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Mollusca Gastropoda : Seguenziidae from New Caledonia and the Loyalty Islands

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Mollusca Gastropoda : Seguenziidae from New Caledonia and the Loyalty Islands JLTATS DES CAMPAGNES MLJSORSTOM, VOLUME 7 RÉSULTATS DES CAMPAGNES MUSORSTOM. VOLUME 7 - RÉSUL1 3 Mollusca Gastropoda : Seguenziidae from New Caledonia and the Loyalty Islands Bruce A. MARSHALL National Museum of New Zealand P.O. Box 467, Wellington New Zealand ABSTRACT Three subfamilies are recognised : Asthelysinae new sub- Iredale, which is transferred from Trochidae. Asthelys niti- family, Seguenziinae Vcrrill, and Guttulinae Goryachev. Two dula sp. nov. is based on type material from Queensland. Jaw tribes are recognised in Seguenziinae. Fifty five seguenziids plates and lateromarginal radular plates are recorded for the are newly recorded from off New Caledonia and the Loyalty first time in the family. Seguenziid species richness and Islands, of which 50 are new to science. These species are western Pacific biogeography are briefly discussed. referable to 13 genera, including 2 new genera and Anxietas RESUME Mollusca Gastropoda : Seguenziidae de Nouvelle-Calédonie raduia est signalée pour la première fois chez les Seguen- et des îles Loyauté. ziidae. Cinquante des 55 espèces présentes sont décrites comme nouvelles de cette région ; une espèce est décrite du Une remarquable faune de Seguenziidae est décrite des Queensland. Ancistrobasis monodon (Schepman) est transféré étages bathyal et abyssal de la région néo-calédonienne des Calliostomatinae, et Anxietas Iredale des Trochidae, aux (bassin des Loyauté et sud de la Nouvelle-Calédonie). La Seguenziidae. Deux nouveaux genres, Eratasthelys et Halys- richesse spécifique de la famille et son apport à la biogéogra- tina, et une nouvelle sous-famille. Asthelysinae, sont créés ; la phie de l'Ouest Pacifique sont brièvement discutés. L'exis- sous-famille nominale Seguenziinae est divisée en deux tribus. tence de mâchoires et d'une plaque latéromarginale sur la MARSHALL, B. A., 1991. Mollusca Gastropoda : Seguenziidae from New Caledonia and the Loyalty Islands. In :A. CROSNIKR & P. BOUOHKT (eds). Résultats des Campagnes MUSORSTOM, Volume 7. Mém. Mus. natn. Hist. not.. (A), 150 : 41-109. Paris ISBN : 2-85653-180-6. Publié le 20 mars 1991. 42 BRUCE A. MARSHALL INTRODUCTION During the last ten years the deep-sea gastro- Abbreviations and text conventions : pods of the family Seguenziidae have received AMS : Australian Museum, Sydney ; considerable attention from systematists, the BMNH : The Natural History Museum, London ; number of genera increasing from 7 to 20, and D : Diameter; the number of Recent species more than dou- H : Height; bling to 139 with the present contribution. MNHN : Museum national d'Histoire naturelle, Through studies of shell structure (BANDEL, Paris ; 1979; BARSKOV, GOLOVINOVA & GORYACHEV, NMNZ : National Museum of New Zealand, 1980), radula (MARSHALL, 1983 ; QUINN, 1983b) Wellington ; and anatomy (QUINN, 1983b), this formerly NMP : Natal Museum, Pietermaritzburg; enigmatic family is now firmly established as a NSMT : National Science Museum, Tokyo ; member of the Archaeogastropoda. SALVINI- TW : Teleoconch whorls (number) ;. PI.AWEN & HASZPRUNAR (1987) have recently referred the family to a new archaeogastropod UD : Umbilicus/diameter as percentage of suborder, Seguenziina. shell