ivist stlin di leontologi e trtigrfi volume IIT noF P ppF IUQEIVV tuly PHIH ri vehsxsex pvye pyw ri gesxe fih @wishi vswiyxiD wyxi ex qsyqsyD sivexhAX ivswsxe i v I P hyvp ygue 8 iivx u egri rd eeivedX perury WD PHIHY eptedX wy Q PHIH uey wordsX vgersttteD widdle rissiD wonte n qiorgioD È sntrodution outhern elpsD lnt fossilsD iltoldusF he gssin eds elong to the fmous fossiliferE ous levels of the widdle rissiwonte n qiorgio estrtF e newly opened exvtion in the gssin eds of the vower weride vimestone @wonte n qiorgio xigy rvD vgersttte @ xigy orld reritge vistD gnton È gnton iinoD outhern elpsA hs yielded smll olletion of iinoD outhern elpsY pigF IAD whih is renowned for vdinin plnt fossilsD together with verterte @mostly fishA nd its rih fun of widdle rissi mrine vertertes @eFgF inverterte reminsF he flor ontins t lest five speiesY onifer uuhnEhnyder IWURY furgin et lF IWVWY furgin IWWWAF È È remins ssignle to the gener iltoldusD oltzi nd celourde possils hve een olleted t the stlin side of the re the most ommon elementsF e new speiesD iltoldus ssine th nF spFD is formlly desriedF goEourring with the onifers re seed entury @gornE wonte n qiorgio sine the midEIW ferns @tilozmitesA nd few puttive ydlen remins @ceniopE li IVSRY toppni IVSUA nd on the wiss side sine IWPR terisAF emong the identified generD only oltzi hs previously een @for review see purrer PHHQAF sn ddition to the mrine reported from wonte n qiorgioF he fossils presented in this pper @nd terrestrilA reptiles nd fishesD highly diverse inE indite tht diversified flor thrived in the region during the verterte fun omposed of mmonoidsD ivlvesD vdininF plorl omposition nd preservtion ptterns re suggestive gstropodsD ehinodermsD forminifers nd rthropodsD of tphonomillyEised reord nd reltively frEwy soure reF s well s vriety of plntsD inluding lgeD lnd plntsD nd undnt plynomorphsD hve een disovE issuntoF n nuovo sito di svo pleontologio reentemenE eredF te perto negli strti di gssin99 del glre di weride snferiore hespite the ourrene of fossils throughout the @wonte n qiorgio xigy rvD gnton iinoD elpi weridioE entire setion rnging from the upper enisin to the nliA h onsentito di portre ll lueD oltre fossili di verterti upper vdininD lnd plnt remins hve not reeived @prevlentemente pesiA e inverterti mriniD nhe resti di pinte ny sholrly ttentionD nd no detiled study hs een terrestriF v mroflor ppre ompost d lmeno S speieD tr ui onduted sine pulition of the works of ordelli prevlgono le onifereD riferite i generi iltoldusD oltzi eceE lourdeF peli on semi @tilozmitesA e presunti resti di ydli @ordelli IVUWD IVWTAF @ceniopterisA sono rionosiuti per l prim volt nel giimento ith regrd to wesozoipleootnyD witzerE del wonte n qiorgioF iene desritt un nuov speieD iltoldus lnd is fmous for single siteD iFeF the grnin flors ssineF hei generi identifitiD solo oltzi er sinor segnlto dl of xeuewelt ner fsel @eFgF reer IVTSY veuthrdt IWHID wonte n qiorgioF uesto primo studio di dettglio dell mroflor IWHQD IWHRY urusel 8 veshik IWSSD IWSWY urusel 8 È È fossile proveniente dl nuovo svo suggerise pertnto un9ssoizioE ne floristi en piuÁ diversifit rispetto qunto desumiile dll hrshmidt IWTTAF ynly few other rissi plnt letterturF snoltreD l omposizione dell flor fossile e l su onserE fossil lolities hve een desried from witzerlnd vzione fnno supporre he l9re di provenienz fosse reltivmente to dte @veuthrdt IWISAF vdinin plnt fossilsD inludE lontnF ing shoots of oltzi nd siliified wood @erurioxyE I wuseo gntonle di tori xturleD ile gttneo RD TWHH vugnoD witzerlnd nd niversite de vusnneD snstitut de qeÂologie et leÂontologieD enthropoleD IHIS vusnneD witzerlndF iEmilX rudolfFstokrdtiFh P xturmuseum udtirolD findergsse ID QWIHH folznoD stlyF iEmilX ivelynFuusttsherdnturmuseumFit È IUR tokr F 8 uusttsher iF pigF I E wp of the wonte n qiorE gio re showing the widdle rissi ronte sequene nd the exvtion site @fter Switzerland tokr PHIHAF Italy Lake Lugano Monte San Giorgio Serpiano Meride Cassina Limestone 45°54' N San Giorgio Dolomite Porto Ceresio Besano Formation Lower Salvatore Dolomite 50m Poncione d'Arzo Meride Besano Arzo 1 km Cassina beds / locality E 8°57' lonAD re known from the eustrolpine rosnto porE the ltest enisin nd vdininD shllowEwter sedimenE ttion ontinued in the northD wheres the formtion of mtion in estern witzerlnd @purrer IWWSD ppF TVPE QHEIHH m deep nd IH to PH km wide intrpltform TVQY qruvogelEtmm et lF PHHQAF sin with restrited irultion in the wonte n qiorE sn PHHTD new exvtion in the gssin eds gio re @fernsoni IWWRY purrer IWWSA resulted in the fossiliferous horizon @pigF IA ws lunhed y the wuE deposition of the fesno pormtionD n qiorgio hoE seo gntonle di tori xturle @wgxAD vugnoF he lomite nd weride vimestoneF fossiliferous horizon ws first