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r . n n M n Wh n nd r r h d v r t & C n rv t n Gr p n ln S n C ntr .O. x 6 , n ln l nd
r .K. Cr b nd Mr .S. d l M n Wh n nd r r h d v r t & C n rv t n Gr p M nt Alb rt r h C ntr r v t 2 0, A l nd l nd
U I E SI IES E ESE A I E r .M. E b r n p rt nt f Ent l .O. x 84, n ln Un v r t l nd
MUSEUMS E ESE A I E Mr . . l t r l Env r n nt p rt nt M f l nd .O. x 46 , W ll n t n l nd
O E SEAS E ESE A I E r . . r n CSl O v n f Ent l G. .O. x 00, C nb rr C t A.C. . 260 , A tr l
SE IES E I O ` AU A O EW EA A Mr C. . v l M n Wh n nd r r h d v r t & C n rv t n Gr p M nt Alb rt r h C ntr r v t 2 0, A l nd, l nd Fauna of New Zealand Ko te Aitanga Pepeke o Aotearoa Number / Nama 34
Anthicidae (Insecta: Coleoptera)
F. G. Werner † p rt nt f Ent C Il f A r It r Un v r t f Ar z n n A 57 1 U S A [†d d]
nd D. S. Chandler p rt nt f Ent C ll f f S n nd A r lt r Un v r t f p h r rh 3 U S A
Manaaki Whenua PRESS n ln C nt rb r l nd 1995 E I O S E ACE Floyd G. Werner died on 20 December 1992, nearly 20 years after he unknowingly initiated this project by agreeing to identify some introduced anthicids for Willy Kuschel. Floyd always enjoyed a challenge, and had nearly completed his study of the entire New Zealand anthicid fauna at the time of his death. One of his last acts was to make certain that this contribution would be finished, by bringing in as co-author his former student Donald S. Chandler.
C p r ht © I995 nd r r h l nd td
p rt f th r v r d b p r ht b r pr d d r p d n n f r r b n n ( r ph l tr n r h n l n l d n ph t p n r rd n t p n nf r t n r tr v l t r th r th t th r tt n p r n f th p bl h r
C t l n n p bl t n WERNER, G Anth d (In t C l pt r / G W rn r S Ch ndl r — n ln C nt rb r l nd M n Wh n r 1995 ( n f l nd ISS 111-53 3 ; n 3 IS - 7 - 5 7- I Ch ndl r S ( n ld 11 tl IlI S r U C 595 7 7 17(931
r p r d f r p bl t n b th S r Ed t r n p t r-b d t xt pr n l t nd pr nt n t nd r r h l nd td Mt Alb rt r h C ntr r v t 9 17 A l nd
M r t xt b Un S rv r n l t n C ntr A l nd
r nt d b G r nt td W ll n t n
bl h d b M n Wh n r nd r r h l nd td C x n ln C nt rb r l nd
r nt v r h b tl d p t d Anthicus hesperi (Ill tr t r W l r Ar K t pīt r n Anthicus hesperi (K - h āh W l r O U A SUMMA Y E W AKA O O GA MA E MA EA
Cl / K r h Insecta Ord r / O t Coleoptera
l / Wh r Anthicidae
Ant-like flower beetles
Ill tr t n: C t r p (Ill tr t r: .W. l r Wh āh : C t r p (K h āh : .W. l r
Anth d r t nl ll d th ` nt-l fl r K n Anth d ōh hān t h `p pōp r
b tl th h n f t r l t v l f p r pā p p ; n t n h n hān h - nl ll t d n fl r nd n l nd p h n p t p t rā īh n h īr n r f nd fr nt n fl r h ll t v b tl t n n h p t p t n K ēn pāp p t h r r d l r n d b th r h rpl n tr t d n r r ōh t t n t t ī t n t h h h ft n b n d th l t r l n tr t n f th t t h n t r t t ō K n t r pr n t h p tt rn f th n tr t n p rf ll o aua ū t n r n n h h n r t t t āh t n nt-l pp r n h h p rt l rl t pōp r t r t t t t h nh n d b th r b h rt-l d t h n l v h r ō rāt p t t ā r n S p n th r ntr r nl ll t d Κ ēt h ēt h t h n h hān t th nt nd r t d ff lt t d t n h h n n pōp r n nō t p t t t ā v n b t n nt r n r bl th th nt h r n n pōp r h p n pr t t v r h l r h n f r f d n h r t n t n pōp r h āh r Ad lt r v n r r pr d t r f d n n ll rāt t ā r p r p nv rt br t r n v t t v d br A f p n K n p h h h h t t rth A r h v b n nv t t d f r th r v l n n p r n ār r tūār r t n t t rān K ēt h b l l ntr l b th f d n th nd ll t Κōt Ā r r n h t t t l rv f v r l n t p t p n h t h h t t n t Anth d r ft n f nd b n th bj t l n n nd n hē n r t n n ār r r tā r l r r nn n b t n v t t n d r n th d K n ī h nō p r ā r nō ōh S p r tr n l t d th t l nd r t h n r h n r t nō n n d n t p h t r r n t ll n n nd pp r ntl h v K n t t Anth d r r n t t n d t l r t th t f th d lt Ad lt fr n p h n t n pū t n n rān h r are tr n l ttr t d t nth r d n th rr t nt pr d d t rā r t n t t K ēt h rāt t n b b tl f th f l M l d nd f nth r d n r āh t t t h t h n n t p t p hau dr n l d b tl n b t ff t v h n tr pp n n pū . d lt Anth d A t n f th h l b l v d K n p t n n n h t h t nh n t n nd d r pr d t n Attr - ` nth r d n h h hōh p t t hān
( nt n d v rl f ( r h r t n Ill tr t n: r d br n (Ill tr t r A C rr Wh āh : r d br n (K - h āh A C rr
n t lpt r f b d pāp p M l d K t p n nth r d n t n l d nd r h r h r pāp p l d n t tōt t ā h p t mau t t p Anth d E h p n t n t h h n t r n p r t t h h t h t n āh p t p t Kāh r nō h āt h pēh t āh p nth r d n n n h n īr n t ā n n ōh t n n īr n mau n h h p n n na t h n h p h — h t n Ah t r r t t n h n t t pā n n āh t n t nth r d n h n t t b n t t d f r th n h n K n tūtūr t ā h n l nd p n pān n t t tūtūr ` n t - nt - x p f Anth d r n n n fr l d n l nd n f th h v b n ntr d d pr b- E r nō n t W n ā hā bl thr h h n r — v n r fr A tr l h t I A Mā K n h h nd th th r t r p l t n p t d th t tāt C t l nth t Τīr n h n ttl nt h v nt n n t v p r fr - n n r pāt t n n t ntl t d th nd p f n t - Ah t r r t n Ā r K n āh t n l d n pl nt h n n t n n ār r t Anth d Τīr n p r f nd nl n th S th I l nd nd f r pāt t n Ah t r r ā h t t n t pān n r f nd nl n th rth I l nd M b r f th n r t ā n h n t n Ā r C t nd l nth r r tr t d t l nd nd r l t t n r f nd n A tr l nd S th A r Floyd G. Werner, h n t h n Ill n h f n l r l t n h p f th l nd Anth d h tōn āt r n t Wh r Wān n rv rd r l rl l t t A tr l nd r r t l t r r r r t n t h tā t h t t 195 I t t t p r t S th A r 195 n Ar z n r r n h t n t n t h n n hān pāp p l d G W rn r b rn n Ill n d t d t rv rd M l d Anth d Ad r d n h n āh Un v r t nd r v d h h th r n 195 In 195 h h n hān t r r ē nō Na t āh p t v d t Ar z n h r h p nt th r t f h l f ēt h t tēnā t p p t h t p l t n th b tl f l M l d t t h hān n t n r r n h t n Anth d nd Ad r d l v d n n r h r th n t n t n t n t ēn ā p t t f l r t d v r nd n ntl th r d/ - h ld n rōp h n n ār r th r d 1 r p p r th f n l t t nd l l tr t nt f th nd n th r r p f Donald S. Chandler h n t t t t h tōn n t h n āt r n t h n C l f rn r t h MS t Wh r W n n Ο Ar z n r r l d W rn r n ld S. Ch ndl r b rn nd n t ll d t d n ā , r r t t h n tā t h t Wh r Wān n C l f rn f ll n th n MS t th Un v r t f Oh St t I t t 19 1 h n t t r ār r Ar z n nd r l d W rn r nd h t h Oh St t t Wh r Wān n Ο p h r ā h t n Un v r t In 19 1 h j n d th p rt nt f Ent l- n r n h pā n n t n n ār r t h Un v r t f p h r nd nt n d Anth d rā l ph d ā n p t n r r h n th t t f Anth d nd l ph d āh t n h r r pāp p n h n t p p n h r nd ff t f d t rb n n f r t fl r b tl h K ia t t h r t ā r p p pā n th r d 55 p p r v r n t t f n l nd - āh tāt n h n h n ār r n t n t n t l l tr t nt f v r l b tl f l ēt h hān pāp p A S AC
h Anth d f l nd r r v d f r th f r t t nd th r t x n ff n t r d d nt - x p h v b n f nd ht f th nd r b d nd ht n n n r d v r d n f th p r ntr d d v n f th r n t n fr A tr l ; th th r t r p l t n nthr p n p M - b r f n C t r ll r tr t d t l nd th n t p n n l nth . h n n n l t b l l t n nd r ph d tr b t n f th p r r d th p nd t n l l t r rd A t th r d nt f t n v n nd t p r ll tr t d
C ECK IS O A A C t h ll n Βr n 19 3 28 pt r n 1 93 9 S bf l A ICI AE pr x r n 1 93 n n n G n Anth ll 179 1 pr b r n 1 1 3 ( Anth n tr t d r l Βr n 1 93 n n n h p r K n 1 9 1 r f r n 1 93 3 r l r K n 1 9 17 n n Βr n 191 n n n (b tr l r p v t t Sh rp 1 77 31 h n p 1 d t n t Βr n 1 93 n n n tr l nt n 19 5 1 l n 1 95 G n r h n n l b rn 1 91 31 ( tr t r p f l d n p 3 l b r K n 1 9 19 n r r n 1 19 l nth n n 33 fl v t r Βr n 191 n n n l t n p 33 t n n p 20 (d S b n O n d M l nt 1 S bf l MAC A II AE fl r l ( nn 175 1 G n M r tr n 1 3 3 f ll x Βr n 1 93 t r n p 3 f r r (G z 1777 1 x l 1 77 35 v rt l Sh rp 1 77 G n S p nt C 1 95 22 fl v p r n 1 93 n n n l nd n n p 22 d tz n p 24 b r rn (Βr n 1 n b n t n 24 nthr n ( r n 1 93 n n n CO E S p ll d p ( r n 1 n b n t n 5 A n l d nt 8 Intr d t n S bf l AG IOI I AE 8 M rph l 8 G n r d r r G r n 1 5 l 1
br n 1 7 26 M th d nd nv nt n 11 S t t 1 S bf l EMO I AE K t n r nd p 13 G n C t Sh rp 1 77 26 r pt n 1 b ll t Βr n 19 3 7 f r n 35 r p ( r n 1 7 Ill tr t n 39 p n t t r n 1 93 n n n tr b t n p 5 rl n p 28 x n nd x 5
— — ACK OW E GME S S t t r n th l nd Anth d r- t nl r fl t th p tt rn r v tr t nt f G ll r K h l ff t v l n t t d th t d h n h l nd nth d n t f l t d p d r pt n nt t d GW n 197 b t d nt f n ntr d d b r n (1 p ( p nd Sh rp ( nth d ll t d d r n t d f th b tl f n f th p fr 1 7 t 19 3 Oth r th n th r l t n n A l nd b rb f nf ld h r t l d v r th t l ( tt n 19 1911 b d n 19 3 l r nt rv n n r t r v n f th nt r l nd 19 b th nl b nt p p r d l n th th nth d f n W th th d th f GW n b r l nd f n th n n f Anth f ll x nd r 199 pl t n f th pr j t nd rt n b SC Anth fl r l b (195 l nt f th r n th l n th p r d f th t d h v nj d pr n t v n C t h b n p rt l rl n t nd r t nd n - p r t n n rr n n l n f p - n f p n f th n x t t d f ll - n n l d n t p b n f nd v d l t t n n l nd r n th r t h M ( t r l v r l n t t t n W h r th n th nd th r l t d t r nd f p l t n Anth d h v v r n th th r r p t v n t t t n n th t n n bbr v - n p n t n In p rt l r ld l t n l d G ll r nt - x p r n n n fr l nd (`W ll K h l r v r Cr b nd Ch rl W tt f th n r d f n t r p t d ntr d t n fr th r l nd Arthr p d C ll t n f r th r nf l n ntr ; v n fr A tr l —A nth l b r, A. h , n r nt nd - p r t n n v l t d A. h p r , A. r l r , A. tr l , S p nt d tz , nd r tl thr h t n f p n l l t th r h n n f l d , th th l t t nd A h th r r d nd n pr p r t n f th p W nd r b d; nd t p l t n Anth fl r l nd l r ( nd r r h A l nd nd G n ll A f r r . v p fτ th r n n v nt n (Un v r t f Ar z n n pr p r d n f th r h r d r b d n h n r S p nt nd h b t ll tr t n M r l v hn r r r h n n r n r n d fr l nd nd (Un v r t f p h r nd hn r n pr n t v n n l nth , d f n d C t , (CSI O A tr l r d ll r p rt f th n r pt th nl th r tr l pr n t v n h v r l p St f n C v r ( rv rd Un v r t d nt f d nt - th th fl ht n r d d t d th tr p l A nth n p Wh l l ttl n n b t p r l t n h p th n SC bl t tr v l t A tr l nd l nd n th l r p l t n n r Anth , S p nt , nd 1993 thr h th rd f Un v r t r v l Gr nt b th M r tr , th th r p th r b l n t n r n p rt nt f E t M n nt Un v r t f t l nd (C t nd l nth r r n n r En l nd Ar d l S W A tr l h r nt f nd nl n A tr l l nd nd th rn S th ll d tr v l t th A tr l n th t h ld t p A r ( r d nd r h n n . C t , r h n t r l f p ntr d d t l nd nd p r t- n , nd l nth r b r f th d n t d v t t th l nd Arthr p d C ll t n n bf l t d v r t th n r nd p f l v l n A l nd ld r n A tr l nh n d thr h A tr l h t n t v p f Anth r h r th rd f h l r h ll h p b th pl d b r f th tr t r p, h h l t Un v r t f En l nd h h ll d d nt t n d v r n A tr l h b r ph ff n t f th f th h b t t f n f th ntr d d A tr l n p l nd nth d f n l rl nd t n n nt r n n n th th A tr l n f n th b r f t f th n r p r l r pr nt d n th rn I O UC IO S th A r M b r f th Anth d th nt-l fl r b tl r t nl t d th r p r n r nd v nn h nd t l r xt nt f r t h r f n MO O OGY n r ll ll n n f r rth A r nd E r nd h t l f r Afr nd S th A r h h f ll n n r l d r pt n f th f l b d n Or nt l A tr l n nd l nd f n h v ll b r f ll th bf l x pt I h l n nd b n l r l n l t d n th rl 19 nd th h Afr n Wh l t n th pl nt f th n p h v b n d r b d th r h v n t b n n r p nd th r d n n th Anth d - nth t tr t nt f n p rt f th f n fr th l nd h n p f r d n r d br n . It r th r th n Kr h-St ld l t d r n th tr t d n nth d n th r v n f ll n nt n d h l pp n (19 5 19 9 n l n f th r d n n th f l b r n
