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Triassic- Jurassic Stratigraphy Of
Triassic- Jurassic Stratigraphy of the <JF C7 JL / Culpfeper and B arbour sville Basins, VirginiaC7 and Maryland/ ll.S. PAPER Triassic-Jurassic Stratigraphy of the Culpeper and Barboursville Basins, Virginia and Maryland By K.Y. LEE and AJ. FROELICH U.S. GEOLOGICAL SURVEY PROFESSIONAL PAPER 1472 A clarification of the Triassic--Jurassic stratigraphic sequences, sedimentation, and depositional environments UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON: 1989 DEPARTMENT OF THE INTERIOR MANUEL LUJAN, Jr., Secretary U.S. GEOLOGICAL SURVEY Dallas L. Peck, Director Any use of trade, product, or firm names in this publication is for descriptive purposes only and does not imply endorsement by the U.S. Government Library of Congress Cataloging in Publication Data Lee, K.Y. Triassic-Jurassic stratigraphy of the Culpeper and Barboursville basins, Virginia and Maryland. (U.S. Geological Survey professional paper ; 1472) Bibliography: p. Supt. of Docs. no. : I 19.16:1472 1. Geology, Stratigraphic Triassic. 2. Geology, Stratigraphic Jurassic. 3. Geology Culpeper Basin (Va. and Md.) 4. Geology Virginia Barboursville Basin. I. Froelich, A.J. (Albert Joseph), 1929- II. Title. III. Series. QE676.L44 1989 551.7'62'09755 87-600318 For sale by the Books and Open-File Reports Section, U.S. Geological Survey, Federal Center, Box 25425, Denver, CO 80225 CONTENTS Page Page Abstract.......................................................................................................... 1 Stratigraphy Continued Introduction... .......................................................................................... -
Eubrontes and Anomoepus Track
Sullivan, R.M. and Lucas, S.G., eds., 2016, Fossil Record 5. New Mexico Museum of Natural History and Science Bulletin 74. 345 EUBRONTES AND ANOMOEPUS TRACK ASSEMBLAGES FROM THE MIDDLE JURASSIC XIASHAXIMIAO FORMATION OF ZIZHONG COUNTY, SICHUAN, CHINA: REVIEW, ICHNOTAXONOMY AND NOTES ON PRESERVED TAIL TRACES LIDA XING1, MARTIN G. LOCKLEY2, GUANGZHAO PENG3, YONG YE3, JIANPING ZHANG1, MASAKI MATSUKAWA4, HENDRIK KLEIN5, RICHARD T. MCCREA6 and W. SCOTT PERSONS IV7 1School of the Earth Sciences and Resources, China University of Geosciences, Beijing 100083, China; -email: [email protected]; 2Dinosaur Trackers Research Group, University of Colorado Denver, P.O. Box 173364, Denver, CO 80217; 3 Zigong Dinosaur Museum, Zigong 643013, Sichuan, China; 4 Department of Environmental Sciences, Tokyo Gakugei University, Koganei, Tokyo 184-8501, Japan; 5 Saurierwelt Paläontologisches Museum Alte Richt 7, D-92318 Neumarkt, Germany; 6 Peace Region Palaeontology Research Centre, Box 1540, Tumbler Ridge, British Columbia V0C 2W0, Canada; 7 Department of Biological Sciences, University of Alberta 11455 Saskatchewan Drive, Edmonton, Alberta T6G 2E9, Canada Abstract—The Nianpanshan dinosaur tracksite, first studied in the 1980s, was designated as the type locality of the monospecific ichnogenus Jinlijingpus, and the source of another tridactyl track, Chuanchengpus, both presumably of theropod affinity. After the site was mapped in 2001, these two ichnotaxa were considered synonyms of Eubrontes and Anomoepus, respectively, the latter designation being the first identification of this ichnogenus in China. The assemblage indicates a typical Jurassic ichnofauna. The present study reinvestigates the site in the light of the purported new ichnospecies Chuanchengpus shenglingensis that was introduced in 2012. After re- evaluation of the morphological and extramorphological features, C. -
The Braincase, Brain and Palaeobiology of the Basal Sauropodomorph Dinosaur Thecodontosaurus Antiquus
applyparastyle “fig//caption/p[1]” parastyle “FigCapt” Zoological Journal of the Linnean Society, 2020, XX, 1–22. With 10 figures. Downloaded from https://academic.oup.com/zoolinnean/advance-article/doi/10.1093/zoolinnean/zlaa157/6032720 by University of Bristol Library user on 14 December 2020 The braincase, brain and palaeobiology of the basal sauropodomorph dinosaur Thecodontosaurus antiquus ANTONIO BALLELL1,*, J. LOGAN KING1, JAMES M. NEENAN2, EMILY J. RAYFIELD1 and MICHAEL J. BENTON1 1School of Earth Sciences, University of Bristol, Bristol BS8 1RJ, UK 2Oxford University Museum of Natural History, Parks Road, Oxford OX1 3PW, UK Received 27 May 2020; revised 15 October 2020; accepted for publication 26 October 2020 Sauropodomorph dinosaurs underwent drastic changes in their anatomy and ecology throughout their evolution. The Late Triassic Thecodontosaurus antiquus occupies a basal position within Sauropodomorpha, being a key taxon