Insecta: Odonatoptera: Meganisoptera) Based Upon New Species and Redescription of Selected Poorly Known Taxa from Eurasia

Home , Meganeura, Meganisoptera, Odonatoptera

Palaeontographica Abt. A 289 Lfg. 4±6 89±121 Stuttgart, September 2009

Revision of Permo-Carboniferous griffenflies (Insecta: Odonatoptera: Meganisoptera) based upon new species and redescription of selected poorly known taxa from Eurasia

by AndreÂ Nel, GuÈnther Fleck,Romain Garrouste,Georges Gand, Jean Lapeyrie,Seth M. Bybee and Jakub Prokop

With 13 plates and 2 text-figures

Summary

The diagnosis of the Meganisoptera is discussed. Paralogus hispanicus sp. nov. is described from the Late Carboniferous-Early Permian of Spain. Those of the Paralogidae and Kohlwaldiidae and the positions of Oligotypus makowskii are discussed. The nomen nudum `Oligotypus britannicus' is transferred from Paralogidae into the Meganeuridae: Tupinae. Solutotherates is transferred in Kohlwaldiidae sit. nov. An emended diagnosis of Meganeuridae is proposed. The diagnoses of Meganeurinae, Piesbergtupinae, and Tupinae are discussed and some important characters redefined. The description of Meganeura monyi is set forth. Meganeura (?) vischerae Zalessky, 1950, is considered as an Insecta incertae sedis stat. nov. Meganeurula selysii and the genus Meganeurula are transferred into the Meganeurinae sit. nov. The genus Gilsonia Meunier, 1909, and the combination Gilsonia titana Meunier, 1909, are restored in Tupinae. The tupine genus Meganeurina Handlirsch, 1919, is restored for Meganeurina confusa Handlirsch, 1919. The diagnosis of the tupine type genus Tupus Sellards, 1906, is set forth, and a new species Tupus gallicus sp. nov. described from the Middle Permian of LodeÁve. A new diagnosis is given for the genus Arctotypus Martynov, 1932, with the following included species, which are either described or revised: four Russian species Arctotypus sinuatus Martynov, 1932, Arctotypus sylvaensis Martynov, 1940, Arctotypus fortis Zalessky, 1950, Arctotypus giganteus sp. nov., Arctotypus diluculum (Whalley 1980) comb. nov. from UK, and the four species from LodeÁve Arctotypus gallicus sp. nov., Arctotypus merifonsensis sp. nov., Arctotypus intermedius sp. nov., and Arctotypus (?) undetermined species A. The new genus Permotupus gen. nov. is described for two new species Permotupus ollieorum sp. nov. and Permotupus minor sp. nov. from LodeÁve. The new genus Curvitupus gen. nov. is described from Curvitupus ariegensis sp. nov. from LodeÁve, Curvitupus verneti (Laurentiaux-Vieira & Laurentiaux 1963) comb. nov. from France, and two Russian species Curvitupus magnificus (Zalessky 1950) comb. nov. and Curvitupus elongatus (Zalessky 1950) comb. nov. The new genus and species Nannotupus pumilio gen. nov., sp. nov. is described from LodeÁve. ªPalaeotheratesºHandlirsch, 1906, and Ephemerites Geinitz, 1865, are Meganisoptera incertae sedis stat. nov. Solutotherates Brauckmann &Zessin, 1989, is transferred in the family Kohlwaldiidae sit. nov. The LodeÁve griffenfly fauna is the most diverse

Adresses of the authors: AndreÂ Nel, GuÈnther Fleck &Romain Garrouste, CNRS UMR 7205, MuseÂum National d'Histoire Naturelle, CP 50, Entomologie, 45 rue Buffon, F-75005 Paris, France. E-mails: [email protected], [email protected]; G. Fleck, Museu Integrado de Roraima, Av. Brigadeiro Eduardo Gomes, Parque AnauaÂ, 69305-010 Boa Vista, Roraima, Brazil. E-mail: [email protected]; Georges Gand, Laboratoire BiogeÂosciences, UMR-CNRS 5561, UniversiteÂ de Bourgogne, 6 Boulevard Gabriel; 21000 Dijon, France.E-mail: [email protected]; Jean Lapeyrie, Corniche de Fontbonne, F-34700 LodeÁve, France. E-mail: [email protected]; Seth M. Bybee, Entomology and Nematology, University of Florida, Natural Area Drive, 32611, Gainesville, FL, USA. E-mail: [email protected]; Jakub Prokop, Charles University in Prague, Faculty of Science, Department of Zoology, VinicnaÂ 7, CZ-128 44, Praha 2, Czech Republic; E-mail: [email protected]

0375-0442/09/0289/0089 $ 20.70 ° 2009 E. Schweizerbart'sche Verlagsbuchhandlung, D-70176 Stuttgart