–8 – t n (1995 S r f r th t n r Cr n b t n h d pr n t nd l tr S r p h v (1955 r n (19 nd r n r tt n (1991 r l p n t r n th l tr h l n th r th p n t r h t tr n f t r f th Anth d b th r nf d d j nt d pp r n f n f th p h h d d l n th th d t n fr l p l p x t l tr l h rpl n tr t d b h nd th t f r n rr n p x In p th bd n xt nd n t bl th pr n t l t l n tr t d l t r ll n r th b nd th l tr b t n t b t v r bl b th pr n t l b n rr r th n th l tr l b xt nd d r ntr t d t n t n l d d n l n th r - th th r j n t n l htl n tr t d nd th l tr l nt p tb l tr n v r pr n ft n p l - l r d r nd r n th v l ff t f f rth r n tr t n h Full description of adult morphology b d t p ll t n br n r d r br n nd fr ntl d d fl x d tr n l n tr t d b h nd t f r h p l b nd r p t n th l tr th h b r f n n -h lf r l f h d dth (b r l n tr t d n d h n nd r rph Ch p n (A - r d n ; f r f h d b t v r bl nd f l tr l nd p - G n b l r d br ll nt n h r t r n p nd n n r E tr n - r d nd p rpl nd Afr n b r f r v r l v l nt r ft n th h rt t r n t n rté-Sén tèr b t ll bl r r n Sp b t n f t Ant nn l b xp d; nt nn n rt d r n fr 1 t 15 n l n th t d f fr n 11- nt d th nt nn r d - t r fr ntl f l n d nt f n p r r n n l n th fr 3rd t l t; t r n l 3 nt b d n d r l l r p tt rn r nt t n f t nd t h rt n d nd/ r br d n d n h t t d n t nd d t n tn f p n t t n C l r p tt rn n l b (M r tr , nd t x r r tr n b t v r bl nd h ld b d th r r - l b ( r d . r nt l p l t r d t n t M nd bl n t n f th t p nd r nt t n f t r t l f r ll tr n l rv d M x ll r p lp th nt r t d nt f t n n n r p th l t ll r f r ( x pt n r d nd th t l t r r t t b r t t ff h r th t ll n nd h r t r t f r f r r p ; nt r- pr j t b v th th r h r h r n r ll p r d t nt th rn r r n pr j t n n f nd pp r t f r rd r d r n th l tr h n th r p n b l rl n In r p th t t l t r r th r x l n t t tr p z d l d t nt r rl t h rt nd d ff lt t ( h n b r f th th l t r l f v d r l t pr x nd t n r n f b n O n d f Anth . h h r th t r h rt r p d f r n nt rn l l r f pr x l v t nd n l d l r th n th t t l t r r f rr d t th ( b nt n r d nd St r p n r x l v t ` t nd r ll h d n r h t n b p n p t r rl ( x pt n n r f E r n n ; d n r p r n d ff r nt r f th l tr r nt d n pr x n l pr n nt nt r n t th d ff r nt d r t n nd r d t h r t r t n l l t r l r n ll r nd d r r l t l n l t ; fr th rf f th b d th l t d r n f th n ll d f n d l ft n r n t n t l t r l f v v r t n f th p n h n t b n r hl h ndl d nd xt nd n p t r d r ll t r d r j t nt r r S t l n l r ll t l rl v bl n th l tr t b ; t d t n t l t r l n tr t n l n l t r l h v r n l f l n dr n fr th t l h ft n r r f th l xt nd n r d r h ll th b t n r th p x n r f r n t th l tr l rf t d p r v n r p ; pr n t l p x th b v n n d r r h r t r d r n r ll fl n d r r ` ll r n M r tr n nd Anth n ; ` ppr d th t p f th l htl rv d t t h n t x n (Anth n th nt r d r l r n tr n l th b d rf ; ` bd b nt l htl r d th pr j t n t f r `h rn d t d n nd nd l n r l n l f th h ft 1 - °; `d b nt l rl (Ch ndl r 19 r d - °; ` b r t - °; r r t -9 ° A M t rn br dl tr n l r r nd d t n l t f r p h th d n nd S p nt n th nt r rl th p t rn n rl t n (r nd d r Anth n h v th rd l r f d n h r th t h v t n ( n l t t ddl f nt r r r n nd th ll d ` nd r t h r ll ppr d nd p t r d l xt n n p r t n x n l t b r nt d n v r d r t n t v rl d pp r- ll r p M t rn l- p t rn l t r d t n t ( x- n ft n n nj n t n th th r nt t n f th t pt n M r tr n t r l nd p t r r r n f tt rn f p n t t n r ft n r t l n p r - p t rn r tl b d r n l d n r p t n t n h h d l p n t r n n rr l n t d - l t v r p r r n r p t h f d n r n l z n n th fr nt r th d b v nl p n t r d l n t t f nd n nt r r r p t r r r n h z d n t nd d pth f th p n t r v r f p t rn nd h n d n l fr n d ft n th
-9- p r r d nd n t v bl th d l f v IO OGY r n t n fr r d p r n S p nt p Anth d r rld d nd r ft n n n nd 1 r f v n th tr t r p f Anth . - r d r r n n fr dl nd t d rt r n M x p n l t r ll M t x th h rt nt rn l f l n f r t Sp fr n b r f n r h v fl n ; t x l v t p r t M t nd t rn t th d pt d t th x r nv r n nt f t l nd nl nd nd t l d h rt r th n l t r l r th l n ; nt r r d n r l n / l l n d fl t h f l l t nd n r n n t l ll b v j n t n th l - t d f nth d h v d lt th th z n t n f p n M t ll ll ll tr n l r ( dr t n f t l b h f n ( n h tt t l. 19 S l ; d n El tr ll th h r l n l C l b n t l. 19 5 1991 It l b l nd b h v r d t n t; fl htl p ( C t nd th r n r f n rthr C n l l n l f th t rn Un t d l n th n l `O pl t (r d r nd d r t St t ( t r n 1973 Ch ndl r 1975 nd n l t n ntr f h l tr n n r b n t pp r nt n fl htl t n ( r hh r 19 Wh l b r f v- p t p n n b r f A nth n rté- r l l r n r h Anth nd S p nt r - Sén tèr (Anth n tr p l tb l tr n v r nl f nd n th r nd n l tt r r nd r l -l n pr n p t r r t pl t ll br d nd v pl nt n r p r n r th r h M r tr b ft n th p l b nd r p t n t pp r nt n n f nd r l n b t n v t t n d r n th d In fl htl p t r l rv f l tr th l n t d - t p r t r n ll nl f p tt r d n l r v n b r f M r tr . l ht n th thr h v r l n r b nl f nd n tr ll d v l p d p t b t l p t h b th p l p r v t t n (th l r l r t n M r h r Ch vr l t pl t b nt r th nt p n lt t t r r n x pt n th h th f n f nd n tr f l n pl p n lt t t r r l b d t xt nt; l th tr p h r d v r In f r t d r f r z l pl t pp nd l t ( GW nd n ( SC f nd th r t t n b r Abd n th 5 f ll xp d fr t rn ( rph l - f p h l b t n / p n tr f l l n tr l l nt 3-7 ; b l t rn f d n r d . M l r t l r n r n h l nl f p ld b f n r p th p d ( rph l l t r f nd n th r nd A l t d rv f th f l f n t l t l htl v bl b t pl t l v r d b t r f nd l d (S nt l lb n Ind pr d d 7 n th r r p M l ft n th 5th v bl t rn t p ( b rd 193 b r n p l d f t n (tr n t pr d r n t th t ft f t St rn t ll pl Ad lt f th p l v n n th r nd r th ht d f d n M r tr nd Anth n ( h t b v n r nd pp rt n t pr d t r f d n n r , A nd Afr St rn t 9 ll Y- h p d r n d br nd ll r n d nv rt br t th t th l n b l t l nd h rt r r ( th t b l t l th n bd (W rn r 19 Ch ndl r 19 h n St r p n ; p l r rt l t d n r p nd nth d p f nd d r n t d f b l l ntr l l t n th r ( - h p d pr nt th n r d n nt n th v t t n f tt n ( tl r 19 Orph n d d r n A d th p r t p r r nd t l. 1971 nd M nt r p n ( n r d t ( nd hr p n n b r f r p h M r tr n 1977 h v b n pr r l pr d f d n n t E r n n nd t x n (Anth n r r f d l r th r ll nv rt br t Wh l n v t t n t f r t n n r p h r d n d n th r l n n t f d n p ll n ( nd hr 1977 n C p b n n d r n nd Anth n ( x pt n x d t fr l nd (S th 19 r n f n l h ph b r f I h r p lp rté-Sén tèr Anth n nd p r ( h r 191 nt n 19 5 W rld h ll b p r t n ll r p x pt rv h v b n f nd n th r nd n d n C p b n n d n nd d r n (f d b t v t t n h r th r n v r r t ph d t n t n d t d r Old W rld n l n r n h ph r p r ( nt n 19 5 K t 19 Y n th n r t f d n d n nd C p b n n 1991b r r pp rt n t pr d t r ( r t Anth Int rn l ft n th p t h f p n f v r n dth h r C n f C r d l d ( v d n nd l n th r r n p r ft n th r t r n W d 19 9 ; r t M lp r n t (S n pt r Anth n p p r (K t 19 h t r f n rth l th t rn t 5 ( rph l l t rn 7 - A r n p t x n d n ( br r n n ll br dl r nd d G n t l ll b d n l n t b r nt t p t t t b r (C thb rt 19 7 t b l r v p t r C x t v r l t th r n dv ntr l h nl d n f r th f l th ` nt-l r n nd rtl d v d d v ntr ll ; t l ll b rn n fl r b tl h t l d n n th t nl n nt d r n pl t l - nt d x t ll n b r f p r t ll fr ntl ll t d n
—1 0 fl r h n tr t n f th n nd pr n t nd nl n n t nt p d h n b n h d h n h d th fr nt r n f th l tr l p tb l pr n ff t v l n r nd d br f v n th pp rt n t th p l b nd ll t d j nt d nd nt-l tf ll tr p n b t pr d t v f r f th pp r n h p r pt n p rt l rl nh n d b p nd fl ht- nt r pt tr p r ll f r th th b h v r f nth d h n l v h h v b th f ll d v l p d n S p n r p r n v t t n h rt-l d t th t t t v f nt v nt ll pr d b r f r p h M r tr , nd Wh l nl ll n b r f p r n n t b p bl l l nth . t n l tr br n h nd r ph l (W nn 1 9 K tn r 19 SC b- b h n b th t pr f t bl ll t n pr d r n rv d b r f th l r tr p l n A nth n tr p l f r t nd dl nd rr n th nt n v t t n (A. nt n W rn r On f th t pr d t v ll t n t hn th nt Cr tο t r p n n r n th r nd (A b d n th ttr t n f n p t ltr v l t nd rn t ( b rd th nt r l p nd r h h t l ht (W rn r 19 b t h v n v r f r x p n M x th t h f nd t l r n p n fr l nd ll t d b th n pp r n nd p tt nn f v nt h th t h h d M n b r f l r n r r ttr t d t dr n d ff lt n r n n th t nth d r pr nt h n l d b tl nd nth r d n n rth A r (Ch n- x n n b t n h t r nt r n dl r 197 E r p (Görn tz 1937 Afr (v n ll 195 C rt n n r f nth d h v n p th t r r hh r 19 nd S th t A (Y n 19 b ttr t d t nth r d n th rr t t n fl d x d d b b t tr p d n d t xpl t th ttr t n (Ch ndl r 197 v r t f bl t r b tl (M l d (Y n 19 S h tz Y n 19 b r t t b t t d n l nd ttn r 199 h d lt ll l ll f d n Sp n r t p ll ll t d nt 7 % th n l nd d d l d b tl r nth r d n r t l t bt n th t r d nt l th r pr p r d A f p n t r l h h th n t r d n th b d C nth r d n n r r l r n h t b p nn d b t t r th r b l v d t t f d n d t rr nt (C rr l E n r l d n tr n l r p nt n th A r n t l r nt d 197 d r n pr d t n n nd v d l th t n f nd n rd th t r- l bl l n th E r p n t l It r E n r (19 h h n th t n th l l r l t d p rt nt h n nt n n rd th t th l d n t f l r hr d l th t nth r d n ll t th l tr h r t t d n th p b n t h v n r d r pr d t v l ll t dr W d f n t l pr f r p nt- nt d p n p r- r r d l th th nth r d n-r h nd v d l nd t l rl h n r n th l r l n n n f n h v th t r l p d t th d r n t n th n th Ex n t n f th l n t l p rt nt t nf r p r t fl d h nth r d n th n n rp r t d nt d nt f t n f f l nd p nd t n b th nd v nt ll th l rv h f d n d t rr nt p bl t r n l f th d n nd th p d n t pr t t th r v ln r bl t M r tr n h n th r nt d n rd n th Anth d r pr d t d r v th b n f t fr p d h dd n b th pr d n t r t nd th f r f n t n nth r d n nd n r d t n h th f fth t rn t n t b n v r th r h ld b b n h n f r p f h h th l nd l ttl pr bl n d nt f n t l nd p n f d n nth r d n r (S hätz ttn r 199 v n f th r nt d n rd r b r f M r tr nd th v t t - r p f C t th l n t l h ld b x n d nt l n ME O S A CO E IO S f l r th th r btl xt rn l d ff r n S ft n th h l p n n h t t r f r t l t 1 n t th n Collecting, preparation, and curation pl t n ll d h n 7 % th n l G ntl l ft p th Anth d t p ll r f nd n th r nd th r r l n l tr nd r p th bd n n r th ddl th p l- f b t r h d n nd r bj t r b l r l n r nd n f r p U n n n t p n th rv d t p r t l t v t t n M b r f Anth , S p nt , r d , p r t n th t r t nd t rn t pl t l n n d nd t ll r xt nt C t r t fr ntl n th h l n t l n th n b t d t n th rv d r nd nd b bt n d thr h t r h p n r r p d th n th r t f f r p nd p ll d t nd r bj t h tt d pl nt r pl nt d br b rd M b r f C t n b d ff lt b th p n nd b h r nd d n h r n ll b t n nth t rn t r l n th bd n nd th l t fr tf l p rt l rl h n th r th r nd r th v r n h v t b nl r d n rd r t pl t l p ll t th t r l l htl d p An p r t r r tt d f n r n t l On th r fr f th b d d t h th n b d t p p p n t pl n v l t d br n nd n nth t rn t nt l th t n Ind v d l n h n d r t n l b t t h v nd ph ll b ( f pr nt n b l rl n l th
