kennedy and Sum - of the collar ribs and constrictions, with near-smooth ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

Kennedy , W.J. & Summesberger , H., New Late Campanian Ammonites ... 25 inner, and distantly ribbed outer flanks. The ornament Type species: Hoplites-Placenticerasplasticus is different from that of previously described species, Paulcke , 1907, p. 186; ICZN Opinion 555, Name suggesting that a new taxon is represented. Reference no. 1629. is made to M atsumoto (1988) for a recent review of Kitchinites. S ummesberger (1979) listed species re­ Hoplitoplacenticeras (Hoplitoplacenticeras) ferred to the genus at that date; other references are preyi sp. nov. H enderson (1970), M acellari (1986) and Stinnesbeck (PI. 3, Figs. 1,4) (1986). Type species The holotype is NHMWzl998/ Occurrence: Late Campanian of the Gschlief- 29/0013, collected by Dr. Prey , incorporated together graben. with the legacy of Prof. Othmar K ühn in the collec­ Puzosiinae gen. et sp. indet. tions of the Museum of Natural History. Discussion: We have seen very large fragments Derivatio nominis: named after Dr. Sieg- of indeterminate Puzosiinae in the Estermann Collec­ mund Prey (1912-1992), brilliant field geologist of tion (Pinsdorf near Gmunden, Upper Austria). the Austrian Geological Survey, who described the geology of the Gschliefgraben (1951, 1983) and col­ Family Kossmaticeratidae S path , 1922 lected the holotype. Subfamily Kossmaticeratinae S path , 1922 Locus typicus: Gschliefgraben near Gmunden Genus Yokoyamaoceras W right & M atsumoto , (Upper Austria), Ultrahelvetic tectonic zone. 1954 Stratum typicum: Gschliefgraben shales (Cre­ Type species Holcodiscuskotoi Jimbo , 1894, taceous). p. 33, pi. 21, fig. 2, by original designation. Description: Crushed and corroded internal mould of a phragmocone with body chamber and ap­ Yokoyamaoceras ? sp. erture preserved. Lateral compaction makes observa­ (PI. 2, Fig. 11) tion of venter almost impossible. Measurements are: Description: NHMW/1998z/29/5 is a tiny dis­ D 95 mm, Wh 53 mm, U 12,9 mm, U% 13,6. The shell torted individual with a diameter of 14.5 mm. Coiling appears to have been compressed, with rapidly increas­ appears to have been moderately involute, with a deep ing whorl height and a rather small umbilicus. The umbilicus, narrowly rounded umbilical shoulder, broad­ umbilical wall is vertical and slightly convex, the um­ ly rounded flanks, more narrowly rounded ventrola­ bilical shoulder narrowly rounded. The flanks are teral shoulder and broadly convex venter. Two narrow, slightly convex, the ventrolateral shoulders gently deep constrictions are visible on the outer whorl, ap­ rounded. The aperture is high oval. On the phragmo­ proximately 180° apart, and associated with narrow cone, relative strong, flexuous closely spaced ribs arise collar ribs, the adapical with tiny ventral tubercles. The in pairs from a tiny umbilical tubercle, crossing the sector of shell between the ribs bears four tiny ventral flank with a double concavity; they are prorsiradiate tubercles; obscure traces of ventrolateral ribs are as­ on the inner flank, and markedly rursiradiate on the sociated with the tubercles. outer. All bear a tiny ventral clavus. Ribbing weakens Discussion This specimen is a microconch on the body chamber, towards the aperture. On the body desmoceratacean, but we are uncertain of its taxonomic chamber pairs of ribs arise without a tubercle or bulla position. The simple style of ornament recalls some of on the inner third of the flank crossing the venter with the diminutive Menuites described by M atsumoto a slight flexure and flexing forwards to join in pairs at (1955), but these do not acquire tubercles until a larger eight large distantly spaced ventrolateral clavi. The last diameter, and have a rounded, rather than subrectan- pair of ribs close to the aperture are fused into a broad gular whorl section. Early acquisition of tubercles and and shallow single rib. The total number of ribs on the whorl section more closely resemble those of Yoko­ outer whorl is about 50. The venter is damaged by yamaoceras W right & M atsumoto , 1954 (see Kenne ­ compaction. Outer ventrolateral clavi are visible only dy , 1986, M atsumoto , 1991, andMAEDA, 1993 for dis­ between the last three pairs of ribs. cussion), to which genus we provisionally refer the Discussion: Despite the poor preservation the specimen. Occurrence: Late Campanian of the Gschlief- specimen appears to be distinct from all other Euro­ graben. pean species. See the extensive descriptions and dis­ cussions of Kennedy (1986:63 ff.; revision of the ge­ Superfamily Hoplitaceae H. D ouvill £, 1890 nus); Kaplan , K ennedy & E rnst (1996:34 ff.) and Family Placenticeratidae Hyatt, 1900 Kennedy & Kaplan (1997:49 ff.) Genus and Subgenus Hoplitoplacenticeras H. (H.) vari (S chlüter , 1876) is more regularly ribbed P aulcke , 1907 without looped pairs of ribs. Umbilical tubercles ex­ ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

