28. RUBUS Linnaeus, Sp. P1. 1: 492. 1753. 悬钩子属 Xuan Gou Zi Shu Lu Lingdi (陆玲娣 Lu Ling-Ti); David E
Flora of China 9: 195–285. 2003. 28. RUBUS Linnaeus, Sp. P1. 1: 492. 1753. 悬钩子属 xuan gou zi shu Lu Lingdi (陆玲娣 Lu Ling-ti); David E. Boufford Shrubs or subshrubs, deciduous, rarely evergreen or semievergreen, sometimes perennial creeping dwarf herbs. Stems erect, climbing, arching, or prostrate, glabrous or hairy, usually with prickles or bristles, sometimes with glandular hairs, rarely unarmed. Leaves alternate, petiolate, simple, palmately or pinnately compound, divided or undivided, toothed, glabrous or hairy, sometimes with glandular hairs, bristles, or glands; stipules persistent, ± adnate to petiole basally, undivided or occasionally lobed, persistent or caducous, near base of petiole or at junction of stem and petiole, free, usually dissected, occasionally entire. Flowers bisexual, rarely unisexual and plants dioecious, in cymose panicles, racemes, or corymbs, or several in clusters or solitary. Calyx expanded, some- times with a short, broad tube; sepals persistent, erect or reflexed, (4 or)5(–8). Petals usually 5, rarely more, occasionally absent, white, pink, or red, glabrous or hairy, margin entire, rarely premorse. Stamens numerous, sometimes few, inserted at mouth of hy- panthium; filaments filiform; anthers didymous. Carpels many, rarely few, inserted on convex torus, each carpel becoming a drupelet or drupaceous achene; locule 1; ovules 2, only 1 developing, collateral, pendulous; style filiform, subterminal, glabrous or hairy; stig- ma simple, capitate. Drupelets or drupaceous achenes aggregated on semispherical, conical, or cylindrical torus, forming an aggre- gate fruit, separating from torus and aggregate hollow, or adnate to torus and falling with torus attached at maturity and aggregate solid; seed pendulous, testa membranous; cotyledons plano-convex.
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