—11— hth t rn t n M r tr . r nd r nt th d lt SYS EMA ICS th th p rt f th l n t l t r d n l r l n r p l p nn d thr h th t pp r b n th th p - C p t n f th f l Anth d h v r d n d r- n l b l r l d n th p nt r rd n t r- bl v r th l t 1 r nd t ll n t pl t l l bl l ttl d h bf l Anth n f r th r f th f l nd b d n Anth ll nd t d Abbr v t n n r n th Anth n pl th n l d d n th t x n h t -l tt r d d t th b nn n f r rd r n ( t x G ffr M n t r rté-Sén tèr nd th r f d tr b t n r f r t r f l nd thr th r n r Unt l r ntl th d r n ( d r rté Sén tèr , d t d r , d f n d b Cr b t l. (197 Abbr v t n f r p t- nd r r d r v d fr W tt (1979 nd Arn tt t l. (1993 thr th r n r r tr t d tr b n th Anth n b t r n r n d d t n t bf l ( r n AMΝΖ A l nd In t t t nd M A l nd t n 1995 Al t ll p p r b nth d p l t l nd ( W E rl h v tr t d j t th thr r p p n th AMSA h A tr l n M S dn S W A - f l f nd n th l t rld t l ( 1911b tr l ( M M ld nd j r f n l tr t nt b n d n (195 M d - A IC A tr l n t n l In t C ll t n CSI O r; 1971-9 r n r ll (19 It l nd C nb rr A C A tr l ( r n Uh nn (199 Ib r n n n l M h t r l t r M nd n U K v r v r t f th r r p h v b n r f rr d t (C M v n p p E th Anth d nd h l r f th pl nt rd (Uh nn 197 197 nth d p l t h v n r d CM C C n d M f t r Ott Ont r th th h r n ll th rr nt p t n v r C n d ( S And r n Gén r l t th t f nd n th l t rld r v n f th f l CM C nt rb r M Chr t h r h - ( rté-Sén tèr 1 h d l d h v ft n b n l nd (C A M r A S v ll r n d d t n t f l ( 1911 Arn tt 19 3 U n ln Un v r t C nt rb r l nd h ld n th bf l d l n E r n n ( M E b r n C A M r Marris) M r tr n nd St r p n Cr n (1955 b d MAMU M l M S dn Un v r t S dn th d l d nt r l th n th Anth d b t th S W A tr l ( S rn n r G h n t n v r ll pt d M r r ntl ll Y n MC C M f C p r t v l rv rd Un - (1991 ff t v l d b r d th f l b d f n t v l v r t C br d M h tt U S A ( tr n f rr n th d l n ( d l h r nd An tr r n Y n t th r hr d nd nf r n pl nt f th th r thr bf l Anth d S n r M M f l nd (f r rl t n l f r rl pl d r hr d r n n d r d t b M W ll n t n l nd ( W nth d h b r f th d n ( l r 1913 br G Ord h l S rv d r tt n 197 nd I h l n (Y n 19 5 nt dd - AC Arthr p d C ll t n Mt Alb rt r h t n r th C p b n n (C p b n r r C ntr A l nd l nd ( K Cr b G r n fr Ch l (Abd ll h 19 9 nd th Afr n G K h l C W tt (Afr v fr th rn Afr ( v 19 SAMA S th A tr l n M Ad l d S th r p th t h v b n br fl pl d n th Anth d nd A tr l A tr l (E G M tth r n n d r d t b b r f th r hr d r UAIC Un v r t f Ar z n n Ar z n U S A th A n th n (= C n n t n n r A n th nd (C A Ol n C n n t ( n 1979 Y n 1991 nd th l - U C Un v r t f p h r rh p lp n (= h n ( ll 199 h rr ntl p h r U S A ( ll t n f S Ch nd- pt d h h r l f t n f r th Anth d v n l r b r n t n (1995 M M für t r nd d r b ldt- h h r n (1 n t ll pl d M r tr n th Un v r tät rl n G r n (M Uhl d l d th l t f th f l f r l nd h v r n d t bl n th l nd l t r t r thr h th C ll t n d t n l n f M r tr b tt n (19 nd d n ll d t f r t r l x n d r v l bl l tr n ll (19 3 n bf l d l n f th Anth d Abd ll h nd h rd p fr AC Abd ll h (19 dd d t th f l b tr n f rr n
—1 — Lagrioida ( r n ll pl d r d nd b ntl h d n nd r d n r b d n th rn M l ndr d O d r d nd C n n t d t th t p r t r p rr n n A tr l l nd Anth d th l n t t nt f th r d n In nd th rn S th A r th th d n xt nd- r p n n th r p d n t b l n n th Anth d n t th tr p n p - G n h bf l b t f ll th l f t n f r n t n h v n t b n th r hl d f n d nd ph l n t (1995 n tr t n t h r Anth d h v th b f th n l f th h h r t r b dl n d d hn h d tr n l n tr t d t f r d t n t n h v r n (CSI O C nb rr A tr l nd SC r fr nt l p l t r th pr n t p l t r l f v n t t n ph l n t n l f th h h r r p th t r th r n f th nt rn l p d p t r rl h bf l f nd n l nd can b p r t d l n th pr x l v t ( x pt n Steropes), nd th b th f ll n f t r f r t t v bl bd n l t rn t r fr l rt l t d • r d n h d f bl n tr t d p t r r t Lagrioida h th h d b nl l htl n tr t d l n t f r n d t n t n ; nt nn th n p l l b fr nt l p l t r l th l t r l pr n t l f v l n l t r l pr n t l f v ; f r t bd n l t rn t nd th f r t t v bl bd n l t rn t r f d h f d; l n t l th ph ll b p n nd p r r r p d h v th h rt nt rn l fl n f th h nd x pr nt f nd n b r f th Anth d h f ll n h rt h f ll n thr bf l h v d t n t n rr h r t r t n f nth d d lt nd l rv d n t n d t n t l t r l pr n t l f v nd fr bd n l n l d th nfl t n f t r f r d v r d n th t rn t . pr d n nt n • Anth n pr n t th n p l ll r n Anth n n t h r t r (l n n t x n ; t rn nd p t rn p - r t ; l tr l n l t r l r v ; l n t l th Ad lt d h rpl n tr t d p t r r t t f r ph ll b pr nt nd p n n t xt nd n nt r rl d t n t n ; fr nt l p l t r pr nt; nt nn l b nd ph ll b n rt n xp d r n t d t nt r rl ; n r p th l t r l n tr t n n r b th l t r l • d n pr n t l n n p l ll r; f v nd t n r n f nt rn l p d f r n p - t rn nd p t rn p r t ; l n t l t r r r n f pr x l v t ; pr x l v t p n l n ph ll b nd p n l n r th n t n xt rn ll l d nt rn ll ( l d xt rn ll f r n r • M r tr n pr n t th n p l ll r; f E r n n p n nt rn ll nd xt rn ll f r t rn nd p t rn nd t n h bl f d; St r p n ; l t r l r n f pr n t ll r nd d l tr th l n t d n l r v t l t r l n l ; l M t rn n rr d nt r rl ; p t rn t n n t l th ph ll b nd p r r pr nt p n r l t t n t nt r r r n Exp d bd n l tr p rt t nd n t xt nd n b nd ph ll b t rn t fr l rt l t d r n (19 v th t r nt d n f th d lt
rv rr tr l b l ndr l l htl l r t d x- KEY O A ICI AE pt f r thp rt nd t p f r ph ( h n pr nt K OW OM EW EA A d ll th n l ll n th r d n- ll th r (Lagrioida th 5 r n n ; p r n l t 1 d nl l htl n rr d p t r r t l n h rt r b nt; n l d n nd r n pr nt d t n t n ( ; pr th r x fl t n r p t r d r l M n-d bl th ll d v l p d p n ll r br h f p n - t pr x l n f v ; pr n t th l t r l l t t b f l Ur ph ( h n pr nt r n tr ht n d r l v l n n tr t n l n p r f d n p t rv l d n r v n b n r b ( r d n p 5 Lagrioida brouni r n (19 nd Y n (1991 b — d h rpl n tr t d p t r r t t f r K t (19 h r t r d th l rv f t n n r n n -h lf r l d h d r ( f Anth n f nd n rth A r h th r b- 1-5 ; pr th r x th f v r d pl p t r d r l t f l th r h v th l rv f b r f nl n pr x ; pr n t t l t l htl n tr t d l t r- n d r b d (St r p n E r n n r n l rv ll n r b r d r b d (C p b n n r d n d n M r tr n (1 El tr th l n t d n l l r n t h r l r bf l r r pr nt d n l nd n l nd xt nd n t l t h lf f l tr l l n th; Anth n r d n d n nd M r tr n nt nn th p l 3 nt d n d t f r
–13– l b ( 5 ; t rn nd t n h bl f d — d b btr n t ; pr n t r l p n t t th p t rn (M r tr n Macratria) .. 3 v r d ; l tr p l th br n b nd r ddl —El tr l n l n t d n l l n r l t r l r- nd br n t b nd p x ( 17 ; l n th 9- n ; nt nn l n d t n t l b ( 1- ; 3 1 (p Sapintus deitzi t rn nd p t rn l rl p r t 9(5 d th t ppr d h rt ( l n nd 3( El tr l t b r t r d t 3 - ° nd p n t r h rt t t l t n n p ; h d nd pr n t l d f n d; l th t rn t th h ll n t h th r t l t r lpt r ; l tr br n t bl t br d p x ( 7 (p 3 Macratria aotearoa r dd h r b ( 1 ( b Omonadus) .. 10 —El tr l t bd b nt r d t b t 15- ° — r nd l r v r bl ll th d t n t t t l nd p n t r h rpl d f n d; l th t rn t th t r l n p b n nd l n r lpt r d p n t h t n rr p x ( n h d nd pr n t 11 (p 35 Macratria exilis 1 (9 M t rn th l t r l r n tr n l b d ( r th r x th p x t ff d t n t ll r ( 1 l t r ll th n t r fr n f t ppr d t 2) (Anth n 5 p t rn ( ; pr n t th p r f ll — r th r x th t t n b r r r nd p x n t b p n r dl n b h nd nt r r r n; l n th n tr t d t b t f r ll r ( 3 1 3 1 -3 7 (p 1 Anthicus floralis ( d n 17 —M t rn th l t r l r n l htl b d l t- r ll l n n fr n f t ; pr n t th 5( r t v bl bd n l t rn th n rr tr n - nt r rl ; l n th 3 1 -3 7 v r v t p t r r t h x th v t v r d (p 1 Anthicus formicarius b d n h r z nt l r f n rv d ll t ; rn t rn th l t r l r n l htl rv d; 11(9 M t rn th l t r l r n tr ht l n p t rn th p t r r r n b r n d n d n t t h nd n l f p t rn fr n f t ( 3 ; l tr th ppr d t (Anthicus n tr t 1 bd b nt nd bl l d r t d nd r t —M t rn th l t r l r n b d r n l rl (Sapintus) .. 6 pr j t n ; h nd n l r l t r l r n f p t rn — r t v bl bd n l t rn l n v t r th r r t ft f t ( 1 13 r f t ; t rn th l t r l r n tr ht r tr n l b d r n l d; p t rn th 1 (11 r r 3 -3 5 l n ( 1 ; l tr l t t l p t r r r n l n d n t r f pp r ntl t r t n p h lf n l n b- d n l t th n t rn th l t r l r n d b nt t (p 1 Anthicus hesperi tr n l b d l t r ll ( - ; l tr l n —S ll r 1 - l n ( 11 ; l tr l t t l nd r t (Anthicus) .. 9 t r t n n p h lf l n bd b nt t (p 17 Anthicus kreusleri (5 El tr l h r b l t ; b d r n ; l tr th br n b nd t b ddl nd p x ( ; l n th 13(11 M t rn br dl l b d l t r ll ( ; - (p Sapintus aucklandensis l t r l r n f p t rn th fr n f t —El tr l h r d t n t 7 l v t d b v rf p rtl v bl fr b v ; d r br n th h t b nd n l tr ; b d l th d th 7( d d r br n th l tr l n r n ( h rt ppr d p b n (australis-group) .. 1 1 ; l ll p l ; l n th 3 7 - —M t rn n l rl pr j t n t n r l tr l r- (p Sapintus obscuricornis n th l n t ft f d r ll d r t d t r n —El tr th p l nd d r r n fr p t rn n r n l t n ( 1 (strictus-group) .. 15 (7 d b r l r; pr n t l d th l t r l p rt n p r l p n t t ddl d t n tl p n t t ; 1 (13 d b l n t p r l n l r nd d; l tr p l th br n b nd r ddl nt r- pr n t l n n nt b l tr n v r l ; l tr r pt d t t r ( 19 v r n t br n th th p t d n h t b nd br d r th n p tb l l t d p l r ; l n th 3 15-3 b nd ( 1 ; l n th 5 - 73 (p 5 Sapintus pellucidipes (p 1 Anthicus troilus