26 Beitr. Paläont. 24, Wien 1999

tend to the aperture. The surface is covered with dense 1997 Bostrychoceras polyplocum (Roemer , 1841); lirae (e.g. Kaplan , K ennedy & E rnst , 1996, pi. 19; Kennedy and Kaplan , p. 52, pis. 39-50 (with full pi. 20; figs. 1, 4, pi. 21; figs. 1, 4). synonymy). H. (H.) coesfeldiense (S chlüter , 1867) is densely Type: Neotype, by the subsequent designation of ribbed throughout ontogeny, ribs being bundled in Kennedy & Kaplan (1997), is the lower part of speci­ threes and fours, umbilical bullae persisting to the body men no. 37092 in the collections of the Natural His­ chamber (e.g. Kaplan , Kennedy & Ernst , 1996, pl.21, tory Museum, London, from the Late Campanian of figs.3,4). Outer ventrolateral clavi connecting the ribs Haldem, Westphalia. Material: NHMW/1998z/29/6; NHMW/1998z/40/l. in twos and threes are described by Kaplan , Kennedy Discussion: Two well-preserved spires, one & Ernst , 1996 (p. 39: „äußere Ventrolateralclavi“). dextral, one sinistral, represent the early whorls of this This feature differs from that of the present species in species. They show the coiling to be relatively tight, that the ribs are dense and equal. and the whorls nearly in contact, while there is only a H.(H.) costulosum (S chlüter , 1867) bears narrow ribs, single row of tubercles, a rather different coiling to and never has distantly spaced ventrolateral clavi (e.g. that shown by the abundant topotype material figured Kaplan , Kennedy & Ernst , 1996, pi. 20, fig. 2, pi. 22, by Kennedy & Kaplan (1997), corresponding, as does figs. 2, 5; pi. 23, fig. 5; pi. 24, figs. 1-6, 8; pi. 27, the tuberculation, to that of Bostrychoceras unituber- fig. 1). Tiny umbilical tubercles or bullae persist onto culatum Blaszkiewicz , 1980 (p. 21, pi. 3, fig. 1-8; pi. 4, the body chamber. figs. 3-6), notably the original of his pi. 3, figs. 2 and H. (H.) dolbergense (S chlüter , 1876) is more evolute 4. Kennedy (1986) and Kennedy & Kaplan (1997) re­ and has much coarser ornament and umbilical tuber­ garded B. unituberculatum as a synonym of a variable cles on phragmocone and body chamber. B. polyplocum’, the present material confirms the tighter H. (H.) marroti (Coquand , 1859), revised by Kennedy coiling of unituberculatum morphotypes during early (1986, p. 70) differs by its coarser and more regular development. ornament, ribs arising in pairs from a coarse bulla and Occurrence: Late Campanian, Germany; Nor­ terminating separately in a ventrolateral clavus (e.g. folk, England; Northern Ireland; northern Aquitaine and Kennedy , 1986, pis. 9, 10). Its coiling is less involute Landes in France; Catalonia and Navarra, Spain; Po­ than in H. (H.) preyi sp. nov. land; Donbass, European Russia; Turkmenia; Kazakh­ H. rejaudryi (de G rossouvre , 1894) has narrower, sharp stan; Tadzhikistan; Iran; Bulgaria; North Africa; Texas ribs arising from long bullae, and a wider umbilicus and northern Mexico. (Kennedy 1986, pi. 7, figs. 1-3; pi. 14, figs. 6, 7). H. gosseleti (de G rossouvre , 1894, pi. 36, fig. 1) is Genus Didymoceras H yatt , 1894 much more evolute and has strong, straight ribs. Type species Ancyloceras nebrascense M eek (M eek, H. vancouverensis 1861) differs by its coarse, & H ayden , 1856, p. 71, by original designation. nearly straight ribs and numerous fine striae (U sher 1952, p. 93, pi. 25, figs. 1, 2). Didymoceras binodosum (Kennedy & Cobban , H. rarecostatum (K hakimov , in A tabekian & K ha - 1993) (PI. 2, Figs. 12, 13; Fig. 1) kimov, 1976, p. 88, pi. 10, fig. 5) has coarse, distant long and short ribs, long ribs sometimes arising in pairs 1963 Bostrychoceras secoense Y oung , p. 42 (pars), pi. 4, from coarse bullae, with strong inner and outer vent­ fig. 8 only. 1980 Didymoceras cf. secoenes (sic) (Y oung ); Blaszkie ­ rolateral tubercles. wicz , p. 24, pi. 5, figs. 4, 6; pi. 7, figs. 16, 19. Occurrence: As for type. 1993 Didymoceratoides binodosum Kennedy & Cobban , p. 92, figs. 8.1, 8.2, 8.5, 8.6, 8.13-8.15, 8.22-8.24, Suborder Ancyloceratina W iedmann , 1966 8.28, 8.29, 8.32, 8.33, 8.35-8.39, 9.1-9.5, 12.1. Superfamily Turrilitaceae G ill, 1871 1994 Didymoceratoides binodosum Kennedy & Cobban ; H yatt , Family Nostoceratidae 1894 Emerson , Emerson , A kers and A kers , p. 314, un­ Genus Bostrychoceras H yatt , 1900 numbered figures. Type species Turrilites polyplocus Roemer , 1997 Didymoceras binodosum (Kennedy & Cobban , 1841, pi. 14, figs. 1, 2, by original designation. 1993b); Larson , Jorgensen , Farrar & Larson , p. 53, unnumbered figures. Bostrychoceras polyplocum (Roemer , 1841) 1999 Didmoceras binodosum (K ennedy & Cobban , (PI. 2, Figs. 4, 5) 1993a ); Kennedy & Cobban , p.72, pis. 2-4; pi. 5, figs. 11-20; text-figs. 3-4. 1841 Turrilites polyplocus Roemer , 1841, p. 92, pi. 14, fig. 1 only, non 2 (= Eubostrychoceras saxonicum Types: Holotype is USNM 441521, from the Pierre (Schlüter , 1876)). Shale, Baculites scotti Zone, USGS Mesozoic locality ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

Kennedy , W.J. & Summesberger , H., New Late Campanian Ammonites 27

D1412, about 1.6 km north-northeast of Oral in the United States Western Interior and Gulf Coast, a re­ NW1/4 NW1/4 sec. 26, T.7 S., R.7 E., Fall River construction of which is shown in Text-fig. A. Kenne ­ County, South Dakota. dy & Cobban (1993) interpreted juvenile body cham­ Material Two specimens: RO/1986/3; NHMW/ bers of this species as adult body chambers of a paedo- 1998z/29/8. morphic derivative of Didymoceras. They have subse­ Description Complete specimens of Didymo- quently demonstrated (Cobban & Kennedy , 1999) these ceras binodosum from the United States show the shell loosely coiled planispiral whorls to be the early growth to consist of a loose open planispire (as in the holotype), stages of a form with helical whorls in middle growth followed by a loose elliptical or circular whorl; the later as is Didymoceras of which Didymoceratoides Kenne ­ whorls form a helix with the whorls barely in contact dy & Cobban , 1993, with D. binodosum as type spe­ except for the last part of the body chamber that is cies, is therefore a synonym. The style of coiling, regu­ slightly uncoiled. Ornament consists of prominent, lar bituberculate ribs and periodic constrictions distin­ narrow ribs, most of which bear two tubercles. Widely guish D. binodosum from other species referred to the spaced constrictions are present (Text-fig. 1). The ma­ genus. terial from the Gschliefgraben includes a specimen Occurrence: Late Campanian of the Gschlief­ with just over two whorls from the loosely coiled early graben, Austria and Poland. Baculites scotti Zone of growth stage (RO/1986/3; pi. 2, fig. 12) with well-de­ South Dakota, Utah, and Colorado; Bergstrom Forma­ veloped constrictions, and bituberculation that varies tion in Travis County, Texas; Annona Chalk near Yancy, from clearly differentiated to more-or-less obscured by Arkansas. Some of the poorly preserved Didymoceras post mortem crushing and abrasion. Much better pre­ sp. from the Wenonah Formation of New Jersey may served is NHMW/1998z/29/8, with parts of three also belong to this species (Kennedy & Cobban , 1994). whorls from the more tightly coiled helical stage, pre­ served to a maximum whorl height of 38 mm (PI. 2, Family Diplomoceratidae Spath , 1926 fig. 13). Ribbing is dense and even, with tuberculation Subfamily Diplomoceratinae Spath , 1926 evenly developed; there are occasional constrictions. Genus Neancyloceras Spath , 1926 Discussio n The Gschliefgraben specimens dif­ Type species Ancylocerasbipunctatum Schlü ­ fer in no significant respects from material from the ter, 1872, p. 98, pi. 29, figs. 1-3, by original designa­ tion.