for documenting how those morphofunctional transitions occurred. Here, we redescribe the braincase osteology and reconstruct the neuroanatomy of Thecodontosaurus, based on computed tomography data. The braincase of Thecodontosaurus shares the presence of medial basioccipital components of the basal tubera and a U-shaped basioccipital–parabasisphenoid suture with other basal sauropodomorphs and shows a distinct combination of characters: a straight outline of the braincase floor, an undivided metotic foramen, an unossified gap, large floccular fossae, basipterygoid processes perpendicular to the cultriform process in lateral view and a rhomboid foramen magnum. We reinterpret these braincase features in the light of new discoveries in dinosaur anatomy. Our endocranial reconstruction reveals important aspects of the palaeobiology of Thecodontosaurus, supporting a bipedal stance and cursorial habits, with adaptations to retain a steady head and gaze while moving. -
Oldest Known Dinosaurian Nesting Site and Reproductive Biology of the Early Jurassic Sauropodomorph Massospondylus
Oldest known dinosaurian nesting site and reproductive biology of the Early Jurassic sauropodomorph Massospondylus Robert R. Reisza,1, David C. Evansb, Eric M. Robertsc, Hans-Dieter Suesd, and Adam M. Yatese aDepartment of Biology, University of Toronto Mississauga, Mississauga, ON L5L 1C6, Canada; bRoyal Ontario Museum, Toronto, ON M5S 2C6, Canada; cSchool of Earth and Environmental Sciences, James Cook University, Townsville, 4811 QLD, Australia; dDepartment of Paleobiology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20013; and eBernard Price Institute for Palaeontological Research, University of the Witwatersrand, Wits 2050, Johannesburg, South Africa Edited by Steven M. Stanley, University of Hawaii, Honolulu, HI, and approved December 16, 2011 (received for review June 10, 2011) The extensive Early Jurassic continental strata of southern Africa clutches and recognition of the earliest known dinosaurian nesting have yielded an exceptional record of dinosaurs that includes complex at the Rooidraai locality. scores of partial to complete skeletons of the sauropodomorph Massospondylus, ranging from embryos to large adults. In 1976 an Results incomplete egg clutch including in ovo embryos of this dinosaur, Our work at the Rooidraai locality has yielded multiple in situ the oldest known example in the fossil record, was collected from clutches of eggs as well as fragmentary eggshell and bones, all from a road-cut talus, but its exact provenance was uncertain. An exca- a 2-m-thick interval of muddy siltstone 25 m from the top of the vation program at the site started in 2006 has yielded multiple in Lower Jurassic Upper Elliot Formation (“Stormberg Group,” situ egg clutches, documenting the oldest known dinosaurian Karoo Supergroup) (11). -
Studies on Continental Late Triassic Tetrapod Biochronology. I. the Type Locality of Saturnalia Tupiniquim and the Faunal Succession in South Brazil
Journal of South American Earth Sciences 19 (2005) 205–218 www.elsevier.com/locate/jsames Studies on continental Late Triassic tetrapod biochronology. I. The type locality of Saturnalia tupiniquim and the faunal succession in south Brazil Max Cardoso Langer* Departamento de Biologia, FFCLRP, Universidade de Sa˜o Paulo (USP), Av. Bandeirantes 3900, 14040-901 Ribeira˜o Preto, SP, Brazil Received 1 November 2003; accepted 1 January 2005 Abstract Late Triassic deposits of the Parana´ Basin, Rio Grande do Sul, Brazil, encompass a single third-order, tetrapod-bearing sedimentary sequence that includes parts of the Alemoa Member (Santa Maria Formation) and the Caturrita Formation. A rich, diverse succession of terrestrial tetrapod communities is recorded in these sediments, which can be divided into at least three faunal associations. The stem- sauropodomorph Saturnalia tupiniquim was collected in the locality known as ‘Waldsanga’ near the city of Santa Maria. In that area, the deposits of the Alemoa Member yield the ‘Alemoa local fauna,’ which typifies the first association; includes the rhynchosaur Hyperodapedon, aetosaurs, and basal dinosaurs; and is coeval with the lower fauna of the Ischigualasto Formation, Bermejo Basin, NW Argentina. The second association is recorded in deposits of both the Alemoa Member and the Caturrita Formation, characterized by the rhynchosaur ‘Scaphonyx’ sulcognathus and the cynodont Exaeretodon, and correlated with the upper fauna of the Ischigualasto Formation. Various isolated outcrops of the Caturrita Formation yield tetrapod fossils that correspond to post-Ischigualastian faunas but might not belong to a single faunal association. The record of the dicynodont Jachaleria suggests correlations with the lower part of the Los Colorados Formation, NW Argentina, whereas remains of derived tritheledontid cynodonts indicate younger ages. -