-14- — d b br dl btr n t th d t n t t p r l (19 r n t l d th d n n t p n t r nd n l ; pr n t th n nt b l tr n v r l ; th nt b l n tr t n n t nt n n r d ; l tr th p t d n nd p tb l h t b nd f br n; l n th 3-7 l dth; l n th 9 (p 9 Cotes optima (p 1 Anthicus gushi — d nd pr n t pp r n p n t t ; nt b l n tr t n xt nd n d r ll r d th n- 15(13 r n t th tr n v r l th n b t d t n t tr t n d t n t n l t r l v 1 r b nd l t r l r n h rpl n l t n d t l h lf n rl r n t ; n f r l t n ( 13 ; 1( r n t l b d r th n d th ll b nd l tr l h r nd n ll d v l p d; l n th - r p tbas l pr n; p b n n b nd n 5 (p 19 Anthicus glaber d d r t d bl l l t r ll nd n d d r t d — r n t th t tr n v r l t b r l d r ll ; l tr l h r pr n nt ( 9 ; l n th r pr nt d b ll v l pr n nd l t r l 5 -5 (p 3 Cotes rufa r n n rr l t br dl r nd d; br n - - r n t l d d b r d r; l tr th t th p l r r n 1 l r t n v r bl p b n ft n d r t d p t- r rl n n t rl nd h r t b nt 1 (15 r n t th ll v l pr n n r b ( 3 pp r ntl r n nt f b l l ; l tr th t l t b r p l r r t pr x l nd d t l th rd ( 1 S ll r 3 - ; b d h n n 3 ( 1 ; h d b br dl btr n t ; l n th - r r 1 -7 5 ; d ll r n t br n 5 (p 19 Anthicus minor — r n t thl rv d n r b th t tr 3( El tr br n ll th d t n t r l r ll f nt b l l ; nt r l br n r l tr b r l r d r n r l t r l r n f d t n t p tb l p l r t b ( 15 ; h d v l b h nd ; l n th pr n ( 7 ; t rn nd p t rn 9- 7 (p Anthicus otagensis th 1 r r p n t r ; l n th 3 -3 (p 7 Cotes bullata 17( r n t th d t n t n rr tr n v r l —El tr th br n b l nd d n b nd nd th r d n r b ( 1 ; b d br n; p rt n f p tb l pr n b r l v d nt ( 3 ; - t rn p r t n x d pr d p t- t rn nd p t rn r l p n t t ; l n th r rl th x pp r n nt thr h 3 - (p 7 Cotes crispi p t r r h lf f l n th; t ll r n t t p x; l n th - 7 ( El tr l d th r t t t l t d t n t r (p 33 Zealanthicus sulcatus th n t l n ( 1 b r t t ( 7 — r n t l n tr n v r l t b p bl ; h r l n l ll b t d t n t; h d b th n xt n n f l t r l n tr t n r d br dl btr n t th t p r l n l r nd d b t ( 3 ; t rn t n t t rn d - d t n t ( ; pr n t l d nv x; l n t l ll l rl p r t n x 1 th t n b r n pr p l l t r l l b ( ; l n th 1 -7 5 (p Cotes gourlayi 1 (17 M t ll h ll l r n t t p x; — El tr l d th r t t t l t h rd t f nd b t pr n t l d th v l n t d n l pr n l n b r t t ; h d b h ll l rv d fl tt n d d r l t l t r l nt b l n tr t n ( l n v rt x l n l t f bl btr n t 5 ; l n th 3 3 -3 (p 3 Trichananca fulgida 5( r n t l d n rl fl t n pl n l tr l —M t ll r nd d t btr n t t p x; pr n t l d nd b d r ; l tr ft n th d n br n d r nd d ft n th l t r l n tr t n nt n n p t nd th ll b t d t n t h r l n l ; l r d r ( 3 7-9 (C t .. 19 n t l th t n b r n pr p l l t r l l b ( ; l n th 5 - 7 (p 3 Cotes proba 19(1 r n t l d th r t l t r lpt r nd — r n t l d h ll l nv x r d b v pl n f p n t r n rl l r f t ; r n -br n; l tr l d ; l tr n l r l n h r l n- l n th 5 1 -5 1 (p Cotes halliana l ; l n t l th l t r l pr p l r n f — r n t l d pp r n p n t t r th d n f n t n p r ll l ( ; l n th 7 - p n t t n (p 31 Cotes vestita
-15- ESC I IO S pr n t n th pr x l th rd th l t r l n l t n f th rn t rn n r t p t r r r n th t ft f d r ll S bf l A ICI AE d r t d t r n fr th p t rn n r th n l t n th r d p r th t h v 1 r G n Anth ll nt rn ll d r t d f v ( 1 th l f nd r t n th l tr nd 1- l t r l t n r th t n p x Anth ll 179 53 Τ p p M l nth r n h l t t p fl r l nd f r r , r - nn b d n t n f n d n (195 rn p l t n ntr d t n nl pl d n th n r nt l p l l r pr nt d b f n l n ; pr n t O n d M l nt b E r p n th r th h th d t n t p l ll r; pr x l v t p n xt r- h r tr t d b n f Anth . M b r f n ll l d nt rn ll ; n t pl r l l br f nd - O n d r n r ll t r n d t n t ll f ntl d f n d b v pr x ; x p r t d r p th th r h rt ppr d t v r h rt t t l b pr j t n f t rn nd t t rn rn; l tr t r lpt r f th f r b d nd n r l l r l n nd r t ; v bl t rn t fr ; 9th t rn t Y- p tt rn v r h f th d t n h n f t r n h p d th rt l t d xt n n t p x f h rt l t r l t n h r d th b r f Anth n r ; t n nd ph ll b p r t th p n h rt r tr t h th t t p bl t pr nt n x l v th n th t b n d h r t r t n f th r p n O n d p d d ff r fr Anth n tr t n l n th p l r Anth d ff lt t h r t r n th t r f th n nth t rn t h h f r pl r d th n d p r t t x r r n ll d r b d n th h rt l t r l xt n n t th p x n nd h v n t b n b ntl r v d It All Anth p h th r ntr d d r n t v r l r nd h t r n r p rId- d nd p - pr b bl d rn l h ntr d d p r tr n l r h It nd bt dl p r ph l t b t tt pt t br t d th d t rb d r n r h n h b t t n t nt n ph l t n t h v th f r b n d pp nt n h f n r l t n th t n b d t h r t r Anth r l t d b v Anth n tr t Anth r pr nt d b n n p n l nd Ant l h p r K n p r pl d n Anth n tr t —h p r nd r l r , b th ntr d t n fr A tr l M b r 1 ; M p 1 f th rld d r p r r n d b th tr ht t h p r K n 1 9 1 (Anth . 1911b 5 l htl b d l t r l r n f th t rn l f 19 7 (A tr l n n n l t d b l nd r t n th l tr pr n t th nl f nt l t r l t r M 1 7 3 7 (Anth . n tr t n pr n f b l tr n v r l th t l 1 95 1 (Anth . n l nt r rl t th l t r l r n t th pr n t l f v nd l f t r v n th f r t v bl n th 3 -3 5 d h n n ; h d d r br n; bd n l t rn t pr th r x r n t br n; l tr d r br n th Anth h nd A tr l r d t n t v l l n rr ll t r n tr n v r b nd ; b nd n p tb l r l t d p l ntr d d fr A tr l nd h r tr n v r pr n t r h n l t r l r n pl d n r p b d n tr l (K n nd th r nd t l t v l nt rr pt d t t r ; l htl p t d n A tr l n p h l rl d n t b l n n A nth b nd n t r h n l t r l r n nd n rr l nt rr pt d n th b f th r br dl l b d t rn th n t t r ; tr n l r d nd v d l t th rnp n n fr n f t n th l t r l r n f th br d r nt r r b nd b t th ft n r d d t v p t rn ( W r n t f l r th ll th nd t n n d r nd v d l th p t r r b nd d t n t; n r n n fr S th t A nd A tr l nd l br n th f r l b r n pr f r t l v th d t n t v r p n Anth nt l d nd pr n t th tt r d h rt r t t t l d f n t v t t nt n b d n t n r pl nt t ; t d b nt d b b dr t th hr p — l b r, n r, nd t n — r - d n nd nt t n; d th th v nl Sp d b r f p - r p pr p d h r th t b d n d t n t p n t r ppr x p rt; fr nt l p l tr t Er h n nd th r A tr l n p th n r l nd t d b f n l n E ll l n bv nd t n pp r t b r tr t d t l nd th t b t n f t l b r n ntr d t n fr A tr l h d t n t v r th r x br d t t d t l f rth; r n l t r p h r t r d b th h rp n tr t n f th v nl nv r nt fr d t p rt t b nl v r
–1 – f bl nv x n d r l v ; p n t t n l r t th t f ll t r n b nd ; b nd r p tb as l tr n - h d; tr n v r nt b l l xt nd n nt r v ntr ll v r pr n t nd n t xt nd p t r rl l n t r t l t r l rn r n t l t r l pr n t l f v t p t d n b nd nd r h n l t r l l tr l r n ; El tr j t p r pt bl d n n t ddl ; h r l p t d n b nd nt rr pt d t t r n d r t nd v d - n l pr rn n nt; p tb l tr n v r pr n nd l nd n t r h n l tr l r n ; l nt nn nd pl t ; t r l r th dj nt f n l t r l thp rt r n l n d f n r d r n d t l h lf; r t t t l t d nd pr n t th t ppr d l n t t l d t n t 1 l n ; t b r t nd d r t d b- t d th b b dr t br dl pr d l l l t r ll l n r d t b t ° t r l d ll ; p n t r ll b t h rpl d f n d ppr x t rn l r n tr ht W n f ll d v l p d 3 p rt b nt fr dl n ; fr nt l p l l r f r ll n n b th x M l th h nd nd t d b a f n l n E th t n t pp r nt tr h nt r d ll n l t n p t r r r n nd l t b t n f t v bl t rn t br dl r n t G n t l t n th r th r x br d t t d t l f rth h r d v nl h rt p l xp n n; nt rn l l n d f - r nd d nd th n ntl nv x b t d t n t l t r l t n n tr t n; p n t r l htl d n r p rt; d t n t tr n v r nt b l l xt nd n nt r v nt- p d t Anth h p r : nt p l b ll d ` rr tt r ll t r n t pr n t l p t K/ Κ35 5 " (AMSA ; l d r b d fr G l r El tr l htl d n d t ddl ; h r l n l pr - S th A tr l A t r d r b d fr G nd h n nt; pl t n t pp r nt; t r l r th dj nt Q n l nd nd A l d r b d fr Q nb n l n t d n l l n t h d tl n n l v t d r t n r S W l nd p n r p r d th th b ; h rt r t t t l t tt r d l n ; t p t r l f A. h p r b SC t d b nt t 1 -15° 7 l n M t rn l rn r n l htl b d l t r ll W n f ll d v l p d Material examined. p p n pl 11 n n-t p M l th h nd tr h nt r br fl n l t n p t- x pl ( U MAMU M AC U IC r r r n; l t v bl t rn t br dl r nd d G n t - U C l nt rn l v r d th l htl l r t d h rt AK C G WA W / S M br d p n C ll t d thr h t th r rd d pr r l fr n r l v l Type data. Holotype labelled "G l r K/ Κ35 57" C n n p t r nd t l f nd nd r (AMSA l nd p n r p r d th th d n h rl t r rd r f r A l nd (1915 nd h l t p b SC l n (1935 nd n tr l n p t rd n l f l tt r nd r d n f rn nd n th n t f n Material examined. Holotype and 50 non-type examples thr h rd ph l l . At nf ld (AK f nd n (NZAC, U C p dd nd r p l f r pr n n nd t n AK C WO / tr b d n b h r (K h l 199 3 C ll t d thr h t th r rd d pr r l fr n r l v l Remarks. A. h p r n r ll l r n pp r n t nd n v r t f h b t t n d t rb d r h A. r l r b t l r r th p n t t n f th h d nd l n p t rd n l f l tt r nd r d n nd n pr n t l r r nd l d n nd th nt r r p l p t r th r rd h fr n t f th rn bl - b nd f th l tr n rr b t th dth th b d ll ( r d n n nd t rl n (St rn brown basal b nd Ind n t A tr l v l r . h rl t l nd r rd fr nf ld (AK n 17 A t 197 nd r h r n n rd n C n n th r nd n b h nd p n r n l f l tt r t nf ld nd n ll f nd n tr Anth r l r K n b d f b h r (K h l 199 34, 66). 11 5;M p Remarks. A. r l r ll r th n A h p r , th th Kr l r K n 1869: 18 (Anth . Pic 1911b: 57. Lea p n t r n th h d nd pr n t d n r th p n t t 1922: 7 dl n f th h d d t n t nd th nt r r p l b nd f n th 1 - d h n n ; h d r d-br n t th l tr t l n th b l br n b nd Indigenous br n ntr t n th r n pr th r x; l tr th t A tr l