disovered in IWQQ y he fesno pormtion @qrenzitumenzone99Y ollegues of the niversity of urihD who lso onE pruenfelder IWITA is n up to IT m thik lterntion duted severl susequent exvtions in IWQUD from of lk shles nd dolomites ontining the enisinE IWUI to IWUQD nd gin in IWUS @see tokr PHIH for vdinin oundry in its uppermost prt @frk 8 ieE reviewAF ell these field mpigns primrily foused er IWWQY frk et lF PHHSAF e volni sh lyer lying on the verterte fossilsF he im of the new nd onE few metres elow this oundry yielded n E miniE going exvtion is to refully reEdoument the level in mum ge of PRIFP HFV w @wundil et lF IWWTAF he order to provide etter hrteriztion of the sinF fesno pormtion yielded most of the spetulr verE vnd plnt remins our within the reovered orytoE terte fossils @reptiles nd fishesAD together with imporE enosisD thus providing first insights into the struture tnt index fossils suh s mmonoids nd donellidsD nd omposition of the vegettion of the reF nd in upwrd diretion it grdes into the TH m thik n qiorgio holomiteF he overlying RHH to THH m thik weride vimestone @purrer IWWSA egins with qeologil setting the vower weride vimestoneD WH m @irz IWRSA to he widdle rissi suession t wonte n ISH m thik @purrer IWWSAF his deposit ontins three qiorgio @pigsF IEPA strts with fluvioEdeltisequene verterte fossil eds @iFeF the gv inferioreD gv dted s lte enisin @fellno pormtionD sllyrinY omE superiore nd gssin edsAD eh with different verE mrug et lF IWWUA tht overlies ermin volni terte ssemlge @nder IWVWY furgin IWWWAF he fosE È suessionF he upper enisin sediments indite proE siliferous eds onsist of finely lminted limestones gressive trnsgression of shllow epiontinentl se with interlted volni sh lyersF nd the relted growth of ronte pltforms @lvE he top of the vower weride vimestone is deE tore holomiteY orn IWUIA north of lnd re uried fined y dolomite ed @holomitnd99Y pruenfelder tody under the o lin @iotti et lF PHHUAF huring IWITAD rehing thikness of out QH m @irz IWRSAF he ldinin flor from the gssin eds @weride vimestoneD wonte n qiorgioD witzerlndA IUS pigF P E ynopsis of the rissiplnt fossils from the wonte n qiorgio reD plotted on the Liassic strtigrphisetionin purE rer @IWWSD modifiedAF ht Tremona Series from different uthors @see Rhaet textAF yther fossils of terresE tril provenne re lso inE ditedF Dolomia Principale Norian Pizzella Marls Carnian Kalkschieferzone Ladinian "Dolomitband" Meride Limestone Cassina beds Cava superiore beds Cava inferiore beds San Giorgio Dolomite Besano Fm. Lower Salvatore Dolomite Anisian Bellano Fm. 100 m Rhyolithe and associated volcaniclastics Permian Triassic 0 m Plant fossils Other terrestrial fossils Voltzia sp. Terrestrial reptiles Horse tails Insects (including Equisetites sp.) Undetermined plant remains Documented occurrence Occurrence questionable or of plant fossils poorly stratigraphically constrained IUT tokr F 8 uusttsher iF he overlying pper weride vimestone shows lterntE VQSAD the ulkshieferzone ontins oltzi remins ing wellEedded limestones nd mrlstones with n inE only in the lower nd middle prtD wheres otherD uniE resing ly ontent towrds the top where strong seE dentified plnt remins re rre @lower prtA to sre sonl vritions of slinity nd wter levelD together @middle nd upper prt of the sequeneAF roweverD with the influene from nery elevted reD re reE these onlusions were sed exlusively on three smll ognizleF he uppermost prt omprises the soElled exvtions rried out in IWWR in the l wr setion ulkshieferzone99D IPH m thikD whih yielded peuE nd on the review of irz9s originl dt nd smples lir fish funsD rustens nd rthropods @eFgF purrer from the sme lolity @irz IWRSD see elowAF purrer IWWSY urzeminski 8 vomrdo PHHIY vomrdo PHHPY @PHHID ppF TEUA lso reported rre findings of iquisetites vomrdo 8 intori PHHRY intori 8 vomrdo PHHUAD @ stem frgment ws lredy figured in purrer IWWWD nd represents the lte evolution of the intrpltform figF PIA from the gv superiore eds nd rre olified sinD followed y the grnin regressive phse @izzelE wood frgments from the gv inferiore eds @oth l wrlsAF levels elonging to the vower weride vimestoneAF pelE er @PHHSD figF IRSA illustrted smll sl ontining shoot frgment sried to oltzi long with smll he terrestril mroflor from wonte n qiorgio plnt frgment ttriuted y the sme uthor to the @inluding gssin edsAX stte of the rt horsetilsF eording to our oservtionsD this speiE men omes from the lower ulkshieferzone nd eE errestril plnt remins our throughout the longs to the irz olletionF htz @PHHSD pF IHHA menE widdle rissi ronte sequene @pigF PA tht ws tions oltzi spF shoots from the lowermost prt of deposited within the sutropil zone t pleoltiE weride vimestone etween the n qiorgio holomite tude of round IVëx @frk et lF IWWWA under