Neancyloceras bipunctatum (Schlüter , 1872) (PI. 2, Fig. 6)

1872 Ancyloceras bipunctatum Schlüter , p. 98, pl. 29, figs. 1-3. 1935 Ancyloceras bipunctatum Schlüter ; Brinkmann , p. 5. 1982 Ancyloceras bipunctatum Schlüter , 1872; Klinger , p. 221, figs. 1 (1-3) (with full synonymy). 1982 Exiteloceras bipunctatum (Schlüter , 1872); Klinger , p. 222 ff., figs. 2-8A-E, 9. 1986 Neancyloceras cf. bipunctatum (S chlüter , 1872a); Kennedy , p. 104, pl. 16, fig. 5. 1993 Neancyloceras bipunctatum (Schlüter , 1872); Kennedy , p. 107, pl. 3, figs. 11-16, 20, 21; text- fig. 3. 1993 Neancyloceras cf. bipunctatum (S chlüter , 1872a); Kennedy & Cobban , p. 139; pl. 6, fig. 17. 1996 Neancyloceras bipunctatum (Schlüter , 1872); Wright , p. 250. ? 1997 Neancyloceras cf. bipunctatum (Schlüter , 1872); Kennedy & Kaplan , p. 61. Types Lectotype, by the subsequent designation of Blaszkiewicz , 1980, p. 29, is the original of Schlü ­ Figure 1. Restoration of Didymoceras binodosum. (Kennedy ter, 1872, pl. 29, fig. 3 (refigured by Klinger , 1982, and Cobban , 1993). Drawing by John R. Stacy is based on figs. 3, 4a), no. 65-10 in the collections of the Mu­ specimens from the Pierre Shale of South Dakota and is seum der Georg-August Universität, Göttingen, from approximately natural size. Figure supplied by Dr. W. A. Cobban (Denver). the Late Campanian of Ahlten, Germany. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien 28 Beitr. Paläont. 24, Wien 1999

Description NHMW/1998z/29/9 is a slightly Type: The holotype, by monotypy, is the original curved shaft 99 mm long, with a maximum preserved of Schlüter , 1872, pl. 31, figs. 1-3, no. 68 in the col­ whorl height of 12.4 mm. The rib index is four, the lections of the Paläontologisches Institut of Bonn Uni­ ribs effaced on the dorsum, narrow, distant, prorsiradi- versity, from the Campanian near Damp, Westphalia. ate and straight to feebly convex across the flanks, and Description NHMW/1998z/29/10 is an exter­ nal mould of a straight to slightly curved shaft 70 mm strong and transverse on the venter, where they bear long, with a maximum preserved whorl height of 17 delicate rounded-feebly clavate ventral tubercles. mm. The fragment shows two rows of tubercles borne Discussion The fragment agrees well with ju­ on a relatively strong, transverse rib, separated by up venile topotypes figured by Klinger (1982, figs. 5, 6, to four weaker nontuberculate ribs. Much larger is 7a-c). NHMW/1998z/29/12. It is a straight shaft with a curved Occurrence: Late Campanian of Germany; Po­ part of the body chamber preserved. The total length is land; the Gschliefgraben, Austria; European Russia; 600 mm from the adapical part of the shaft to the apex northern Aquitaine, France. Specimens from the Mons of the curvature. From the apex to the preserved adoral Basin, Belgium, described by Kennedy (1993), are said part of the body chamber it measures 176 mm. An ad­ to be from the Early Maastrichtian Craie phophatee de ditional fragment of the body chamber (NHMW/1998z/ Ciply. 29/11) that is tentatively referred to the present spe­ cies measures 201 mm in length. The general shape Subfamily Polyptychoceratinae Matsumoto , 1938 seems to consist of parallel straight shafts connected Genus Pseudoxybeloceras W right & by curved portions. Initial whorls are unknown. Whorl section may have been rounded or oval. The fragment M atsumoto 1954, p. 119 of the body chamber described below suggests a high Type species Hamites quadrinodosus Jimbo , oval section. On the straight portion of the shaft about 1894, p. 39, pi. 7, figs. 3, 4, by original designation. 24 narrow, straight and rectiradiate ribs per 100 mm cross the venter and dorsum without interruption. The Subgenus W iedmann , 1962 Schlueterella ribs are linked together in pairs by tubercles that are Type species Ancyloceras pseudoarmatum clearly spine-bases. These are arranged into four regu­ Schlüter , 1872, p. 99, pi. 31, figs. 1-3, b original des­ lar rows, in mid-flank and ventrolateral positions. Tu- ignation. berculated ribs are separated by one, two or three nontuberculated ones. There is a distinct change of Pseudoxybeloceras (Schlueterella) pseudoarmatum ornament on the final hook. Ribs become stronger and (Schlüter , 1872) are grouped in pairs, with interspaces of about the same (PI. 2, Fig. 10; PI. 3, Figs. 3, 5) width as one pair of ribs; the tubercles increase in strength. Ribs become rursiradiate from the apex of 1872 Ancyloceras (?) pseudoarmatum Schlüter , p. 99, pl. 31, figs. 1-3. the curvature on. This style of ribbing is even coarser 1876 Ancyloceras pseudoarmatum Schlüter ; on the large fragment of body chamber (NHMW/ Schlüter , p. 164, pl. 43, figs. 5-9. 1998z/29/l 1) that may also belong to this species. 1913 (?) Anisoceras pseudoarmatum Schlüter sp.; About 18 double ribs per 100 mm are fused on the N owak , p. 385, pl. 40, fig. 4. flank to broad flat ones. Tubercles are relatively weak 1925 Anisoceras pseudoarmatum Schlüter ; D ie­ and not every rib is tuberculated. ner , p. 73. Discussion We follow W right (1996, p. 253) ? 1974 Neocrioceras aff. pseudoarmatum (Schlüter , in treating Schlueterella as a subgenus of Pseudoxy­ 1872); N aidin , p. 167, pl. 57, fig. 2. beloceras rather than of Neocrioceras because A. pseu­ ? non 1982 Neocrioceras (Schlueterella ) aff. pseudo­ doarmatum, , type species of the subgenus, consists armatum (Schlüter ); Szäsz , p. 48, pl. 1, mainly of parallel shafts whereas Crioceras spinigerum fig. 1. Jimbo , type species of Neocrioceras, is crioceratiticone. 1986 Neocrioceras (Schlueterella) pseudoarmatum The closest specimen described is N. (S.) aff. pseudo­ (Schlüter , 1872a); Kennedy , text-fig. 37. armatum (Schlüter , 1872a) figured by Kennedy & 1996 Pseudoxybeloceras( Schlueterella) pseudoar­ Kaplan (1997, pl. 63, figs. 1-3). It differs in the larger matum (Schlüter , 1872); Wright , p. 253, fig. 196/1 a, b. distances between tuberculated ribs and consequently ? 1997 Neocrioceras (Schlueterella) aff. pseudoar­ greater number of intercalatories. The ornament of the matum (S chlüter , 1872a); Kennedy and body chamber hook is unknown. The shaft of the Kaplan ; p. 60; pl. 59, fig. 4; pl .62, figs. 1- holotype, refigured by Kaplan & Kennedy (1997, 3; pl. 63, figs. 1-3. pl. 65, figs. 1^1) and material from the Schlüter Col­ 1997 Neocrioceras (Schlueterella) pseudoarmatum lection (Bonn) refigured by Kaplan & Kennedy (1997, (S chlüter , 1872a); Kennedy & Kaplan ; pis.64,65) shows coarser ribbing with only 1-2 inter­ pl. 64, figs. 1-3; pl. 65, figs. 1-5. calatories. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