— — tr l r p d r br n h th l t r l t p r l r n p t r r t Anth h n p th l htl rv d nd th p t l t p x f th t n n rl d th t n b A. tr l is ; M p 3 br n th l t r l t p r l r n r p r ll l nd th n th 9 - 9 d d r br n; pr n t th p t l t t n p x h n rr r b t n -th rd ll r nd b p t r r t nt b l l r n ; l tr th dth f th t n b th p tb l nd p t d n h t b nd th p t d n A. h p r t d fr A. tr l b t br dl b nd l htl th n rr r n th r b nd r h n t r btr n t h d b tr p z d l pr n t b l tr n - b th r h n l t r l r n f l tr nd d r b v r l n th pr n t l v t f r nd th p l d th f ll n p tbas l b nd; f r br n b nd n th l tr th t r ll d All thr p r n t b ; r nd r f l nd nt nn r n t h v l r pp r n b d n th r h r d l r r dd h br n; t d n nd ppr d p l br n; p tt rn ppr d d r t pl d th v r h rt t t l t p r nd r t d ff lt t t t l t d n p n t t n nd f r nd t l p tt rn d th b br dl btr n t r n p t r r f th t rn nd p t rn t l htl rv d t br dl r nd d t p r l n l ; h p n d f r th ll t r f th t p r d nv x p l h d; p n t r ll d t n t ppr x G h p rt; fr nt l p l t r f nt nd t d f n l n E ll pr n nt pp r ntl l n t b t n f t nt n r n t tr p z d l d t t d t l f fth th l t r l Anth tr l r n l n rl nv r nt t l ht l t r l nt b l 1 7; M p n tr t n; d t n t tr n v r nt b l l n l d l n 1 95b 7 ( r . t Anth nt r v ntr ll t l t r l r n t p l h d d pr n l n St v n 1 1 1 1911b 5 b v pr x ; d l htl nv x th l t r l r n tr l nt n 19 5 (Anth . pl nt n h rpl r nd d t d l n r f r t t dth f r l n j n r nd r h n n El tr fl tt n d n d f ntl r r t l t n pr x - Anth . l h lf; l t r l r n bp r ll l p t r r t pr n nt n th 5 - 7 d d r br n d ll; l tr th h r ; pl t nd p tb l tr n v r pr n p l ll p tb l nd p t d n b nd ; p t d n f nt; p r br dl nd nj ntl r nd d; t t l b nd th d r r h n l t r l r n f l tr nd t 1 l n ; t l n br fl rl d r n br dl t t r ; p tb as l b nd nt rr pt d t l t r ll n r f p tb l pr n M t rn t r ; nt nn r n ; l th f r br n t b th l t r l r n br dl l b d l t r ll th fr n nd t r r n ; d r n t l nd d n l p n t t ; f l n p r t n l t r l nd p t r r d n t t t l t r t v r h rt 1 l n ; d n t ppr d t n rr l xp d p r r l r p l br n h rt nd ppr d l v t W n f ll d v l p d M l th l t t rn t br dl tr n t G n t l p x d th b l n t v nl r nd d l n t - br dl p t l t n rl d t n b ; nt rn l p r l n l ; d f ntl l n t d n ll tr ; p n t r th z n f d n p n n pr x l th rd nd n fr n ppr x 1 p rt n rr l b nt fr n th r j t b nd ddl ; 9th t rn t th p l rt - dl n ; fr nt l p l l d t n t E ll pr - l t d r v r h rt n nt; t b t n l r f t b t 1 l n r n t l n t d t t d t l f rth; l t r l r- Type data. Holotype l ( M nd 1 l paratype n l rl n t t f nt l t r l nt b l n tr t n; (MAMU), [GB] K r n r 1991 G h d n rl fl t d r l t n tr t n f ntl r r t l t ; tr n v r nt b l l b nt Material examined. p p n nl El tr fl tt n d n d br dl nd p r t l r nd d
G / — p ll ; l t r l r n l htl d n n t ddl ; h - C ll t d n n r r pr n nt; pl t nd p tb l tr n v r pr - nd b n th d d h d h (Er n r p . n nd t n t; t d r t d l t r ll n r f p tb l pr n M t rn th l t r l r n br dl Remarks. C p r n f A. h th nt p fr l b d th fr n f l n p r t n l t r l nd rr tt f A tr l (K n — p d pr d n p t r r d th n t ppr d t p r r A tr l — h d th t b v r l r A. h is l n t l n r n t ll n W n f ll d v l p d
–18– M l th l t t rn t br dl tr n t G n t l t t l t h rt nd r t 3 l n ; t ppr d t n th p x br ptl h d v ntr ll ; nt rn l l n tl d r t d bl l l t r ll M - th br d z n f d n th n p n n d t l h lf t rn th l t r l r n n l rl pr j t n t n r n rr n t ddl b ll l tr l r n M p t rn th l n t ft f t n r t rn l n l t n W n f ll d v l p d p d t . r b d fr rth t rn A tr l p M l th l t t rn t br dl r nd d G n t l t r l (SAMA x n d b SC n 1993 t n th l t r l r f p n n r p x; nt rn l th br d z n f h rt tr n t p n r d n t l n M t r l x n d. p t r l pl 11 n n-t p th n p n t b nd p x nd th n rr z n t p n ( ΜΝΖ AC U C ddl l n p n —/ ΚA, MC C ll t d n r br r p d t . l t p l b ll d "G l r K Κ35 " rd d fr n r l v l (AMSA l nd p n r p r d th th nd n nd d l n l r n nd n t h l t p b SC l tt r M t r l x n d. l t p pl 17 n n-t p x - r . A. tr l t d t n t n l n b l pl ( AC U C tr n v r l n th pr n t n th l n t h d AK C / — th th b v nl r nd d nd th d p t d n C ll t d O t b r– l b nd t n br dl t th l tr l t r It rt nl rd d fr l v l t l t A tr l n th f r f th t rn nd th M t nl f nd n rb n r n rd n l f d n p n t t n ppr d t nd v r h rt t t l l tt r nd b p n r nd d h rl t t v r th b d th p r l rl n t b r n n l nd r rd 5 n r 1975 fr f Anth , b t nn t pl th n r ppr pr - nf ld (AK h r p n h v b n f nd n t n t th t p dd n n r d t nd n t C rt d r nd nn t (K h l 199 3 5
tr t r p r . A. l b r d t n t b t p l l r nd Anth l b r K n fl tt n d pr n t l d th th l t r l r n tr n l n l t It f rl n n A tr l 13 ; M p 5 l b r K n 1 9 1 (Anth . 1911b 5 K h l 199 5 Anth n r r n
n th - 5 d r n ; l tr h t 1 1 9; M p tr n l nt nd l n r n ; l nd l tr ll n r r n 1 93 (Anth . tt n 19 19 d nd pr n t th d fl tt n d t ppr d 1911b 1 d n 19 3 3 5 nd n n p ; h d b v nl rv d t d t n t fl v t r r n 191 119 (Anth . d n 19 3 t p r l n l ; fr nt l p l t r d t n t; p n t r 3 5 n n ll b t d t n t ppr x t l p rt b t dl n n rr l p n t t E pr n nt l n t b - n th - 5 d h n ; h d nd pr n t t n f t r d-br n t br n th h d ll l htl d r r; r n t nt r r t n tr t n xpl n t ; d n rl l tr br n th b nd ll n b r p t d n r l r fr ll r t n tr t n n d r l v b nd t n d t t d t l th rd; l t r l r n r n t t r t t d nd pr n t th t p r ppr d t dth f d ; b l tr n v r l n l d nt r rl t d b nt nd t t l t p r h rt d th b l l t r l r n t f v b v pr x ; p n t r l r r n n rl tr ht p t r r t f r n -f rth f nd d t n t p rt l rl n pr x l th rd f d 3 h d l n th t p r l n l br dl r nd d nd b p rt br dl btr n t ; d br dl nv x th; p n - El tr th d fl tt n d l n pl t r t r f n p rt b t b nt fr fr nt l dl n ; p tb l tr n v r pr n; h r l n l d t n t; t d r t d ll 5 l n ; t t l t b r t p n t r f nt r th n n pr n t p rt f n r l n ; fr nt l p l t r th n E ll n t t rd p x; r d t r l r n d t n t n d t l h lf; pr n nt l n v bl t b t n f t
– r th r x d t t d t l th rd; r n l t - Anth t n n p r l r fr ll r t nt b l n tr t n; d n r- 15 3 ; M p 7 r l r nd d l t r ll ; d fl tt n d; ll v l f v n r f b l tr n v r l ; p n t r d p r th n n n th 9- 7 d h n n d r br n h d p ll l n dl n fr b t ddl d nd pr n t th d v nl r nd d; t El tr th d fl tt n d rf th; h r d - ppr d; t t l t h rt nd r t v bl nl n h d t n t; pl t nd p tb l tr n v r pr n n t nd pr n t ; p n t r p r nd f n d b th pp r nt; d th f n p n t r p rt d p r r n p t r r t n rl tr ht f r d t n t rd b ; t r l htl l v t d n d t l th rd th v l nt t l n th th n br dl r l r n f n l n t d n l l n t th r d r n b rd r f tl n l n t p r l n l ; p n t r p rt r d r ; t t l t n t pp r nt; t p r nd b nt fr fr nt l dl n ; t t l t l n ; d r t d bl l n d bd b nt t -3 ° 9 fr nt l p l t r j t p r pt bl E ll n t pr - l n W n ll r d d n nt pp r ntl l n t b t n f t M l th h nd t b j t p r pt bl rv d thr h r n t d t t d t l th rd; l t r l r n n rl d t l h lf f fl x r rf th t l n t ; h nd r l r fr ll r t nt b l n tr t n; d tr hant r n l t n p t r r r n; l t v bl t rn t r nd d; b l tr n v r l b nt br dl tr n t t d fl t r l htl nv x G n t l El tr th r n n rl p r ll l; d v nl r nd d t n th n l p n n r ddl f l t r l r n ; nd th; p n t r d r t l f n f n r t rd p x nt rn l th n rr z n f th n p n t b nd p rt; pl t nd p tb as l tr n v r - th p t h f h rt br d p n n r p x pr n b nt; h r l v d nt; l tr l t r l htl l v t d n d t l h lf; t bd b nt t p d t Anth n r: l t t p f l l b ll d b t ° 9 l n f n nd d r d r t d bl l "1 7 / [AK] / n r" ( r n C ll M h r l t r ll n d t r l t rn l r n tr ht d n t d b GW l b d p t r rl th t ft f t n p t rn n r Anth fl v t r : h l t p f l l b ll d "3 37/ l b W n r d d t n rr tr p [S ] W ll t n 15 1 / Anth fl v t r " (Βr n M l th h nd t b f bl x v t d b nd ddl C ll M x n d b GW f fl x r rf th t ft f l n t j t b f r x v t n b t 1 l n ; l t v bl t rn t br dl tr n t th v tr n v r pr n b f r p x M t r l x n d p p n f n r nd fl v t r , pl 1 9 n n-t p x pl (AMΝΖ U G n t l t n th l t r l p n b f r br ptl MAMU M AC U IC U C d n d p x; nt rn l l n p n hr K n I / AK WO G O I WI p d t l t p l [CO] Cr ll tl - W /S M C KA MC MK CO S /Ch th rv 17 M r h 1975 C W tt l tr l 75-1 9 Is. ( AC C ll t d thr h t th r r t p ( 1 l 1 f l ; AC U C rd d fr l v l t 1 7 S th I MC A hb rt n (1 CO Al x ndr Kn bb nd n d v r t f h b t t ll n th n 9 S p 19 d l l ( ; d n r nd n r nd r t pl nt b t l n l nd r 9 S p 19 d l l tr lί ( ; t nt b r b t n t n l h n n C rd l n fl r v 19 n nd n l ( Cr ll 1 S p n d d M hl nb , nd n ph tr l . nd 1 O t 19 d l t pl nt (3 ; 19- v 197 W tt p t tr p (5 ; t p l l t W tt- 17 M r 1973 r S p n fr n rth rn l l t r l tr l ( 17 v 1977 p t tr p (1 ; C t r p l r A n r l r n pp r n t Anth tr t d 17 v 1977 (1 ; 9 M r 1979 nd r d d Er h n fr A tr l th n t n th r p n n t (1 ; 13 M r 1979 l tt r nd d br ( M n A tr l n nd th pr v t b t r (1 Old M n n d l nd 1 S p 19 t p h n ph l n t n l p rf r d h pl nt (9 ; 1 v 19 l t nd l tt r (1 W n h l t p f tr t ( M x n d b SC nd t M r 19 7 W l r t pl nt (1 nf r d th d t n tn f th t p h l tr l t f tr t r n t ntl h rt r nd d n r nd th M t r l x n d p r nl l tr l p n t t n rr p nd n l d n r —/ MC CO
- - C ll t d S pt b r—M r h br ptl n tr t d n r br d p x; nt rn l th M t nl ll t d nd r l tr l l h v l l r t d rv d r t r n r pr r n p r n nd nd r t pl nt Type data. M l fl r l : t p t r l ("E r p " r h l f b l tr n v r l n th pr d t b n th ll t n f th nn n S t f pr n t th d r l r nd th thl rv d pr n t l nd n d th r nd d l t r l r n r d l p r t A Anth f ll x: h l t p l l b ll d " 7 / [AK] t n fr th th r b r f th tr t r p. / Anth f ll x" ( r n C ll t n M h n t n fr th pr v n f Ot Material examined. l t p f f ll x, pl n n-t p x pl (AMΝΖ U M AC ND, AK, WO, BP, TO, HB, WI, WA, WN/NN, NC, SC, S b n O n d M l nt & S FD. Anth (O n d fl r l ( nn C ll t d O t b r— n rd d pr r l fr n r l v l 1 31; M p M t n fr rb n nv r n nt nt n (19 5 fl r l nn 175 (M l . rté-Sén tèr r p rt d th t b th d lt nd l rv ld f d n 1 15 ( n p rt (Anth . 1911b 7 d n d n pl nt nd n t p r th f n l h ph nd 19 3 3 5 n d n 1953 1 1-1 3 f 5 7 ( x- l f d n th r n f n t nd t C ll t n t n v b bl r ph n n K h l 199 5 r rd n l nd r fr pr h t rd n f ll x r n 1 93 11 (Anth . tt n 19 19 f t d nd p t r C n n th r nd n d n 19 3 3 5 195 9 ( n n nd r p n r h nh tr rd n p t nd p l f fl r l . pr n n n p dd t nf ld (AK (K h l 199 3 n th 3 1-3 7 n n d r d-br n; pr n t r n ; 5 h rl t n n l nd p n r n l tr th b l b nd r n r nd r r d-br n t t p f f ll x, ll t d t n p r r l br n; l r d-br n t r n p l r b ll ; t t l t r t h rt nd p r v r b d l n ; t Remarks. A. fl r l p r t d fr A f r r b v r h rt nd ppr d v r b d 3 l n t br d r h d l t r ll b d t rn l r n nd d nd pr n t r r t l t r f ntl n th pr n f t l b p n r th nt r r r n f pr n t d rd t ; b d t tp t r r r n f th pr n t It p l t n p th t pr b bl ; r n l htl nv r n b ll b f r br dl r n t d fr Afr r nd d t p r l n l th br d d n b l nd n- t t n; fr n th ll d t n t p n t r 3 p rt th l d n n n rr dl n ; fr nt l p l l Anth (O n d f r r (G z d t n t E l n v bl t b t n f t 3 ; M p 9 r th r x tr p z d l br d t t d t l f rth h r f r r G z 1777 7 (M l . 1911b 9 d l htl n v t f nt l t r l n tr t n j t n d n 1953 1 3 (Anth ( xt n v b bl r - nt r r t b ; h rpl d f n d b l tr n v r l ph nd n n n l d nt r rl t l t r l r n nt n tr t n; br d- t p rt f d th p r f l d n b p ; p n t r n th 3 1 -3 7 d r d-br n t br n; pr n t l r t th n h d r d-br n r d n t r n n r b ; l tr r n n El tr th d l htl nv x; h r pr n nt; pr x l f rth br n p ll ; l r n th p l pl t ntl nv x; p n t r l r r th n n f r b d p rt n f f r d r r; t t l t p r v r b d p rt p rt l rl tr n n f nt p tb l - h rt nd r t 3 l n ; t h rt nd ppr d pr n j t p t r r t pl t b n nd t n t v r b d l n t rd p x; t r fl n d th f n l n nl n d t l d nd pr n t r r t l t r f ntl b t th rd M t rn th l t r l d br dl nd t ll d t n tl n pr n t d th r n b h nd r l rl xp nd d n t r d b v l v l f br fl p r ll l; t p r l n l br dl r nd d t p t rn fr n d th h rt t 3 l n h ll d n n t h; d th d t n t d n p n t r ppr d t p t rn W n f ll d v l p d 3 p rt b t dl n n rr l p n t t ; fr nt - M l th l t v bl t rn t br dl r n t ; l p l t r d t n t E l r th v r h rt t t t l t r l n l l htl l n t G n t l t n v bl b t n f t n v ntr h lf 1 l n