The Origins of Dinosauria: Much Ado About Nothing
View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by RERO DOC Digital Library [Palaeontology, Vol. 57, Part 3, 2014, pp. 469–478] FRONTIERS IN PALAEONTOLOGY THE ORIGINS OF DINOSAURIA: MUCH ADO ABOUT NOTHING by MAX C. LANGER Departamento de Biologia, FFCLRP, Universidade de S~ao Paulo, Av. Bandeirantes 3900,14040-901, Ribeir~ao Preto, SP Brazil; e-mail: [email protected] Typescript received 19 February 2014; accepted in revised form 7 March 2014 Abstract: Research this century has greatly improved our early members of the main dinosauromorph lineages and knowledge of the origin and early radiation of dinosaurs. requires a more meticulous assessment of characters and The unearthing of several new dinosaurs and close out- homologies than those recently conducted. Presently, the groups from Triassic rocks from various parts of the oldest uncontroversial dinosaur records come from Late world, coupled with improved phylogenetic analyses, has Triassic (Carnian) rocks of South America, southern Africa set a basic framework in terms of timing of events and and India, hinting at a south-western Pangaea origin of the macroevolutionary patterns. However, important parts of group. Besides, macroevolutionary approaches suggest that the early dinosauromorph evolutionary history are still the rise of dinosaurs was a more gradual process than pre- poorly understood, rendering uncertain the phylogenetic viously understood. Obviously, these tentative scenarios position of silesaurids as either non-dinosaur Dinosaurifor- need to be tested by new fossil finds, which should also mes or ornithischians, as well as that of various early help close the major gaps recognized in the fossil record of saurischians, such as Eoraptor lunensis and herrerasaurs, as Triassic dinosauromorphs. -
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A couple of partially-feathered creatures about the The Outside Story size of a turkey pop out of a stand of ferns. By the water you spot a flock of bigger animals, lean and predatory, catching fish. And then an even bigger pair of animals, each longer than a car, with ostentatious crests on their heads, stalk out of the heat haze. The fish-catchers dart aside, but the new pair have just come to drink. We can only speculate what a walk through Jurassic New England would be like, but the fossil record leaves many hints. According to Matthew Inabinett, one of the Beneski Museum of Natural History’s senior docents and a student of vertebrate paleontology, dinosaur footprints found in the sedimentary rock of the Connecticut Valley reveal much about these animals and their environment. At the time, the land that we know as New England was further south, close to where Cuba is now. A system of rift basins that cradled lakes ran right through our region, from North Carolina to Nova Scotia. As reliable sources of water, with plants for the herbivores and fish for the carnivores, the lakes would have been havens of life. While most of the fossil footprints found in New England so far are in the lower Connecticut Valley, Dinosaur Tracks they provide a window into a world that extended throughout the region. According to Inabinett, the By: Rachel Marie Sargent tracks generally fall into four groupings. He explained that these names are for the tracks, not Imagine taking a walk through a part of New the dinosaurs that made them, since, “it’s very England you’ve never seen—how it was 190 million difficult, if not impossible, to match a footprint to a years ago. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
Perennial Lakes As an Environmental Control on Theropod Movement in the Jurassic of the Hartford Basin
geosciences Article Perennial Lakes as an Environmental Control on Theropod Movement in the Jurassic of the Hartford Basin Patrick R. Getty 1,*, Christopher Aucoin 2, Nathaniel Fox 3, Aaron Judge 4, Laurel Hardy 5 and Andrew M. Bush 1,6 1 Center for Integrative Geosciences, University of Connecticut, 354 Mansfield Road, U-1045, Storrs, CT 06269, USA 2 Department of Geology, University of Cincinnati, 500 Geology Physics Building, P.O. Box 210013, Cincinnati, OH 45221, USA; [email protected] 3 Environmental Systems Graduate Group, University of California, 5200 North Lake Road, Merced, CA 95340, USA; [email protected] 4 14 Carleton Street, South Hadley, MA 01075, USA; [email protected] 5 1476 Poquonock Avenue, Windsor, CT 06095, USA; [email