Kennedy , W.J. & Summesberger , H., New Late Campanian Ammonites 29

Occurrence Late Campanian Polyplocum Zone to the adult aperture is characterized by an abrupt loss of the Gschliefgraben, early Late Campanian of NW of ribs, the surface bearing growth lines and irregular Germany. constrictions and folds. The aperture is marked by a pronounced collar-rib. These macroconchs have whorl Subfamily Polyptychoceratinae M atsumoto , 1938 heights of 14.8 mm at the aperture. The microconch is Genus and subgenus Polyptychoceras Y abe , 1927 represented by SK. 1985.20 (Pl. 2, figs. 1-3), which Type species: Ptychoceraspseudogaultinum shows exactly similar ornament on the penultimate Y okoyama , 1890, p. 181, pi. 20, figs. 1-3, by original shaft, the ornament weakening around the curved sec­ designation. tor before rejuvenating. A marked constriction sepa­ Polyptychoceras (Polyptychoceras) rates the ribbed portion of the final shaft from the es­ obliquecostatum (Schlüter , 1872) sentially smooth section that precedes the adult aper­ (PI. 2, Figs. 1 -3, 7-9). ture, which is prolonged into a blunt ventral rostrum.

1872 Hamites obliquecostatus Schlüter , p. 187, pl. 29, Discussion These specimens confirm the ge­ fig. 6. neric identity of Schlüter ’s Hamites obliquecostatus. 1876 Hamites Berkelis Schlüter , p. 45. As we have noted elsewhere (1984, p. 169), the non 1890 Ptychoceras pseudogaultinum Y okoyama , pl. 20, paralectotype of P. (P.) pseudogaultinum (Y okoyama , fig. 3 only. 1890) (pl. 2, fig. 3) from the Santonian of Japan that non 1953 Polyptychoceras sp. juv. ind., Spath, p. 18, pl. 7, fig. 5. was referred to the present species by W iedmann (1962) ? 1962 P. (Polyptychoceras) obliquecostatum (Schlü ­ has much more prorsiradiate ribs, and widely separated ter ); Wiedmann , p. 85, pl. 10, fig. 6. shafts. The juvenile Polyptychoceras from Antarctica 1965 P. (?) obliquecostatum (Schlüter ); Howarth, p. figured by Spath (1953, p. 18, pl. 7, fig. 5) was also 385. referred to P. (P) obliquecostatum by W iedmann 1984 Polyptychoceras (Polyptychoceras) obliquecos­ (1962); we prefer Spath ’s more cautious assignation. tatum Schlüter , 1872); Kennedy & Summesber ­ ger , p. 169. The specimen from the Santonian of Spain described 1984 Polyptychoceras (Polyptychoceras ) sp.; Kennedy by Wiedmann (1962, p. 185, pl. 10, fig. 6) cannot be & Summesberger , p. 169, pl. 10, figs. 2, 3. afforded even generic assignation with any confidence Type: Holotype, by monotypy, is the original of in our view. Schlüter , 1872, p. 187, pl. 29, fig. 6, from the Late Occurrence Late Campanian of the Münster Campanian near Coesfeld, Westphalia, Germany. Basin, Germany and the Gschliefgraben, Austria. Material four specimens; NHMW/1998z/29/13- Polyptychoceratinaeindet. 16. (Pl. 3, Fig. 6) Description We have four specimens in addi­ tion to the two described in 1984. Of the new material, Material a single specimen, SK/1998/1. three are interpreted as macroconchs (e.g. Pl. 2, figs. 7- Description partly preserved initial whorl with 9) and one as a microconch (Pl. 2, figs. 1-3). NHMW/ a portion of the succeeding straight shaft. The diam­ 1998z/29/15 (Pl. 2, fig. 9) is a 101 mm long, partially eter of the whorl is about 23 mm. The preserved length septate shaft with a maximum preserved whorl height of the shaft is 33 mm including the preceding part of the coil, Wh is 9,8 mm at the preserved end of the frag­ of 13.4 mm. The whorl section is compressed oval, ment. The whorl seems to have been circular, deformed with a whorl breadth to height ratio of 0.79, the degree post mortem into an ellipse. The original shape might of compression probably modified by post-mortem have deviated from planispiral coiling, and been some­ crushing. The rib index is 8-9, the ribs weak and trans­ what helicoidal. The whorls are preserved as a rusty verse on the dorsum, strengthening across the dorsola­ film only, with a very small fragment of internal mould, teral margin, straight, feebly prorsiradiate, low, rounded which is smooth. Ornament may have been lost during and of approximately the same width as the interspaces fossilisation. Visible ribbing starts on the first quarter on both flank and dorsum, where they are transverse. of the coil and extends over the whole preserved length NHMW/1998z/29/16 (not figured) consists of two of the shaft. Ribbing is dense (17/cm) slightly subparallel shafts joined by a narrowly curved section rursiradiate, convex and circular. There are no constric­ of shell, the shafts closely spaced and nearly touching tions visible and no tubercles. at the apertural end of the larger shaft. NHMW/1998z/ Discussion This initial stage of a heteromorph 29/14 (Pl. 2, figs. 7, 8) is essentially identical, but lacks ammonite can be separated from fragments of equal the penultimate shaft. Rib direction shifts from feebly size of Polyptychoceras (P) obliquecostatum (Schlü - prorsiradiate to recti- to feebly rursiradiate around the ter) by their parallel shafts and constrictions. We can­ curved sector before reverting to feebly prorsiradiate. not tell if it belongs to one of the other taxa described The final shaft is very short, and the final sector prior from the Gschliefgraben. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

