.A. BollettinoMalacologico, Roma, 37 (9-12):223-224, 2001 - SocietàItaliana di Malacoloaia

Bernard Landau & Robert Marquet

KEYWORDS: Chi/eutomia,, Pliocene, Zanclean, Mediterranean

ABSTRACT A few shells attributable to Chi/eutomia miranda (Dautzenberg, 1925) (Eulimidae, Prosobranchia, ) have been discovered in che early Pliocene (Zanclean) deposits at Velenn near Estepona, Southern Spain. This is che first fossil record of this in che Meditetranean Neogene.

RIASSUNTO Pochi nicchi atttibuiti a Chileutomiamiranda (Dautzenberg, 1925) (Eulimidae, Prosobranchia, Gastropoda) sono stati rinvenuti in depositi del primo Pliocene (Zancleano) a Velenn, vicino a Estepona (Spagna meridionale). Questo è il primo reperto fossile neogenico di questo genere nel Mare Mediterraneo.

B. LANDAU, Inrernarional Healrh Cenrres, Avenida Infanre de Henrique 7, Areias SaoJooo, 8200 Albufeira, Porrugal. R. MARQUET,Instirur royal des Sciencesnarurelles de Belgique, c/o Constirutiesrraar 50, 2060 Anrwerp, Belgiurn.

INTRODUCTION More recently LoZOUET(1999) considered Auricu/igerina a synonyrn Only in recentyears bave the highly fossiliferousPliocene (Zan- of Chi/mtomia, and we fully agree with this opinion, since alI these dean) depositsaround Velerin, Estepona(Malaga, Spilla) appeared genera display a strong anal notch, absent in Oceanida.Furrherrnore, in the literature (e. g. Vera-Pelaezet al, 1995).These Pliocene depo- the presence of the umbilicus depends on the greater or Iesserdeve- sits, consistof coarsesands (representing out nearshore and/or beach Ioprnent of the columellar Iip and cannot be used as a character to deposits),fine dayey sands(deposited in a relatively deepershelf diff"erentiate between the genera. environrnent)and coarseconglomerates, alI within a half kilometre radius. The conglomerateshost the most diversified macrofossil assemblages;however the most interesting small gastropodsafe found in fine sandyday especial1yat the locality ofVelerin Carrete- ra. Here the preservationof calcareousfossils is exquisite and even fragile speciesrarely found as fossil (such as Aglaja depictaRenier, 1807) occur in such sediments.The presentpaper documentsthe first recordof shellsbelonging to Chileutomiamiranda (Dautzenberg, 1925) in the MediterraneanNeogene. Our recordis basedupon two shells sourcedfrom Velerin Carreteraand two more found within coarsesand deposits outcropping nearbyat Velerin Antena.

SYSTEMATICPALAEONTOLOGY Family Eulimidae Rafinesque, 1815 Genus Chileutomia Tate & Cossmann in Tate, 1898 Synonyms Hoplopteropsis de Morgan, 1915; AuriculiKerina Dautzenberg,1925

Discussionof genus:The genusHoplopteropsis, Mogan, 1915 was synonymisedwith ChileutomiaTate & Cossmannin Tate, 1898 by COSSMANN(1921). WARÉN(1984, 1986) discussedthe close relationshipbetWeen Auriculigerina Dautzenberg, 1925, Oceanidade Folin, 1870 and ChileutomiaTate & Cossmannin Tate, 1898. Accor- ding to this author Chileutomiais restricted to the Tertiary of Australia and differs from the previoustWo in having a deepumbili- cus. Auriculigerina is characterisedby a greater development of thickened scarsfrom earlier positions of the outer lip presentin all PLATE 1: Chi/eutomiamiranda (Dautzenbetg, 1925) Lowet Pliocene, Zanclean; Velenn conglometates, Velenn Estepona,Spain. thesegenera, into wing-like varices.lndeed he suggestedAuriculige- Fig. 1: Fronta! view; x 15.2. Fig. 2: Side view; x 15.2. Fig. 3: Protoconch api, rina miranda could be an aberrant speciesof the genus Oceanida. view; x 45.8. Fig. 4: Protoconch; x 45.4. Fig. 5: Protoconch detail; x 56.8.