— 1— r n t tr p z d l d t t d t l f rth l htl t n n tr t n t f v b v pr x th n- n tr t d l t r ll j t nt r r t b ; h rpl d f n d tr t n l d f n d l t r ll n t xt nd n nt d ; b l tr n v r l n l d nt r rl t l t r l r n d nv x; n t pl r l l h rpl d f n d b v nt n tr t n; d d n l p n t t x v r bl xt nd d p t r rl ; pr x l v t El tr th pr n nt h r ; pl t d t n t; p n xt rn ll l d nt rn ll p tb l pr n f nt; p n t r d p r nd l r r M t ll tr n t t p x El tr th p n t r th n n pr n t p rt l rl n p tb l pr n nf d; l n dj nt t t r xt nd n fr t ll pr r v l f nt r t rd p x; t r fl n d b f n t p x t r l t rn l r n n rl tr ht th l n x pt n pr x l th rd M t rn th l t r l l ht l t r l rv t ddl t r l r n f r n tr ht l n fr n f t l n j n t n p t rn th fr n f l n t v r n p - th p t rn W n f ll d v l p d r ; p r d pl r d v t xt nd n M l th f r tr h nt r n l t n r p x; l t fr x t h r l n l th d t n t f v v bl t rn t br dl r n t th r n t n th pr j t n nt b d v t n nv n t n fr r d tr ht l d t ddl th r p f l n r p r M x l v t p r t d b t rn t t l t r l n l G n t l nt rn l l n p n h h n rr l t t t rn d ll r t t rn t th n rr tr n v r v t d t l p t r r t p d t r b bl d r b d fr E r p t n f t x nv n t d f v th n b r n fr n t p t r l n n n t f t M l n t l th 9th t rn t - h p d th l t r l p l r rt l t d; ph ll b nd t n M t r l x n d nt - n n n-t p x pl d t n t th p n h rt r th n th t b n d t v ( AC l nd p ll th l t r l pr p l r f t AK, WI/M MC n t n C ll t d v b r-Apr l rd d pr r l fr n r l v l r S p nt l r n t d v r n th nd n p t h p d t h p nd n G nd n nt n nt nd p n tr t n t l t d xt nt r r d nt rth A r nd th rn E r It r d l r n d b th b n t n f t th nd r t n r A f r r v r l r t A. fl r l , b t th l tr th d p l n r r v t th p t r r r n f th pr n t l th ll nt r r b p th l t r l th p t rn th t nv n t d f v nd b- t rn l r n r tr ht nd l t l fr n nd r d b fr n f t fr th p t rn nd th th l n t l r v r d ff r nt l n th pl x tr n v r r v t th b f th f r t v bl bd n l r t r f nd n r th pr r n p r n fl r l . t rn t th t r b r d b p b n nd r l C p l t n nd t p ll f nd t d th v t - nv n t d n l f v n th r d bl d br n rb n nv r n nt Sp rld d r t p ll t d th t l f l tt r n f r t d r p r n h b t t ft n th n t pr nt G n S p nt C S p nt C 1 95 73 p - p Anth p b S p nt l nd n n p n rté-Sén tèr b b nt d n t n f W rn r (19 93 33; M p 1 d th t b r t t r t nd t t l t r "Anth l nd n " n ll t n ; pp r ntl r t t d ff lt t p r t fr t El tr n v r d r b d th h rt nd r t ppr d t bd b nt ft n n th - d h n n r n t p l br n; d r t d bl l l t r ll l tr th b d n b nd nd br d p l z n d th fr nt l p l l th n l n ; x ll r br n th d n b nd ft n nt rr pt d t t r nd n t p lp th nd nt l n t 3rd h rt nd n l r n r h n l t r l r n ; nt nn br n th b l nd l r n th d t t ddl th p x bl p l nt nn r r n E l r th l v bl rv d t r n b t n d th d th nv x; b l r n bp r ll l f t th fr p t r r h lf f l n t br fl b h nd th t p r l n l r nd d nd r n t th n p l ll r; b l tr n v r l v rt x l htl tr n t ; fr n th p n t r f n b r l n l d nt r rl t l t r l r n d n d nd d n l v bl ppr x 5 p rt b nt fr n rr
— — l n t d n l z n n dl n ; fr nt l p l t r b r l 19 (1 M r h 7 O t 197 W tt dr nd t d n rf l rl nd t d d r l n ; t t l nd (1 I M t h r p p I r p t tr p n— t tt r d; t b r t E th l r t A 197 (1 ; O t- 197 (9 ; O t- 1979 ( ; l n 1979 — b 19 (1 O r n 19 1 Sp ll r (3 ; r n t th rf nd t l r t th f h d M r 19 M K nz (1 p n r C ll 13 b t p n t r ppr x p rt; l t r l r n M r 1915 (1 ; 13 M r 19 5 ( t r n 1 v 191 v nl r nd d fr tr n ll r t nt b l n tr - r C ll (1 ; 5 Apr 19 M W C rt r (1 W h t n d t t d t l th rd; b l tr n v r l d p I 9 Apr 19 Sp ll r n b h (1 W dh ll 7 b xt nd n v ntr ll nt n tr t n h r p b n 197 t h r p tf ll tr p ( 9; 11 n p t r 15 n r r b r b th t nd pr n t l f v p t r ; 9 M r 197 ( 1 t h r C ttl rr r I El tr h t nfl t d; h r ; pl t 11 M r 197 d l l tt r (11 K r I v f nt; p tb l pr n b nt; p n t r n l tr 197 M r n l tt r (7 TO. n n n h l r r d t n t n pr x l h lf ppr x p rt 3 M 197 M M r ntl f ll n d rp f n r t rd p x; t t l t l n nd r t 1 p t ( . K l th r t t 19 M 19 1 l n ; t b r t t b t 5° 13 l n ; ppr d M S xt n fl x l tt r (1 W M r 1 O t 1919 nd r t l n M t rn th l t r l r- A C O C nn r (1 ; 1 A 1935 d n C ll (1 n l htl b d t rd W n r d d t tr p r h M 193 d n C ll ( t h th x th d n br h f t n p t r r 7 19 1 Sp ll r (1 W ll n t n t n l r n f 1 t h nd t r r M l th h nd t b G rd n O t 1 7 d n C ll ( ; n r n 1 b r n d n l n r p b n n f bl x v t n A 1 9 d n C ll (1 fr b f r ddl t p x f fl x r rf ; t rn S th I. NN. C bl p n n l ff M r t r d l l r t d br dl r n t t p x th l t A 19 5 W l r n nd (3 r ll Sp t r l n l b r n l n rv d t G n t l t n Apr 19 7 W tt M hl nb pl x n nd (1 th l t r l r f t n d t l h lf; nt rn l th l n 3 O t 19 3 G rl (1 h n [n h] h rt br d p n n d t l f fth rb rn (1 ; 31 M r 1937 nd 5 v 1951 G rl ( ; O t 197 W tt ( h r h 19 n 1973 p d t l t p l [AK] nf ld r p n K h l ( W W t Ev l l r h r v 9 S pt b r 197 G K h l l tt r n 7 /5 O t 1971 l tt r ( W t v n Inl t nd ( AC O t 19 I n nd l tt r (1 M h d Paratypes: 5 ( U M AC U IC U C M M b t n C n p (5 MC Chr t h r h 13 hr K n I Gr t I.: C t C p l tt r v M 195 tr p n r nd rth n r fr En - 197 K h l (7 9 v 197 (1 ; n l l nd (1 5 v 197 K h l l tt r ( North!. ND. K 1 M 1951 d n v - Material examined. p r pl 13 n n-t p x - t t n ( M h n I r I M r 197 K h l pl (AMΝΖ (3 ; n l I M r 197 K h l ft d l tt r nd r nd hr K n I / AK C O K W / ΝΝ pl nt ( ; n th r d t ( b h 13 1951 M MC A E r ( ; b 197 C t h r p t tr p nd r C ll t d thr h t th r ( Sp r t 17 n 19 l tt r (1 t rd d pr r l t r n r l v l 1 S p 19 C W tt l tt r nd n t ( Wh n r nd pr r l t h r r nd n t l r b W t rn ll 3 O t 19 r (5 Wh n r th f p n ll t d v r l l tr nl nd d O n h 7 195 W nd r M hl n Ind v d l h v b n f nd b n th t n l tt r b ( r I l t O t 19 W tt l tt r nd nd r d n r t n th r nd r p rt l rl t n ( AK. A l nd 9 M r 19 1 Sp ll r (1 ; Apr n n th r nd nd r G hn t f l nd A 19 7 nd r rd n r bb h ( ; v- b h xt r ( rn n b h r b t nf ld (AK (K h l 199 Gr ft n G ll 15 O t 19 1 Sp ll r (1 n r n 3 "Anth p 1" ( h r h A 19 1 M M nd r dr d n (1). nf ld r p n r v K h l 9 Remarks. S. l nd n r d l r n d b th A 197 (1 ; 7 197 h r (1 ; 15 n 1977 n rl p n t t h d nd pr n t th ntr t n r (1 ; 7 b 1977 (1 Mt Alb rt 191 r (1 ; r n p l nt nn r nd th l r l r n l r M 19 1 Sp ll r nd r t n (3 ; S p 19 p tt rn h n t n fr th n r pt n f Sp ll r v d ( Mt W ll n t n l v f ld 1 v M r f r th p
— 3— S p nt d tz n p p f S p nt . d f r tz ll t r f 17 3 ; M p 11 th t p r n th 9 -3 d r n ; l tr th b d n b nd nd p x br n S plnt b r rn ( r n n b n t n d nd pr n t th t d b nt d th 1 35; M p 1 b br dl btr n t ; b l r n br fl p r ll l t br dl r nd d t p r l n l ; d t n t th p n - b r rn Βr n 1 11 (Anth . tt n 19 t r n d ppr x p rt r d n t f n 1 9 1911b d n 19 3 3 5 p t r rl th th d l l n n fr n b r l nthr n r n 1 93 11 (Anth . tt n 19 nd t d E l r nd pr n nt; t b t n f t 19 1911b 33 d n 19 3 3 5 n- l n n r n t d t t d t l th rd; l t r l nt b l n- n th 3 7 - d d r br n th t l tr l tr t n f bl nd t d; d r l p n t t nd l htl r n ; pr n t t p l r p t r r t nt b l r r t l t th p n t r ppr x 3 p rt; l ; l nd nt nn p l br n nt b l l d t n t v r d b t n r f l t r l d nd pr n t th t d b nt; t t l t f v p r nd r t; p n t r f n d th l t r l r n El tr bp r ll l; h r pr n nt; pl t ; p t r r t br fl p r ll l t br dl r nd d t p- p tbas l pr n nd t n t; p n t r ll d f n d n r l n l ; b br dl btr n t ; p n t r n d pr x l h lf ppr x 5 p rt r p t r rl ; p rt; t t l t 1 l n ; t 1 t t l t 1 l n n rl p rp nd l r; t l n E l r ; rv d l r t v bl n p t r r l htl rv d r d t b t 35° 11 l n ; h lf f l n nd r t ppr x l n d b nt nd v r r n t d t t d t l th rd; p n t r d ff lt t n n p d r t d bl l l t r ll t l t r ll x pt n r b ; nt b l l h rpl d f d b- M t rn th l t r l r n l htl b d t- r d b t n r l t r l f v rd W n f ll d v l p d El tr l n t bp r ll l; h r l n l pr n nt; M l th d t l h lf f h nd t b h ll l x v t d pl t n t v d nt; p tb l pr n f nt; p n - n nn r rf th d n l n r t p b n n t r l r nd d t n t n pr x l t -th rd x v t n; t r th h rt fl t p b nt z n t p rt r d n t f nt p ll ; t t l t l n t p x; t rn h rt th p x l htl r n t nd t n l d ff lt t p r t 1 l n ; t G n t l t n th p r ll r nd l d tr b t d r d t b t ° 1 l n ; nd r t bd - t n l t r l r n n t f r n d r t r ; nt rn l b nt d r tl l t r ll n d l n M t rn l n p n th l t r l r n d t n tl b d t rdl W n f ll d v l p d Type data. Holotype: l [AK] n rth f r th x th 1 t h nd t r r b r n d n l 197 tz ( AC br h f l t r ll d r t d t n p t r r margin. M l Paratypes ( f l ; AC U C North I. AK. th t rn t br dl r nd d to barely subtruncate. S d t h l t p (1 ; d t x pt n rth f G n t l t n th d n row of setae on lateral r l 197 (1 r n n d t l th rd
Material examined. p r nl Type data. Anth b r rn : lectotype male lab-
AK / — ll d "7 5/ [C ] r " ( r n C ll., ΒΜ Η n l C ll t d p n n l p r l t t p th l b l plus "Anthicus obscuri- rd d fr n r l v l cornis" ( r n C ll BMNH), here designated by FGW. Anth nthr n : lectotype (sex undetermined) Remarks. S. d tz d t n t n t br d h d r l l b ll d " 73/ [BR] Boatmans Reefton/ Anthicus anth- p n t t nd r r t l t pr n t nd n r ll r- r n " ( r n C ll BMNH), and paralectotype with n l r t n It h l n n nd n t h b n f r t l b l ( r n C ll BMNH), here designated ll t d nl n th A l nd r p t th t t n b GW ntr d d p In p n t t n t t n nd n r l f r t h l r t n b r f A tr l n nd Material examined. Type specimens, plus 132 non-type Or nt l p th n t th th r thr l nd ex pl (AM Ζ, U , M , AC, U IC, U C .