30 Beitr. Paläont. 24, Wien 1999

Occurrence: Late Campanian of the Gschlief- Pseudoxybeloceras (Schlueterella) sp. ? graben. Pseudoxybeloceras (Parasolenoceras) wernickei (Wollemann ) Aptychus sp. indet. Pseudoxybeloceras (Parasolenoceras) interruptum (PI. 3, Fig. 2) (Schlüter ) Material A single specimen, NHMW/1998z/ Pseudoxybeloceras (Parasolenoceras) phaleratum 41/1. (Griepenkerl ) Description: This double winged lower jaw of Polyptychoceras (P) obliquecostatum (Schlüter ) an ammonite is badly corroded, as a result of which Polyptychoceras (P.) cf. pseudogaultinum (Y okoyama ) the black substance of the surface is only partially pre­ Polyptychoceratinae indet. served and is of coaly lustreless condition. The two Baculites sp. 1 wings are displaced along the symphysis. The rostrum Baculites sp. 2 is somewhat damaged; the wings show concentric un­ Baculites 3 dulations. Trachyscaphites pulcherrimus (Roemer ) Measurements: length 23,45 mm, width 20,4 mm. Accompanying fauna and flora: Discussion The specimen cannot be assigned to a specific ammonite taxon. Plankton: Coccolithophores (Prey 1983, p.101, W ag - REICH, 1999). The Late Campanian ammonite assemblage of the Planktonic foraminifera (Prey 1983, p.101) Gschliefgraben: One fragment of driftwood, no other plant remains. Ammonites (250 specimens, 32 taxa) comprising 18 Nekton: Nautiloidea (rare, one taxon) heteromorph species, 14 non heteromorph ones are the Ammonites (abundant; diverse with prevailing Pachy­ dominant elements in the Gschliefgraben fauna. As discus and heteromorphs) Belemnites (rare, a single taxon; Christensen 1998) already stated by Kennedy & Summesberger (1984) Fish teeth (Prey 1983, p. 101) the fauna is comparable to the Late Campanian faunas Benthos: Sponges (fairly common) of northwestern Europe. Genera such as Phylloceras, Corals (extremely rare, three small solitary corals) Tetragonites, Anapachydiscus are more widely occur­ Brachiopods (? a single disciniscid) ring also in the Tethyan realm or as with Polypty- Gastropods (very rare; four specimens, three taxa; one choceras cf. pseudogaultinum, in circumpacific re­ Pleurotomariidae gen. et sp. indet.(det. H.A. Koll - gions. mann ), Inoceramids (large and common; some articulated; Faunal list Tröger, Summesberger & Skoumal , 1999) Phylloceras (Hypophylloceras) sp. Ostracods (Prey 1983, p. 101) Saghalinites cf. cala (Forbes ) Cirripedia (a single specimen; Fraaye & Summesber - Tetragonites cf. obscurus (Schlüter ) ger , 1999) Gaudryceras jukesi (Sharpe ) Other Crustacea (two specimens, two taxa; Fraaye & Desmophyllites larteti (Seunes ) Summesberger , 1999) Kitchinites sp. Regular echinoids (one specimen) Puzosiinae gen. et sp. indet. Crinoids (rare skeletal elements only) Hauericeras fayoli De Grossouvre Infauna: Bivalves, other than inoceramids (small and Yokoyamaoceras? sp. very rare, 5 specimens; Pholadomya cf. cordata Tate, Pachydiscus (Pachydiscus) haldemsis (Schlüter ) Microchlamys wisniowskii Pasternak ; Cuspidaria Pachy discus (Pachydiscus) perfidus De Grossouvre acutissima (J. M üller ); Cardiidae indet.; identifica­ Pachydiscus (Pachydiscus) cf. subrobustus Seunes tions by A.V. Dhondt ). Anapachydiscus arialoorensis (Stoliczka ) Burro wers: Hoplitoplacenticeras (Hoplitoplacenticeras) preyi sp. Irregular echinoids ( Echinocorys , Micraster and oth­ nov. ers; very common; Jagt , 1999) Bostrychoceras polyplocum (Roemer ) Ichnofossils (rather common in the field). Nostoceras (Nostoceras) sp. Didymoceras donezianum (Mihailov ) Palaeoecology Didymoceras binodosum (Kennedy & Cobban ) The fauna is dominated by a nektonic population of Neoglyptoxoceras cf. retrorsum (Schlüter ) cephalopods. The diversity of the benthic fauna is low. Neancyloceras bipunctatum (Schlüter , 1872) Pseudoxybeloceras (Schlueterella) pseudoarmatum The plankton is extensively represented by foraminif­ (Schlüter ) era and coccolithophores which probably lived close ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

Kennedy , WJ. & Summesberger , H., New Late Campanian Ammonites ... 31 to the surface. The soft substratum of the seafloor sup­ Acknowledgements ported an abundant population of burrowing echinoids We are deeply indebted to Mag. Herbert H ütter , who and other, unidentified groups. There are no hardground gave us access to his outstanding collections and fi­ settling communities present. The only cirripede (pi. 3, nally presented his Gschliefgraben collection to the fig. 5) was found within the body chamber of a large Museum of Natural History in Vienna. Dr. Peter heteromorph ammonite. There is no evidence for shal­ Skoumal donated the specimen figured in pi. 2, fig. 5 low water nor for tropical or subtropical conditions. to the NHMW collection. Mr. Ferdinand E stermann Temperate conditions and a relatively deep continen­ (Pinsdorf near Gmunden) gave us access to his collec­ tal slope environment are indicated. tions, and presented the Aptychus figured in pi. 3. fig. 6. We thank Dr. Gerhard N iedermayr and Dr. Franz Sedimentology Brandstätter from the Dept, of Mineralogy and Pe­ trology of the NHMW for a sedimentological analy­ Pale greenish-grey to yellowish fine sandstones with sis. Dr. Annie V. Dhondt (Brussels) identified the non calcitic cement alternate with shales with high carbon­ inoceramid bivalves, Dr. Heinz A. Kollmann (NHMW, ate and rather low clay mineral content. They were Vienna) the gastropods. We thank the staff of the Mu­ deposited under offshore conditions. Post-sedimentary seum of Natural History, Vienna and of the University transportation by slumping or suspension turbidites Museum of Natural History, Oxford for technical as­ cannot be demonstrated. sistance. We also thank Prof. Jake M. H ancock (Lon­ don) and Mr. John Jagt (Maastricht) for carefully re­ Stratigraphic conclusions viewing the paper. The ammonite fauna of the Gschliefgraben was inter­ preted as Late Campanian Bostrychoceras polyplocum References Zone by Kennedy & Summesberger (1984), the index species occurring abundantly. Hauericeras fayoli De A rkell , W.J., 1950. A classification of the Jurassic ammonites. — J. Paleont. 24: 354-364. Tulsa. Grossouvre and Pachydiscus (Pachydiscus) perfidus A tabekian , A.A. & Khakimov , F.K., 1976. Campanian and De Grossouvre are also recorded from the highest Maastrichtian ammonites in Central Asia. — Akad. Campanian Nostoceras hyatti Zone (H ancock & Nauk Tadzh. SSR, Inst, geol., 146 pp, 13 pis., Kennedy , 1993), whereas Pseudoxybeloceras (Para- Dushanbe (in Russian). solenoceras) phaleratum is the index species of the Blaszkiewicz , A., 1980. Campanian and Maastrichtian lower Late Campanian Phaleratum Zone of the Vistula ammonites of the Middle Vistula Valley, Poland: a valley (Blaszkiewicz 1980). As the total range is still stratigraphic-paleontological study. — Prace Inst. not of many of these taxa known, we follow Kennedy Geol. 92:1-63, 56 pis., Warszawa. & Summesberger (1984) in interpreting the bulk of the Brinkmann , R., 1935. Die Ammoniten der Gosau und des fauna as from the Late Campanian Polyplocum Zone. Flysch in den nördlichen Ostalpen. — Mitt. geol. Staatsinst. Hamburg, 15:1-14, Hamburg. The highest Late Campanian Hyatti Zone is also rep­ resented and now confirmed by nannoplankton ages. Christensen , W.K., 1998. Upper Campanian Belemnitella from Austria. — Beitr. Paläont., 22:13-21,1 pi., Wien. Nannoplankton samples taken from Trachyscaphites Coquand , H., 1859. Synopsis des animaux et des végétaux pulcherrimus indicate nannoplankton zones CC 20- fossiles observés dans la formation crétacée du Sud- 21a-22c; from Bostrychoceras polyplocum CC22b; Ouest de la France. — Bull. Soc. géol. Fr., 2)16:945- from Hauericeras fayoli CC22c (Wagreich , 1999). 1023, Paris. This coincides with the known range of the ammonites. Diener , C., 1925. Ammonoidea neocretacea. — Fossilium A Late Campanian age is also confirmed by echinoids Catalogus (1: Animalia), 29:244 pp., Berlin. (Jagt , 1999) and inoceramids (Troger et al., 1999). D ouvillé , H., 1890. Sur la classification des cératites de la Craie. — Bull. Soc. géol. Fr. (3)18:275-292. Paris. Palaeobiogeography Emerson , B.L., Emerson , J.H., A kers , R.E. & A kers , T.J., 1994. Texas Cretaceous ammonites and nautiloids. Occurrence of belemnites and Trachyscaphites make — Texas Paleontology Series, 5:439 pp. it likely that the Gschliefgraben fauna belongs to the F raaye , R.H.B. & Summesberger , H., 1999. New Northern Temperate Realm. Heteromorphs seem to be crustacean records from the Late Campanian of absent in contemporaneous Tethyan sediments of the Austria. — Beitr. Paläont., 24:1-6, 1 Text-fig., Wien. Gosau Group (S ummesberger et al., 1999) with Gill, T., 1871. Arrangement of the Families of Mollusks. pachydiscids as dominating element of the fauna. — Smithson. Mise. Colins., 227: xvi + 49 pp. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