22~ ~ Bemard Landau & Rober1 Marquet

1925 -Auriculigerina miranda Dautzenberg, 1925: 7, fig. 5-6. speciesshows distinctly carinatedapical whorls, as in Menan',placed 1926 - Auriculigerinamiranda Dautzenberg: Dautzenberg, 1927. by him in the Rissoacea.Neither the figure nor the description 170, pi. 5, fig 14-17 given by de MORGAN(1915, p. 340, fig. 36) suggest carinated 1927 - Auriculigerinamiranda Dautzenberg: Warén 31, fig. 46-47. whotls and due to the lack of further material, we prefer to keepthe 1928 - Auriculigerinamiranda Dautzenberg: Bouchet & Warén, 440, fig. 1042-1047. ' originai genericdesignation. ACKNOWLEDGEMENTS MATERIALSTUDIED We thank P. Lozouetof the Museum nationald'Histoire natutel- Two specimens from the Velenn sands. two from Velenn Carre- le, Paris and A. Warén, Swedish Museum of Natural History, reca, Estepona, Spain. AlI Zanclian, Lower Pliocene (no forrnation Stockholmfar the loan of photographsand helpful commentson the name). manuscript. We afe grateful to L. Cilis and K. Wouters, lnstitut Dimensions: figured specimen: 6.9x3.9 rom, others: 7.lx2.7 royai desSciences naturelles de Belgique, Brussels,far the useof the rom, 5.Ox2.2 rom, 3.4x3.4 rom. SEM. Description: Shell small, thin, turriculate, antero-posteriorly flattened, seven to eight whorls, height of the largest specimen REFERENCES 6.9rom, width 3.9 rom (height/width ratio 1.76). Protoconch pau- BOUCHET P. & WARÉN, A., 1986. Revision ofthe northeast Atlantic bathyal and cispirai, about one and one half whorls, smooth, first whorl bulbous, Abyssai Aclididae, Eulimidae, Epitoniidae (, Gastropoda). Bolleti- junction with teleoconch not clearly delineated. Teleoconch about no della SocietàMalacologica Italiana, Milano, Suppl. 2: 300-576. six whorls, devoid of sculpture except far very faint, closely set ~ M., 1921.Essais de pa/éocOn&hoIogie compa~, 12. Paris, PressesUniversitai- growth lines. Whorl profile is almost straight with a well-developed res, 348 pp. suturai ramp, giving the shell profile a stepped appearance. Two strong varices afe present on the periphery of each whorl, placed DAUTZENBERG P., 1925. Mollusques nouveaux provenant cles croisières clu Ptince almost in a linear fashion above each other, developing from the scar Albett Jet cle Monaco. Bu/letin ti. l'Imtitut ~nographique ti. Monaco, Mona- of the outer aperturallip. These varices afe detached from the whorl co, 457: 1-12. and most developed adapically, where they join the suture forming a DAUTZENBERGP., 1927. Mo1lusques provenanc des campagnes sciencifiques du Pcin- sinus. The whorl profile is therefore broadest adapically, tapering sli- ce Albett Jer de Monaco dans l'Océan Aclanrique ec dans le golfe de Gasco- ghtly towards the lower suture. Aperture elongated, ovai, with the gne. Résultats deJcampagoo scientifiques .Albert ler. 72: 1-401. outer lip slightly flared abapically. Outer lip thickened; well-develo- ped anal sinus; narrow, clearly delimited parietal callus. In some spe- MORGAN J. de, 1915. Observationssur la sttatigtaphie et la paiéontologiedu Falu- cimens a srnall umbilical cleft is formed between the columella and nien de la Toutaine. Bu//etin de la SociétéG6J/ogique de France, Paris, (4) 15: the last varix on the body whotl. The thickness of the outer lip and 217-241. strength of the umbilical cleft varies between specimens. LozoUET P., 1999. Nouvelles espècesde Gastétopodes(Mollusca: Gastropoda)de REMARKS l'Oligocène et du Miocène inférieUt d'Aquitaine (sud-ouestde la Fmnce). Partie 2. Cossmanniana,Paris, 6 (1-2): 1-68. Auriculigerina miranda is a rare taxon. Originally described from deep water off the Azores and, more recently, recorded also from VERA-PELAEZJ.L. et al., 1995. Estudio preliminar de la malacofaunadel Pliocenod! shallower depths offshore Sicily (BOUCHET & W ARÉN, 1986, p. Estepona(Mlilaga, Espafia).Ikrus, 13 (2): 93-117. 440). Three european Tertiary species bave been described, alI from W ARÉNA., 1984. A generic revision of che fami1y Eulimidae. Journal o/ Mo/luscan French Atlantic deposits under the various generic narnes mentio- Studies,Londtm, supp1. 13: 1-%. ned above. The oldest is Chileutomiapaulinemis Lozouet, 1999, recor- ded from the Late Oligocene, Hoplopteropsispontileviensis de Morgan, 1915, from the Pontilevien, Middle Miocene and the Late Miocene (Redonian) Chileutomia morgani Cossrnann in de Morgan, 1915. Chi- leutomiapaulinemis Lozouet, 1999, differs from C. miranda by having a multispiral protoconch with about one extra whorl; the teleocon- chs of both species afe remarkably similar and characterised by strongly developed varices. Chileutomia pontileviensis (de Morgan, 1915) has a paucispiral, bulbous protoconch (like C. miranda), but differs by having a more elongated, less stepped teleoconch and dif- fers from from both C. paulinensisand C. miranda by having lesser developed varices. Judging from the original figure Chileutomia mor- gani Cossrnann in de Morgan, 1915 has even weaker varices than C. pontileviensisand a larger and more oval aperture than any of the pre- vious species.According to W ARÉN(1984, p. 49) 'The figure of that Lavoro accettato il 24 marzo 2001

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