-2424--- AK WO WI W WA / S C tr n t t p x G n t l l t r l r n f t n th W O S l n r f t n d t l th rd; nt rn l th h rt C ll t d thr h t th r br d p n n d t l h lf rd d fr l v l t h h r l v t n M t nl ll t d t n ht n l hn nd p d t t t p f l th nt r r p n nd th r pl nt ; d r n th d f nd nd r b r nd t n p r l t t p f l th p t r r p n n d bl rd nt l b ll d "7 / [AK] " ( r n C ll r S. b r rn d t n t n t d r l r M h r d n t d b GW br dl btr n t h d b nd l t r ll d r t d l tr l nd r t G K h l ( n l tt nd t th t th p M t r l x n d p p n pl 1 n n-t p t d th v t t n f n t p S. x pl (AMΝΖ AMC CM C U MC C M p ll d p , b t th t pp r ntl d n t r t th r AC U IC U C AK C WO G K WA W / S DN. C ll t d thr h t th r. S p nt p ll d p ( r n n b n t n rd d fr l v l t 1 19 3 3 ; M p 13 nd pr r l n l f l tt r r n f ll n d d tr f rn (C th d ll r d r n th d nd t n ht - p ll d p Βr n 1 1 (Anth . tt n 19 nl ll t d n f l f v r pl nt n l d n 1 9 1911b d n 19 3 3 5 l hn pr r . At nf ld (AK r r l l- n th 3 15-3 d l n t h n n ; h d nd n nt r d n n t h r n t pr n t r n t br n; l tr r n t p l br n b r n , G hn t f l , G. l r , C r x p nd th br n d n b nd nd p rl d f n d b l b nd; Un n p nd t p rt l rl b nd nt n b l b nd nn t d t d n b nd l n d nd th S th A r n C rt d r j b t r n n th b h n ll l n t r ; d n b nd t d l (K h l 199 3 nt rr pt d t t r nd th r l r t v l r n p t n d t l f rth f l tr ; d r r p n th p l b nd r S. p ll d p d t n t n t r l r tr n l nt br n h v p l r r b r l pp r nt; h d b l r p tt rn nd l n r t t t l t n th l ll h t r n th t b nd t r d n d r l tr nd v d l ; nt nn r n d nd pr n t th t d b nt nd t t l t p r r t d th b r l r l n S bf l AG IOI I AE t p r l n l ; d th f n tt r d p n t r 5 p rt b t n rr l b nt t dl n ; t t l t 1 G n r d r r & G r n l n ; t 1 l n E d r t l pr n nt r d r r G r n 1 3 l r 19 9 th p t r rl rv d t 3 l n t j n t r f p p n t d n t d f t n p t r r h lf r d f th r ( p ll n 1 7 5 tt n r n t th d d t t d t l th rd; d l t 19 1 d n 19 3 3 v nl nv x fr ll r t nt b l n tr t n; p n - t r l d f n d n d t l h lf r d t n t t rd d l htl n tr t d j t p t r r t l n b ; nt b l l d t n t b r d b t n r d t n t n ; fr n fl tt n d; nt nn l b xp d; l t r l f v ; pr n t l ll r th l t r l l n f t nt nn r 1- n rr nd l n t th p l 3 n- l n b l r d th p n lt t tr n v r ; fr nt l p l l El tr th h r l n l d t n t; pl t f nt; b nt E l r th h rt t b t n f t M x- p tb l tr n v r pr n b nt; p n t r l r ll r p lp th nd nt l n t 3rd h rt nd nd d t n t n pr x l t -th rd r d n t nd t n t tr n l r th l r nd bl l r f r p ll p rt n r b ; t t l t p r 1 r n t d t t nt r r f rth rv n br ptl t l n ; t r d t b t 5 ° 1 l n ; nd r t p x; r n l htl nv r nt t b th f nt bd b nt 5 l n d r t d bl l l t r ll l t r l n tr t n n r b ; d h ll l rv d; l n r t r M t rn th l t r l r n l htl r n t n t l t r l r n f b br fl n l d nt r - b d t rd W n f ll d v l p d v ntr ll ; l n n th r v ntr l t r l l ; l n p t th x th 1 t h nd t r r b r n h rt br h r d pl d r l t pr x ; l n t r t rn l l f t n p t r r r n M l th t rn br dl r x l v t p n xt rn ll l d nt rn ll
— 5— El tr th tt r d p n t r ; t t l t h rt; r All l nd r d r pl d nd r nd r t b nt M t ll n rr l r nd d t br n , th h th r n d r bl v r t n n th dth p x M t rn tr n l r th tr ht l t r l r- l n th nd n l f th l tr l t t l t nd n b d z n ; d n pr j t n p r t n x bl ntl All p n fr rthl nd r nt r l ll Sp - t n tr n t xt n n f t t rn M p t rn n fr n r A l nd n l d f p l nd v d l th l t r l r n tr ht M p r l r n lt - b t n r br n v r ft n th p l r p t r nd th t t r r l n t l b l b d Abd n th f r t t n p n t r th t b r th t t l t n th l tr h v bl t rn t f d th r nd r fr G n t l 9th p l p t n b nt r nn t d t th xt nt th t t f t rn t Y- h p d th p l r n t rt l t d; t n th rf p l r nl th t r br n h In th nd ph ll b d t n t th f th rth I l nd nd th l n r ll r br n th th p l p t n th l tr f ll r d t r nd r M b r f r d r n A tr l r th r d ll x pl fr th ll r r r v r l nd nd t p r t S th A r h r th r d r th nl f bl nd t n f th p l p t t d th t l nd d n t l t r v r l n n rthl nd l n thr h t t f th r t f th r n b t h rt n th ll r x pl h l tr l t r t th r r t t n l n rthl nd n rl ppr d v r th r t f th rth r d br n I l nd nd n th ll r r b t nt r d t n l n 37; M p 1 Sp n fr rf l I l nd l pp r t b th p r n 1 7 5 ( r d [ ] tt n 19 1 d n 19 3 3 l r 19 9
n th -5 5 d r n th h d r dd h r S bf l EMO I AE h d nd pr n t br n th l tr ttl d ll nd br n; d r t p n br n th l tr l b p l r; G n C t Sh rp l nd nt nn ll t r n ; nt nn l l b - C t Sh rp 1 77 9 p - p C t v t t Sh rp b t d r r n t r d t n t v r d r nd v nt r r n l n t p t l t p r h rt r t; t h t ppr d t d b nt v r b d d th b h rt br dl btr n t t d t l n t r n h d nd pr n t p rt E r nd d p t r r t ; fr nt l p l l br d; d th t b t n f t 1 l n El tr l p n t r l t r ll th h ll l n t d n l pr n xt nd n 7 p rt n r b ; t t l t n l tr 5- 1 fr ddl f t n r nt nn l b E l r l n ; t - 1 l n pr n nt ll th v bl t r n t n b t n f t Ant nn l n r l t v l t t M x ll r p lp p d t t t p nd p r l t t p l b ll d " th nt nd 3 b r n - l n t nt 3 [C ] r " C ll t n ( M h r d - n l t v ntr ll nd nt n rr l r f r n t d b GW r pt n b d n f v p n fr r th r x th th n r d r t p x l n " b h t r " f h h th t r ll d t n t ll r; nt b l n tr t n d p l t r ll ft n th th l tr br n h n th d n th d t l h lf; n nt n d h ll l r d ; l r p t t v ntr l p n h n nd t n f p l p tt n n th br n h r n f n tr t n d r l t pr x ; t r t rn l r th r p n h v b n n th th p rt - l pr nt nl d r l t pr x ; d thl l r l r p tt rn nv x; pr x l v t l d nt rn ll p n xt r- n ll M t r l x n d p p n pl 1 3 n n-t p El tr th p n t r b r l t r l n pr x l h lf x pl ( U MC C AC U C U IC th nd r t n dd t n t t t l t nd t ; f n AK C WO G K WI/ MC S l n l n l tr l t r d t n t n d t l h lf S t ll SI / Ch th I l n t br dl r nd d t btr n t t p x M - C ll t d O t b r—M t rn th l t r l r n l htl rv d l t r ll rd d nl fr n r l v l M p t rn n rr l t n d ll nt r r t Str n l t d th t l nd d n n b h t rn M p r l n t p r ll l- d d r h- C nl f nd b n th d br n th b h r n nd n ddl x l v t ; p t r r r n th fr n f d n r h rt t M x p r t d b t rn h h
— — narrowly meets median point of metasternum. Sternites row of 3 setae laterally near narrowly pointed apex. Fem- free. ales with last sternite broadly rounded. Male genitalia 9th sternite V-shaped, with apical arms not articulated; phallobase absent; penis extremely long Type data. Described from "[ND] Pakarau, North Auck- and thin, 2-4x as long as tegmen; tegmen with lateral rows land." The holotype could not be found in the Broun of setae near apex. C ll t n ( M b t th Βr n C ll t n n AC contains one male without a label, which is probably the Remarks. This precinctive New Zealand genus is very holotype, and another specimen labelled "[WO] Ohaupo / close to the Australian genus r h n n . Members of 545 / Cotes n. sp." C t usually have the lateral constriction of the pronotum continued as an impression across the dorsum near the Material examined. Probable holotype and 9 non-type base, the mesoscutellar apex broadly rounded to subtruncate, examples (AMNZ, NZAC, UNHC).
and the pronotal disc convex. Species are generally quite ND, AK, CL, WO / — distinct and easy to recognise, but the three species forming Collected January, February, and May-July. thevestita-group pr b , rl , and v t t — r much Recorded from sea level to 100 m. more similar in appearance, and difficult to separate. Found in leaf litter of forest sedges such as C r x, The various species of C t are invariably associated Un n , and S h n in bush areas at Lynfield, AK with vegetation native to New Zealand, and are found in (Kuschel 1990: 34, 66). leaf litter or on fallen dead trees in the bush. Remarks. C. b ll t is closest to C. r p in its deep pronotal constriction that continues dorsally across the C t b ll t r n disc, small size, and relatively long, suberect elytral pubes- Fig. 7, 38; Map 15 cence. It is distinguished by the distinct humeral angles of the elytra, elytral setae raised to about 40°, and lack of dark b ll t Βr n 19 3 9 (C t . Hudson 1923: 385. elytral bands. Length 3.20-3.60 mm. Body orange to red-brown, shining; head and pronotum sometimes darker; legs orange; elytra of most specimens with a smooth, raised, yellow mound C t r p ( r n near lateral margins of postbasal impression. Fig. 3, 39; Map 16 Head and pronotum with disc polished; setae subdecum- r p r n, 1880: 412 (Anth —1893: 1165 (C t . bent on head, decumbent on pronotum; tactile setae sparse; Hutton 1904: 189. Pic 1911b: 23. Hudson 1923: 385. punctures fine and sparse. Head with base shortly and p n t t Broun, 1893: 1165 (C t . Hutton 1904: 189. Pic broadly rounded; discal impressions between eyes faint, 1911b: 23. Hudson 1923: 385. New synonymy. oval. Eyes with setae between facets 0.03 mm long. Pronotum widest at distal third; margins curved to a Length 3.40-4.20 mm. Head and pronotum orange to red- deep, smooth lateral constriction at proximal third, this brown; elytra orange, with base and median band brown; continuing across disc to a depth of 0.04 mm; base slightly distal third obscurely darker in some specimens; legs narrower than greatest width across disc; tergosternal orange. sulcus faint; pit at ventral margin of constriction outlined Head and pronotum with disc polished; punctures fine, by setae. sparse; setae subdecumbent on head, decumbent on Elytra with humeral angles distinct; omoplates well pronotum; tactile setae long. Head with base briefly and defined; transverse postbasal impression weak; a series of broadly semicircular posterior to eyes; punctures on disc deep punctures in proximal third 0.09 mm apart, grading to 0.04 mm apart; lateral discal impressions between eyes fine posteriorly, absent or sparse on omoplates and on faint, oval. Eyes with setae between facets 0.02 mm long. raised lateral mounds in postbasal impression; tactile setae Pronotum widest at distal fourth; margins curved to deep erect, 0.18 mm long; setae bushy, suberect at about 40°, constriction at proximal third; punctures dense and coarse 0.14 mm long; undersetae 0.07 mm long, directed ob- in constriction, which extends across disc as a saddle to liquely near suture from base to postbasal impression, 0.04 mm depth on dorsum; base narrower than maximum otherwise directed posteriorly. Wings straplike, shorter width across pronotal disc; tergosternal sulcus faint; pit at than elytra. ventral margin of constriction glabrous. Males with last sternite broadly rounded apically, with a Elytra narrow; humeri obsolete; omoplates and postbasal slight truncation at middle. Genitalia: tegmen with a short impression faint; large serial punctures distinct in proximal
-27--27- th rd r d n t nd t n t p t r rl ; t t l t r t El tr th h r l n l d t n t b t ll; pl t 1 l n ; d n t b r t t b t 3 ° 1 nd p tb l pr n f nt; d fl tt n d t ddl ; l n ; nd r t bd b nt 1 l n l htl p n t r r l nd d t n t n pr x l th rd p r nd rl d l t r ll n p tb l pr n W n tr p-l nd t n t p ll ; t t l t r t nd b r t n lt r- h rt r th n l tr n t n r l t r ll fr l tr l t r 1 l n ; M l nd f l th l t t rn t br dl btr n t t d b nt t -3 ° 7 l n ; nd r t t p x G n t l t n th l t r l r f b t t ppr d 7 l n W n l htl r d d n r n r p x h h t p r t bl nt p nt p x Abd n l t rn t th d t n t l t r l p t h f r d t p d t Anth r p : h l t p ( x nd t r n d M l th l t t rn t br dl btr n t t h ll l l b ll d "7 7/ [ ] Ρ r / C t r p " (Βr n C ll r n t G n t l t n th l t r l pr p l l b ΒΜΝΗ fr n d th t l th l t t rn t br dl C t p n t t : l t t p f l l b ll d " / [AΚ] r nd d / C t p n t t " ( r n C ll M h r d n t d b GW p d t l t p l [ ] ll 5 [75 ] 5 br r 1957 E S G rl ( AC M t r l x n d p p n pl n n-t p r t p ( 1; AC U IC U C S th I x pl (AMΝΖ A IC U M AC U IC C n n 17- n 19 9 r (1 Mt Arth r 3 U C [9 ] M r 1935 G rl (1 ; 3 -35 [9 -1 5