32 Beitr. Paläont. 24, Wien 1999

G rossouvre , A. de, 1894. Recherches sur la Craie tion in Central Texas. — Acta Geol. Polonica, supérieure, 2. Paléontologie. Les ammonites de la 49(l):67-80, Warsaw. craie supérieure. — Mém. Serv. Carte géol. dét. Kennedy , W.J. & H ancock , J.M., 1993. Upper Maast- France, 264 pp., 39 pis., Paris. richtrian ammonites from the Marnes de Nay between Hancock , J.M. & Kennedy , W.J., 1993. The high Cre­ Gan and Rébénacq (Pyrénées-Atlantiques), France. taceous ammonite fauna from Tercis, Landes, France. — Géobios, 26:575-594, 7 pis., Lyon. — Bull. Inst. Sei. Nat. Belgique, 63:149-209,20 pis., Kennedy , W.J. & H enderson , R.A., 1992. Non-hetero- Brussel. morph ammonites from the Upper Maastrichtian of Henderson , R.A., 1970. Ammonoidea from the Mata Pondicherry, South India.— Palaeontology, 35:381- Series (Santonian-Maastrichtian) of New Zealand. — 442, 18 pis., London. Spec. Pap. Palaeont., 6:82 pp., 15 pis., London. Kennedy , W.J. & Kaplan , U., 1997. Ammoniten aus dem H owarth , M.K., 1965. Cretaceous ammonites and Campan des Stemweder Berges, Dämmer Oberkrei­ nautiloids from Angola. — Bull. Brit. Mus. (Nat. demulde, NW-Deutschland. — Geol. Paläont. West­ Hist.) (Geol.), 10:335-412, 13 pis., London. falens, 40:31-245, 82 pis., Münster. Hyatt , A., 1889. Genesis of the Arietidae. — Smithson. Kennedy , W.J. & Summesberger , H., 1984. Upper Cam­ Contrib. Knowl., 673: xi +239 pp., 14 pis., London. panian ammonites from the Gschliefgraben (Ultra­ helvetic, Upper Austria). — Beitr. Paläont. Öst., Hyatt , A., 1894. Phylogeny of an Acquired Characteristic. — Proc. Am. Phil. Soc., 32:349-647, pis. 1-14. 11:149-206, pi. 14, Wien. Klinger , H.C., 1982. Revision of Ancyloceras bi- Hyatt , A., 1900. Cephalopoda, pp. 502-604 [in:] Zittel, K.A. von 1896-1900, Textbook of Palaeontology, punctatum Schlüter , 1872 (Cephalopoda, Ammo­ transi. Eastman , C.R., London - New York (Mac­ noidea) and discussion of the validity, phylogeny and millan). limits of the genus Neancyloceras Spath , 1926. — Ann. S. Afr. Mus., 90:219-239., Cape Town. Jagt , J.W.M., 1999. Late Campanian echinoids and crinoids from the Gschliefgraben (Ultrahelvetic, Kossmat , F, 1895-1898. Untersuchungen über die Austria). — Beitr. Paläont., 24:7-22, Wien. Südindische Kreideformation. — Beitr. Paläont. Öst. Jimbo, K., 1894. Beiträge zur Kenntniss der Fauna der Ung. Orients, 9: (1895):97-203 (1-107), pis. 15-25 Kreideformation von Hokkaido. — Paläont. Abh. (1-11); 11 (1897a):l-46 (108-153), pis. 1-8(12-19); (N.S.) 2:47-194, pis. 17-25, Jena. 11 (1898):89—152 (154-217), pis. 14-19 (20-25), Wien. Kaplan , U., Kennedy , W.J. & Ernst , G., 1996. Strati­ graphie und Ammonitenfaunen des Campan im süd­ Larson , N.L., Jorgensen , S.D., Farrar , R.A. & Larson , östlichen Münsterland. — Geol. Paläont. Westfalen, P.L., 1997. Ammonites and other cephalopods of the 43:133 pp., 41 pis., Münster. Pierre Seaway. — xi + 148 pp. (Geoscience Press), K ennedy , W.J., 1986. Campanian and Maastrichtian Tucson. ammonites from northern Aquitaine, France. — Spec. Macellari , C., 1986. Late Campanian-Maastrichtian Pap. Palaeont., 36:145 pp., 23 pis., London. ammonite fauna from Seymour Island (Antarctic K ennedy , W.J., 1993. Campanian and Maastrichtian Peninsula). — Mem. Paleont. Soc., 18:55 pp., Tulsa. ammonites from the Mons Basin (Belgium). — Bull. Maeda , H., 1993. Dimorphism of Late Cretaceous false- Inst. R. Sei. Nat. Belgique, 63:99-131, 7 pis., puzosiine ammonites, Yokoyamaoceras Wright & Brussels. Matsumoto , 1954, and Neopuzosia Matsumoto , Kennedy , W.J., B ilotte, M. & M elchior , C., 1995. 1954. — Trans. Proc. Palaeont. Soc. Japan, n.s. Ammonite faunas, biostratigraphy and sequence 169:97-128., Tokyo. stratigraphy of the Coniacian-Santonian of the Matsumoto , T., 1938. A biostratigraphic study on the Corbières. — Bull. Centres Rech. Explor. Prod. Elf- Cretaceous deposits of the Naibuchi Valley, South Aquitaine, 19:377-499, 30 pis., Pau. Karahuto. — Proc. Imp. Acad. Japan, 14:190-194, K ennedy , W.J. & Cobban , W.A., 1993. Campanian Tokyo. ammonites from the Annona Chalk near Yancy, Matsumoto , T., 1955. The bituberculate pachydiscids Arkansas. — J. Paleont., 67 83-97, Tulsa. from Hokkaido and Saghalien. — Mem. Fac. Sei. Kennedy , W.J. & Cobban , W.A., 1993. Upper Campanian Kyushu Univ., Ser. D (Geol.). 5/3:153-184, pis. 31- ammonites from the Ozan-Annona Formation 37, Fukuoka. boundary in Southwestern Arkansas. — Bull. geol. Matsumoto , T., 1988. A monograph of the Puzosiidae Soc. Denmark, 40:115-148, 8 pis., Copenhagen. (Ammonoidea) from the Cretaceous of Hokkaido. — Kennedy , W.J. & Cobban , W.A., 1994. Ammonite Fauna Palaeont. Soc. Japan Spec. Pap., 30:1-179, Tokyo. from the Wenonah Formation (Upper Cretaceous) of Matsumoto , T., 1991. The mid-Cretaceous ammonites of New Jersey. — J. Paleont., 68(1):95—110, Tulsa. the family Kossmaticeratidae from Japan, Part 1, atsumoto Kennedy , W.J. & Cobban , W.A., 1999. Campanian (Late General remarks, p. 3-20. [in:] T. M and Cretaceous) ammonites from the Bergstrom Forma­ others. The mid-Cretaceous ammonites of the family ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