diameter; USNM : National Museum of Natural History, Through the generosity of Philippe BOUCHET, Washington DC ; I now have the privilege of recording by far ZMA : Zoologisch Museum, Amsterdam. the richest seguenziid fauna known, comprising 55 species in 13 genera. The fact that all are new Height precedes diameter in all given dimen- records is testimony not only to the great sions. All shell measurements were taken on the diversity of the fauna occurring off New Caledo- longitudinal axis or at right angles to it. In nia and the Loyalty Islands, but also as to how descriptions of the posterior notch in the outer little is actually known of the deep-sea molluscan lip, " retraction depth " is the depth from the fauna occurring off tropical and subtropical adapical insertion to the back of the notch, while western Pacific islands in general. " protraction depth M is the depth from the back Preserved specimens of several species recorded of the notch to the tip of the forward-swinging living were received too late for inclusion of abapical part of the lip. Unless otherwise stated descriptions and illustrations of their animals these measurements were taken at the lip rim of and radulae. mature specimens. STATION DATA Species taken alive are denoted by asterisks. Station DW08. 20°34' S, I66°54' E, 435 m, Station number prefix CP = chalut á perche 12.8.1985 : Anxietas exigua, Ancistrohasis (beam trawl), DS = drague type Sanders (epiben- monodon, A. tiara, Calliobasis spectrum, C. thic sledge), DW = drague type Waren (rock nepticula, C. merista, Fluxinella polita. dredge). Station CP13. 20°19'S, 167° 18' E, 3 690- 3 740m, 13.8.1985 : Basilissa superba*. BIOCAL campaign stations : N. O. " Jean- Station DS 14. 20°18'S, 167" 18' E, 3 680- Char cot 3 700m, 13.8.1985 : Seguenzia platamodes*, Fluxinella tenera. Station DS04. - 21°16'S, 166°40' E, 2 340 m, Station CP17. — 20"35'S, 167°25'E, 3 680 m, 11.8.1985 : Asthelys depressa, Halystina cale- 14.8.1985 : Basilissa superba. donita*, Seguenzia emmeles. Station CP23. 22°46'S. 166°20'E, 2 040 m, MOLLUSCA GASTROPODA : SEGUENZIIDAE 43 28.8.1985 : Quinnia laetifica*, Seguenzia richeri. Station DW64. - 24°48'S, 168°09'E, 250 m, Station CP 26. 22°40'S, 166°27' E, 1618- 3.9.1985 : Calliobasis festiva. 1 740 m, 28.8.1985 : Carenzia serrata, C. acan- Station DW66. 24-55'S, 168°22'E, 505- thodes, Quinnia limatula, Seguenzia eidalima. 515 m, 3.9.1985 : Ancistrobasis monodon. Station DW 33. — 23°10' S, 167°10' E, 675-680 m, Station DW 70. 23°25' S, 167°53' E, 965 m, 29.8.1985 : Aneistrobasis monodon, Fluxinella 4.9.1985 : Fluxinella polita, F. asceta, F. runci- asceta*. nata, Seguenzia eutyches. Station DW 36. 23°09'S, 167°11'E, 650- Station CP72. 22°10'S, 167°33'E, 2 100- 680 m, 29.8.1985 : Fluxinella asceta. 2 110 m, 4.9.1985 : Fluxinella brychia, Carenzia Station DW 38. 23°00' S, 167°15'E, 360 m, ornata, Quinnia patula, Seguenzia emmeles, S. 30.8.1985 : Calliobasis phimosa, C. spectrum. levii. Station DW41. 22°45'S, 167° 12' E, 380- Station CP 75. — 22°19' S, 167°23' E, 825-860 m, 410 m, 30.8.1985 : Calliobasis spectrum*. 4.9.1985 : Seguenzia chelina, S. chariessa*. Station DW44. 22°4T S, 167° 14' E, 440- Station DW 77. 22°15'S, 167° 15' E, 440 m, 450 m, 30.8.1985 : Aneistrobasis tiara*, A. 5.9.1985 : Ancistrobasis bouchetì*, Fluxinella caledonica*, Calliobasis phimosa*, Fluxinella membranacea. polita*. Station DW 79. 20°40' S, 166°52' E, 1 320- Station DW46. 