protected] 6 Department of Evolutionary Biology, University of Connecticut, 75 North Eagleville Road, U-3403, Storrs, CT 06269, USA; [email protected] * Correspondence: [email protected]; Tel.: +1-413-348-6288 Academic Editors: Neil Donald Lewis Clark and Jesús Martínez Frías Received: 2 February 2017; Accepted: 14 March 2017; Published: 18 March 2017 Abstract: Eubrontes giganteus is a common ichnospecies of large dinosaur track in the Early Jurassic rocks of the Hartford and Deerfield basins in Connecticut and Massachusetts, USA. It has been proposed that the trackmaker was gregarious based on parallel trackways at a site in Massachusetts known as Dinosaur Footprint Reservation (DFR). The gregariousness hypothesis is not without its problems, however, since parallelism can be caused by barriers that direct animal travel. We tested the gregariousness hypothesis by examining the orientations of trackways at five sites representing permanent and ephemeral lacustrine environments. -
Massospondylus Carinatus Owen 1854 (Dinosauria: Sauropodomorpha) from the Lower Jurassic of South Africa: Proposed Conservation of Usage by Designation of a Neotype
Massospondylus carinatus Owen 1854 (Dinosauria: Sauropodomorpha) from the Lower Jurassic of South Africa: Proposed conservation of usage by designation of a neotype Adam M. Yates1* & Paul M. Barrett2 1Bernard Price Institute for Palaeontological Research, University of the Witwatersrand, Private Bag 3, WITS, 2050 Johannesburg, South Africa 2Department of Palaeontology, The Natural History Museum, Cromwell Road, London, SW7 5BD, U.K. Received 17 February 2010. Accepted 12 November 2010 The purpose of this article is to preserve the usage of the binomen Massospondylus carinatus by designating a neotype specimen. Massospondylus is the most abundant basal sauropodomorph dinosaur from the Early Jurassic strata of southern Africa. This taxon forms the basis for an extensive palaeobiological literature and is the eponym of Massospondylidae and the nominal taxon of a biostratigraphical unit in current usage, the ‘Massospondylus Range Zone’. The syntype series of M. carinatus (five disarticulated and broken vertebrae) was destroyed during World War II, but plaster casts and illustrations of the material survive. Nonetheless, these materials cannot act as type material for this taxon under the rules of the ICZN Code. In order to avoid nomenclatural instability, we hereby designate BP/1/4934 (a skull and largely complete postcranial skeleton) as the neotype of Massospondylus carinatus. Keywords: Dinosauria, Sauropodomorpha, Massospondylidae, Massospondylus, Massospondylus carinatus, neotype, South Africa, upper Elliot Formation, Early Jurassic. INTRODUCTION same taxon, possibly even the same individual, as at least Richard Owen described and named Massospondylus some of the syntype series of Massospondylus carinatus. carinatus (1854, p. 97) with carinatus as the type species of Their initial separation from Massospondylus carinatus the genus by monotypy. -
Dinosaurs British Isles
DINOSAURS of the BRITISH ISLES Dean R. Lomax & Nobumichi Tamura Foreword by Dr Paul M. Barrett (Natural History Museum, London) Skeletal reconstructions by Scott Hartman, Jaime A. Headden & Gregory S. Paul Life and scene reconstructions by Nobumichi Tamura & James McKay CONTENTS Foreword by Dr Paul M. Barrett.............................................................................10 Foreword by the authors........................................................................................11 Acknowledgements................................................................................................12 Museum and institutional abbreviations...............................................................13 Introduction: An age-old interest..........................................................................16 What is a dinosaur?................................................................................................18 The question of birds and the ‘extinction’ of the dinosaurs..................................25 The age of dinosaurs..............................................................................................30 Taxonomy: The naming of species.......................................................................34 Dinosaur classification...........................................................................................37 Saurischian dinosaurs............................................................................................39 Theropoda............................................................................................................39