Kennedy , WJ. & Summesberger , H., New Late Campanian Ammonites 33

Kossmaticeratidae from Japan. — Palaeont. Soc. Spath , L.F., 1921. On Upper Cretaceous Ammonoidea Japan. Spec. Pap., 33:143 p., Tokyo. from Pondoland. — Annls. Durban Mus., 3:39-56, Meek, F.B., 1861. Description of new Cretaceous Van­ pis. 6,7, Durban. couver and Sucia Islands. — Proc. Acad. Nat. Sei. Spath , L.F., 1922. On the Senonian ammonite fauna of Philadelphia, 10:314-318, Philadelphia. Pondoland. — Trans. Roy. Soc. S. Aff., 10:113-147, Meek, F.B. & Hayden , F.V., 1856. Descriptions of new pis. 5-9, Cape Town. species of Gastropoda and Cephalopoda from the Spath , L.F., 1926a. On new ammonites from the English Cretaceous formation of Nebraska Territory. — Proc. Chalk. — Geol. Mag., 63:77-83, table, London. Acad. Nat. Sei. Philadelphia, 8 (for 1856):70-72, Spath , L.F., 1929. Corrections of cephalopod nomen­ Philadelphia. clature. — Naturalist, 871:269-271. Mojsisovics , E.v. & Schlönbach , U., 1868. Das Verhalten Spath , L.F., 1953. The Upper Cretaceous Cephalopod der Flyschzone zum Nordrande der Kalkalpen fauna of Grahamland. — Sei. Rep. Br. Antarct. Surv., zwischen dem Traun- und dem Laudach-See bei 3:1-60, pis. 1-13, London. Gmunden. — Verh. G.R.A., 1868 (10):212-216, Stinnesbeck , W., 1986. Zu den faunistischen und pal­ Wien. ökologischen Verhältnissen in der Quiriquina Forma­ N aidin , D.P., 1974. Ammonoidea. Pp. 158—195, pis. 53— tion (Maastrichtium) Zentral-Chile. — Palaeonto­ 70 [in:] Krymogolts , G. Ja. (Ed.). Atlas of Upper graphica A, 194:99-237, pis. 1-15, Stuttgart. Cretaceous Fauna of Donbass. „NEDRA“, Moscow Suess , E., 1866. Über Ammoniten. — Sber. Akad. Wiss. [in Russian], Moscow. Math. Nat. Cl., 52:19 pp., Wien. N owak , J., 1913. Untersuchungen über die Cephalopoden Summesberger , H., 1979. Eine obersantone Ammoni- der oberen Kreide in Polen. III Teil. — Bull. Acad. tenfauna aus dem Becken von Gosau (Oberöster­ Sei. Cracovie. CI. Sei. Math. Nat., Sér.B, Sei. Nat., reich). — Ann. Naturhist. Mus. Wien, 82:109-176, 1913: 335-415, pis. 40-45, Cracow. 15 pis, Wien. Paulcke , W., 1907. Die Cephalopoden der oberen Kreide SzÄsz, L., 1982. Les Ammonites hétéromorphes cam- Südpatagoniens. — Ber. naturw. Ges. Freiburg i.B., paniennes des Carpates méridionales (Roumanie). — 15:167-248, pis. 10-19, Freiburg i.B. Dari de Seama sedint.: Paléontologie, 66 [for 1979]: Prey , S., 1951. Helvetikum und Flysch. [in:] Geologischer 45-56, pis. 1-4, Bucharest. Führer zu den Exkursionen. — Verh. Geol. B.A., Tröger, K.-A., Summesberger , H. & Skoumal , R, 1999. Sonderheft A: 38-48, pis. 7, 8, Wien. Inoceramidae from the Campanian (Upper Creta­ Prey , S., 1983. Das Ultrahelvetikum-Fenster des Gschlief- ceous) of the Gschliefgraben (Ultrahelvetic; Austria). grabens südsüdöstlich von Gmunden (Oberöster­ — Beitr. Paläont., 24:41-61, Wien. reich). — Jb. Geol. B.A., 126/1:95-127,4 Text-figs., Usher , J.L., 1952. Ammonite faunas of the Upper 1 geol. map, Wien. Cretaceous Rocks of Vancouver Island, British Roemer , A., 1840-1841. Die Versteinerungen des nord­ Columbia. — Bull. Geol. Surv. Canada, 21:1-182, deutschen Kreidegebirges, — 145 pp., 48 pis. (1^48, 31 pis, Ottawa. pis. 1-7,1840; 49-145, pis. 8-16,1841) (Hahn’sche Wagreich , M., 1999. Calcareous nannofossil assem­ Hofbuchhandlung), Hannover. blages from the Gschliefgraben near Gmunden (Ult­ Schlüter , C., 1867. Beitrag zur Kenntniss der jüngsten rahelvetic, Austria). — Beitr. Paläont., 24: 63-67, Ammoneen Norddeutschlands. — 36 pp., 6 pis., Wien. Bonn. Ward , P.D. & Kennedy , W.J., 1993. Maastrichtian Schlüter , C., 1871-1876. Cephalopoden der oberen ammonites from the Biscay Region (France, Spain). deutschen Kreide. — Palaeontographica, 21:1-24, — Paleont. Soc. Mem., 34, 58 p., Tulsa. pis. 1-8(1871); 21:25-120, pis. 9-35 (1872); 24:1- Wiedmann , J., 1962. Unterkreide-Ammoniten von Mal­ 144 (121-264) + x, pis. 36-55 (1876), Cassel. lorca. 1. Lieferung: Lytoceratina, Aptychi.— Abh. Seunes , J., 1892. Contribution à l’étude des Céphalopodes Math.-Naturw. Kl. Akad. Wiss. Lit. Mainz, 1962 du Crétacé Supérieur de France. I. Ammonites du (1): 1-148, 10 pis, Mainz. Calcaire à Baculites du Cotentin (Suite). II. Ammo­ Wiedmann , J., 1966. Stammesgeschichte und System nites du Campanien de la région sous-pyrénéenne. den posttriadischen Ammonoideen; ein Überblick. — Départment de Landes. — Mém. Soc. géol. Fr. N. Jb. Geol. Paläont. Abh., 125:49-79, pis. 2; 127, Paléont., 2: Mém. 2:8-22, pis. 12-15 (3-6), Paris. 13-81, pis. 3-6, Stuttgart. Sowerby , J., 1812-1822. The Mineral Conchology of Wright , C.W., 1996. Cretaceous Ammonoidea [in:] Great Britain. 1, pis. 1-9 (1812), pis. 10-44 (1813), Kaesler , R.L. (ed.), Treatise on Invertebrate Pale­ pis. 45-78 (1814), pis. 9-102 (1815); 2: pis. 103-14 (1815), pis. 115-50 (1816), pis. 151-86 (1817), pis. ontology, Part L., Mollusca 4, revised; 4: 362 pp., 87-203 (1818); 3: pis. 204-21 (1818), pis. 222-53 216 figs., 2 tabl., Boulder, Lawrence. (1819), pis. 254-71 (1820), pis. 272-306 (1821); 4: Wright , C.W. & Matsumoto , T., 1954. Some doubtful pis. 307-18 (1821), pis. 319-83 (1822), London. Cretaceous ammonite genera from Japan and ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