22°53'S, 167° 17' E, 570- 1 380 m, 5.9.1985 : Asthelys nitidula, Fluxinella 610 m, 30.8.1985 : Aneistrobasis tiara*, Fluxi- runcinata, F. euphanes, Carenzia nitens, C. nella polita*, F. asceta*. acanthodes, Halystina farinata, Quinnia lima- Station DW48. - - 23°00'S, 167°29'E, 775 m, tula, Seguenzia wareni, S. engonia, S. praeceps, 31.8.1985 : Eratasthelys corona, Aneistrobasis S. chariessa. tiara, A. caledonica, A. adonis, Basilissopsis Station DW80. 20°32' S, 166°48'E, 900- charcoti, Fluxinella asceta, F. ruminata. 980 m, 5.9.1985 : Fluxinella asceta, F. runci- Station DW49. 23°03'S, 167°32'E, 825- nata, F. megalomphala, Seguenzia chelina, S. 830 m, 31.8.1985 : Aneistrobasis adonis*, Fluxi- metivieri, S. matara, S. chariessa, S. stegastris. nella ruminata. Station DW 83. - 20°35'S, 166°54'E, 460 m, Station DW51. 23°05'S, 167°45'E, 680- 6.9.1985 : Ancistrobasis tiara. 700 m, 31.8.1985 : Aneistrobasis scitula*, Fluxi- Station DS98. 21°24'S, 166°30'E, 2 365- nella asceta*, F. stirophora. 2 470 m, 7.9.1985 : Carenzia ornata, Quinnia Station DW 53. — 23°09'S, 167°43' E, 975- patula, Halystina caledonica, Seguenzia cha- 1 005 m, 1.7.1985 : Fluxinella polita, F. asceta, riessa. F. runcinata. Station DW 56. 23°35'S, 167°12'E, 695- Station DW 106. - 21°36'S, 166°29' E, 625- 705 m, 1.9.1985 : Fluxinella stirophora. 650 m, 8.9.1985 : Fluxinella xysila. Station CP 57. — 23Ü 44'S, 166°58' E, 1490- R. V. "Vauban" 1978-79. 1 620 m, 1.9.1985 : Asthelys nitidula*, Hadro- conus grandiosus, Carenzia nitens, C. serrata. Station 40. - 22°30'S, 166°24'E, 250-350 m, Station DS 59. — 23°56'S, 166°41'E, 2 650 m, 7.6.1979 : Anxietas inspirata, Fluxinella mem- 2.9.1985 : Asthelys semiplicata, Fluxinella bry- branacea, Halystina vaubani, Seguenzia iota. chia, Basilissa superba, Carenzia ornata, Seguen- zia chariessa, S. emmeles, S. levii. 44 BRUCE A. MARSHALL SYSTEMATIC ACCOUNT Subclass PRO S O B R A N C H I A Milne Edwards, 1848 Order ARCHAEOGASTROPODA Thiele, 1925 Suborder SEGUENZIINA Salvini-Plawen & Haszprunar, 1987 Superfamily SEGUENZIOIDEA Verrill, 1884 Family SEGUENZIIDAE Verrill, 1884 Seguenzidae Verrill, 1884 : 186 (emended). nal anatomy ; three of these groups are here allocated subfamilial status, the nominal sub- The family Seguenziidae is divisible into four family containing two tribes, and are defined as rather well .defined groups characterised by dis- follows : tinctive combinations of shell, radula and exter- Subfamily ASTHELYSINAE nov. TYPE GENUS. - Asthelys Quinn, 1987. and/or minute punctations on subsequent whorls. Axials seldom present. Posterior notch shallow, INCLUDED GENERA. — Anxietas Iredale, 1917 ; Thelyssina Marshall, 1983; Asthelys Quinn, 1987; no tooth on inner lip. Snout tip blunt. Central Er at asthelys gen. nov. and lateral teeth stout, rigid, outer marginals each with fine cusps that extend around tip of DIAGNOSTIC CHARACTERS. — Teleoconch with cutting area. anastomosing dendritic threads on first whorl Subfamily SEGUENZIINAE Verrill, 1884 TYPE GENUS. - Seguenzia Jeffreys, 1876. DIAGNOSTIC CHARACTERS. Teleoconch micro- sculpture usually present, consisting of minute INCLUDED GENERA. — Seguenzia Jeffreys, 1876 ; granulations. Axial sculpture present or absent. Basilissa Watson, 1879 ; Ancistobasis Dall, 1889 ; Posterior notch very shallow to very deep, inner Basilissopsis Dautzenberg & Fischer, 1897 ; The- lip with or without tooth.
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