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Saghalien. — Mem. Fac. Sei. Kyushu Univ. (D), Y oung , K., 1963. Upper Cretaceous ammonites from the Geol., 4:107-134, pls. 7, 8, Fukuoka. Gulf Coast of the United States. — Univ. Tex. Bull., Y abe , H., 1927. Cretaceous stratigraphy of the Japanese 6304: ix + 373, 82 pls., Austin. Islands. — Sei. Rep. Tohoku Imp. Univ., (2)11:27- Zittel, K.A.v., 1884. Handbuch der Paläontologie, 1, Abt. 100, pis. 3-9, Sendai. 2; Lief. 3, Cephalopoda, pp. 329-522 (R. Olden­ Y okoyama , M., 1890. Versteinerungen aus der ja­ bourg), München & Leipzig. panischen Kreide. — Palaeontographica, 36:159- Zittel, K.A. v ., 1895. Grundzüge der Paläontologie 202, pls. 18-25, Stuttgart. (Paläozoologie). — vii + 972 pp. (R. Oldenbourg), Munich & Leipzig.

Plate 1

Figs. 1-3: Phylloceras sp.; NHMW/1998z/29/l Fig. 4: Kitchinites sp.; NHMW/1998z/29/3 Fig. 5: Kitchinites sp.; NHMW/1998z/29/4

Figs. 6-9: Desmophyllites larteti (Seunes 1892); NHMW/1998z/29/2 All from the Upper Campanian of the Gschliefgraben; all figures x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

Kennedy , WJ. & S ummesberger , H., New Late Campanian Ammonites 35

PLATE 1 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

36 Beitr. Paläont. 24, Wien 1999

Plate 2

Figs. 1-3, 7-9: Polyptychoceras (Polyptychoceras) obliquecostatum (Schlüter , 1872); figs. 1-3 is SK 1985/20, figs. 7,8 is NHMW/1998z/29/14, fig. 9 is NHMW/1998z/29/15.

Figs. 4, 5: Bostrychoceraspolyplocum (Roemer , 1841); 4 is NHMW/1998z/29/6; 5 is NHMW/1998z/40/l.

Fig. 6: Neancyloceras bipunctatum (Schlüter , 1872); NHMW/1998z/29/9.

Fig. 10: Pseudoxybeloceras (Schlueterella) pseudoarmatum (Schlüter , 1872), NHMW/1998z/29/10. Fig. 11: Yokoyamaoceras? sp.; NHMW/1998z/29/5.

Figs. 12, 13: Didymoceras binodosum (Kennedy & Cobban , 1993). 12. is RO/1986/3; 13 is NHMW/1998z/29/ 7, 8. All from the Upper Campanian of the Gschliefgraben; all figures x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

Kennedy , W.J. & S ummesberger , H., New Late Campanian Ammonites ... 37

PLATE 2 ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

38 Beitr. Paläont. 24, Wien 1999

Plate 3

Fig. 1, 4: Hoplitoplacenticeras (Hoplitoplacenticeras) preyi sp. nov. Holotype, NHMW/1998z/29/13. Fig. 2: Aptychus sp. indet. Lower jaw of an indeterminate ammonite; NHMW/1998z/41/01; uncoated x 1.

Fig. 3: Pseudoxybeloceras (Schlueterella) pseudoarmatum (Schlüter , 1872), NHMW/1998z/29/12; x 0,43. With a cirripede in the body chamber.

Fig. 5: Pseudoxybeloceras (Schlueterella) pseudoarmatum (Schlüter , 1872), NHMW/1998z/29/ll; x 0,43. Fig. 6: Initial growth stage of Polyptychoceratinae indet.; SK/1998/1; uncoated. All from the Upper Campanian of the Gschliefgraben; The figures 3 and 5 are reduced x 0,43, the remainder are x 1. ©Verein zur Förderung der Paläontologie am Institut für Paläontologie, Geozentrum Wien

Kennedy , W.J. & S ummesberger , H., New Late Campanian Ammonites ... 39

PLATE 3

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