AK WO I WI WA W / — ] 13-1 19 1 n nd W d ( ll C ll t d thr h t th r 1 v 19 9 O C nn r (1 ; 5 [75 ] 5 b nd 7 rd d fr l v l t 335 M 1957 G rl (1 ; 19 b 1957 G rl (3 C nl f nd n r l f nd br n h l tt r n th Gr th 19- 1 n 1957 G rl (1 f r t fl r nd ll t d n l r n b r fr nb t d 35 [1 5 ] -1 1957 G rl ( p tf ll tr p At nf ld (AK f nd n l f l tt r b n th h r t n x, A t l b n , nd G hn t f l M t r l x n d p r pl 3 n n-t p x - (K h l 199 3 pl ( AC U IC U C —/ r C. r p p r t d fr C. b ll t b t C ll t d v b r-M b l t l tr l h r l tr l t t b t 3 ° nd rd d fr l v l t 11 ; t p n t l tr th br n b nd d n b nd r n nd t d 75 -11 n h r n l d r pt n th t th p n d f r h nl b tr t nf r t n r rd fr th ll t r Mr A Cr p " th f pp." t l n t n l r (U IC pr b bl fr l f l tt r
C t rl n p r C. rl t l r t C. pr b ( ; M p 17 r nd r pr b . It r d l r n d b th d d dl nv x pr n t l d br dl btr n t h d n th 1 -7 5 d ll t br n n- b nd th d t n t r t t b r t t t l t n th ll th h d nd pr n t d r r th n l tr l tr l d th t r t l n th d b nt t d nd pr n t th d n bd b nt t d - d f r E S G rl h ll t d h f th t p b nt nd r t nd d b nt t b r t t r t th r ft n f r n rl d p tt rn ; t t l t d - t n t r t d th b h rt br dl btr n t ; d th n t b r p n t r ; v rt x b l n p t r r C t h ll n r n t ; nt r nt nn l pr n d t n t E th t t ll tr t d; M p 1 t j n t n f f t l n r n t th d tr n l nv x d t n tl x d- h ll n r n 19 3 9 (C t . d n 19 3 3 5 n pl n f l tr l d ; d d t t d t l t -th rd ; n th 5 1 -5 1 d r n ; l p l r p n t r n t pp r nt; l t r l n tr t n nt n d d d ll n pp r n ; t d n b r t; t t l tr n l r d d p n d r ; p t t t h rt r t; l t r l r n f b l n rl nv r- v ntr l r n f n tr t n b r d b t nt t br dl r nd d d n n l t n; p n t r n
- - d p r h ll b t p rt; l t r l d l rv d p t r r t ; fr nt th; d th tt r d pr n b t n l n t xt nd n nt r rl p n t r ll nd d t n t p rt d n r n n t b t n nt nn l b E th t b t n f t r j t p t r r t b f nt nn d l n t d d ll l n b n bl h ll d pr n; n th p n t r r n t d t t d t l th rd; d d t n tl r - h l r r r t r f r E l r ; t b t n f t r t l t ; p n t r n r b r t r f r ppr x 1 l n p rt l r r nd d n r th n n h d; l t r l r n r n t th d n f n p n t r d t t d t l r nd d t d p l t r l n tr t n n r b th n- f rth; b l htl n rr r th n x dth r t n n r d b r l p r pt bl r v ppr x d ; l t r l n tr t n v r d p j t p t r r t d- d p t l t r l p rt n r l p n t t ; p t t dl ; d n l t r l pr f l ntl nv x fr d t p rt t v ntr l r n f n tr t n b r d b fr n f t n tr t n th n fl tt n d p t r rl t b ; l br p t El tr f ntl r r t l t bp r ll l p t r r t b v pr x th rr nd n fr n f t b l t h r ; d ntl nv x th n nd t n f El tr ll n t ddl l n d t n t h r nd p tb l tr n v r pr n r pl t ; p n t r th t p tb l pr n r pl t ; l t r l r- d t n t n rl t p x tr n t nd n r n pr x l n v nl nv x; p n t r l r r th n n pr n t b t h lf 9 p rt n r b ; t t l t r t 1 d t n t nl n pr x l f rth h r b r l nd 9 l n ; t d b nt t b t 35° 1 l n ; p rt n r b ; t t l t r t l n ; t nd r t ppr d l n ; t nd nd r t d b nt t 5° l n nt r x d th nd r- d r t d p t r rl n rl n f r v r nt r rf t ; nd r t 1 l n l htl r d pr d W n r d d t tr p h rt r th n l tr th n t ; p b n l htl rl d bl l n pr x - M l n n n l th l t t rn t br dl l f rth nd d t l h lf (p tt rn b t n b v n btr n t t p x l ht n W n r d d t tr p th l n t d n l v n nd n xp nd d p t b t l p t h p d t l t p l b ll d"[W ] r nth 5 11 191 / M l th l t t rn t n rr l btr n t t p x C t h ll n " ( r n C ll ΒΜΝ l n l ft nt nn G n t l t n l n t thl n rr d t bl nt b nd 5th nt p x th l t r l r f b t 1 t n r p x - l th l t t rn t l r t l b t n rr l r nd d M t r l x n d l t p pl 1 n n-t p x pl t p x ( AC W /S p d t C t pt : d r b d fr "[AK] " C ll t d n v b r h h l t p h ld b n M v r th r n h n n-t p p n ll t d fr l f l tt r C ll t n ( M nt n nl n l l b ll d " 7/ [Β ] p r / C t pt " S n th l l t d n t r C. h ll n t d t n t b th d n rr p nd th th n p bl h d n th r n l d r p- r t r f r p n t r n th pr n t nd t n t nt n - t n GW d d n t r th p n h l t p t n f th nt b l n tr t n r th d l tr l C t pr x : h l t p l l b ll d " 71/ [C ] t t 35° nd nd r t ll d r t d p t r rl M h n / C t pr x " ( r n C ll ΜΝ
M t r l x n d l t p f C t pr x , pl 1 r n C t pt n n-t p x pl ( U M AC U C 1; M p 19 AK C WI W /— C ll t d n ll nth x pt A t pt r n 1 93 11 5 (C t . tt n 19 1 9 rd d fr n r l v l 1911b d n 19 3 3 5 A t d th d d nd h ll d- t tr nd n pr x r n 1 93 11 7 (C t . tt n 19 1 9 h ll M tr d r x l (K h l 199 3 n 1911b 3 d n 19 3 3 5 n n d d A th tr l , n f ll n tr tr n nd r tt n n th 3-7 d nd prothorax r dd h br n; t p nd nd r l ( r n 1 93 11 5 l tr br n h d n t r dd h br n n r p x; l r n r C. pt d t n t n t l r z l f d nd pr n t th t d n d b nt; t t l tr n v r r v d r l t th l t r l n tr t n n th t d t n t r t d th b h ll l nd v nl pr n t l d nd th d n f n p n t r n th pr n t
– 9– C t pr b r n C n n f l f A t l b n , nd n v t - ; M p t n d r n th n ht t nf ld (AK (K h l 199 3 C d r l . Al ll t d fr p d rp-br dl f pr b Βr n 1 1 91 (C t . tt n 19 1 9 f r t l tt r l f l tt r nd n n p 1911b d n 19 3 3 5 d r l r n 1 93 11 (C t . tt n 19 1 9 r C. pr b d ff lt t p r t fr t th r 1911b d n 19 3 3 5 n n p h h t th r r h r d n t d th v t t n th 5 - 7 d t n t r n th l t r l r p. It h r th C. rl th r t nt n f th r n f pr n t nd n ll nt r b br n; h r l n l f th l tr nd th l t n th l tr th v bl n br n p t t ddl n d pr p l t l t r l l b C. pr b h th pr n t l nd l t r l r n d r r n r b nd l t r l t d l d fl tt n d nd n th pl n th l tr l d th r ; l ll h d b h ll l nd v nl rv d b t n th d nd pr n t th d n nd r t ppr d t nd th l tr l t nd t t l t l r n l n th th n rr l bd b nt nd t d n nd bd - th t t l t ll n n p n th d Oft n t b nt th t th r pr d n rl d p tt rn n t h ntr l br n p t n th l tr nd th pr n t l b p n ; t t l t p r r t d d ll n pp r- d r r th n th l tr n f ntl r r t l t n d l n d t n t p n - t r ; b h ll l nd v nl rv d; nt r nt nn l pr n rv d d t n t; v rt x b l n p t r r t C t r f r n ; t b t n f t l n 9 3; M p 1 r n t th d n rl fl t n pl n l tr l d d t t nt r r f rth; l t r l n tr t n nt n- r f r n 1 93 11 7 (C t . tt n 19 1 9 d r d d t n t pr n 5 d p n 1911b 3 d n 19 3 3 5 d r ; r r t l t n d t n t n d ; p n t r n n r n 191 3 (C t . d n 19 3 3 5 h ll l nd p rl d f n d p rt; p t t v ntr l n n r n f n tr t n b r d b t n th 5 -5 d br n; l tr th br d El tr th pl t f nt; p tb l tr n v r - ll t r n b nd n p tb l pr n; l tr l pr n ; h r l n l h rt b t d t n t; d t br n t b ll n b nd nd br n n d t l fl tt n d; p n t r tr n t n pr x l th rd h lf th n rr tr n v r b nd f h t p b n t f nt t p r pt bl n d t l h lf; t t l t n rl d t l th rd r t 15 l n ; t b r t t 3 ° 1 l n ; d nd pr n t th p n t r n t pp r nt; t t l d n nd r t ppr d t bd b nt 7 t l n r t d b btr n t th br dl l n d r t d l t r ll n r b bl l l t r ll n r nd d v rt x l n l ; d th d t n t l t r l l n t - d t l t -th rd f l n th W n f ll d v l p d r d n l pr n d l t nd nt nn l b ; n rl Abd n l t rn t th r l r p t h f th t b t n f t b t 1 l n ; n r d t n r l t r l r n r l p n t t M l th p l r n f l t t rn t br dl tr n- r n t th d d t t d t l f rth l htl t G n t l t n th l t r l t l b n r p x n rr r th n b ; r n r nd d t d p nt b l l th l t t rn t br dl r nd d n tr t n; h ll n tr t n d t n t r d 3 d p n d r ; p t b v pr x n l d b p d t C t pr b : l t t p l l b ll d "1 15/ t ; t l t r l n d d r t d t rd ddl f [W ] W ll n t n/ C t pr b " ( r n C ll M nt r r l b t ddl d r t d p t r rl h r d n t d b GW El tr th l t r l r n p r ll l; h r pr n nt; C t dorsale: l t t p l l b ll d "[AK] Cl v d n/ pl t nd p tb l tr n v r pr n d t n t; 9/ C t d r l " ( r n C ll M h r d - p n t r h ll nd r l b ll nd t n t n d t l n t d b GW h lf p rt n r b ; t t l t v r bl n l d t b t 5° 13 l n ; t d n ppr d t M t r l x n d p p n pl 1 n n-t p l htl r d 1 l n ; nd r t ppr d 9 x pl (A IC U M AC U C l n ; p b n n h r d r t d l t r ll ; d n AK O Ι W / ΝΝ b nd th t n d d r t d l t r ll t ddl f C ll t d v b r—M l tr nd fr th r t l t r l r n th t d r t d rd d fr l v l t 75 ll th bl f t t n t f r p t r rl
—3 — r nt d l n ; h t p b n n d t l th rd d r t d d p n d r ; d d r t l r nd d x d n bl l t ddl f d W n f ll d v l p d pl n f l tr l d ; p n t r h ll p rl d f n d th x th l t t rn t br dl r nd d G n t l b t p rt; p t t v ntr l r n f n tr t n t n br ptl n tr t d n r p x t f r n l n t b r d b t p nt th l t r l r f b t 1 t n r p x El tr l n h r l n l pl t nd p tb l pr n; d h ll l nv x; p n t r ll r l Type data. C t r f : d r b d fr " t l l n r d pp r n n d t l th rd; t t l t r t n r ll [W ] W ll n t n " h r n C ll t n ( ΜΝΗ n- p r n d 1 l n ; t b r t t -3 ° t n n l f l l b ll d " 7 / [C ] M h n / 1 n n l n ; d n d b nt nd r t n f nt C t r f " h p n h 7 nd nt nn l nt r nd rl thr h ddl p rt n f h l tr n p nh l n th r ht l tr n nd v r r bb d h l n Abd n l t rn t th l t r l r l r p t h M h n l b l pp r d nt l th th n n th f r d t nd t n t h l t p f C. pr x r n (1 93 h h l M l th p x f l t t rn t br dl btr n t fr Mr S nd r h nt th p n t r n h G n t l t n th pr p l r n tr ht b r n p n pr b bl th h l t p b t n t l b ll d h rt l t r l pr p l r f t l th l t b GW t rn t br dl r nd d C t n n : h l t p f l l b ll d "3 5 / [W ] K t W ll n t n/ C t n n " ( r n C ll M Type data. C t v t t : h l t p ( x nd t r n d l b ll d "C t v t t p S [MC] rt n v Material examined. p p n pl 7 n n-t p 73" n th rd th th p n ( M x pl ( M AC C t d t n t : t p l l t n t v n h n l C RI, WA, WN / BR. f l (? p n n th r n C ll ( M l b ll d C ll t d n v b r nd n " 7 / [W ] W ll n t n/ C t d t n t " pr b bl th rd d fr n r l v l h l t p h b t t nf r t n n l b l Material examined. p p n pl 39 n n-t p Remarks. C. r f n t bv l l t n th r x pl (AMΝΖ U AC M U C p It d t n t n t b n t n f d n t nd ND, TO, HB, RI, WI, WN / SD, NN, MC SC nd r t th l r p tt rn f r d b th t nd C ll t d v b r—M th 1 r rd n A t th ll d r t d t f th l t r l p rt n f th rd d pr r l fr n r l v l b t f nd p tπ p tb l pr n b t t nf r t n p r th f f r t nd fr ; n p n ll t d b l ht tr p n v b r C t v t t Sh rp Remarks. C. v t t th t nl ll t d ; M p p f th v t t r p, nd v rl p n r n th C. v t t Sh rp 1 77 9 (C t . r n 1 11 tt n pr b n th th rn h lf f th rth I l nd nd th C. 19 1 9 1911b 3 d n 19 3 3 5 rl n th n rth rn S th I l nd It p r bl fr d t n t Broun, 1 93 11 (C t . tt n 19 1 9 th t p b t l f l tr l h r nd th 1911b d n 19 3 3 5 n n tr ht pr p l r n f th l t n n th 7 - d t n t r d-br n ft n th h d nd pr n t br n d nd pr n t th d n ppr d nd r t G n r h n n l b rn nd d b nt t bd b nt t ; t t l t p r r h n n l b rn 1 91 3 1 p - p r t t d t n t n p n d pr d nd r h n n v t r n l b rn b r n l n n p n th r d th b br dl nd n t p h ll l rv d; d th n t b r p n t r ; v rt x b l n p t r r t ; nt r nt nn l pr n d th v rt x h rt p t r r t br ptl n- t nd t n t; t b t n f t tr t d t br d n ; fr nt l p l t r r n t l n pp r nt; l p fr nt l r nv x th n l n t r n t d t t nt r r f rth; l t r l n tr t n bl l t r l pr n l t nd p t r r t nt n d tr n l r d t pr n nt nn l b E l r th rv d t d t n t n
—31— p t r r h lf M x ll r p lp th nt 3 n l t l ; p n t r n d ppr x p rt; t t v ntr ll ; l t nt l r r f r Ant nn r l - j t r f f t r t ppr x l n t v l t t r n t th nt b l n tr t n d p l t r ll r th r x th n b r th n pr d r t p x nd n v ntr ll n p t b r d b t b v th l n ll r nd l n tr n v r nt b l l ; p n t r -fr pr n; d th d n l n t d n l pr x l v t p n xt rn ll l d nt rn ll ; t r- d pr n; p n t r ppr x p rt t rn l l d t n t d r l t pr x xt nd n t El tr th r n n rl p r ll l v l r n n r pr th r b n p l rl xt nd n t p tb l r ; h r pr n nt; d ntl nv x; pr n t l p x; nt b l n tr t n d p l t r ll nd pl t f bl nd t d; p n t r l r nd r l n xt nd n t l r p t b v pr x n t nt n n l tr l d r b t d t n t t rd p x r d r ; d br dl fl tt n d t l htl l n t d - p rt n r b ; t t l t br tl n 1 l n ; t n ll pr d b r t t b t 3 ° 1 l n ; nd r t b- El tr th p n t r r l r n rl x pt n d t l d b nt l n W n f ll d v l p d th rd th nd r t n dd t n t t nd t t l t ; M l th l t t rn t br dl r nd d; f r t r f n l n l n t r d t n t n d t l h lf W n f ll l htl xp nd d th 1 t t r r th nt r r p l d v l p d S t ll br dl tr n t t r n t n l pr d d G n t l t n br dl nd n rr l M t rn n rr l d v d n x br fl f r d b l b d t p x th r f 3 r t n l t r l r- t p x t n d n p nt f t t rn ; l t r l n n r p x; p n tr n l rv n v ntr ll nt r r t r n f t rn tr ht M p r l n t t n b l th l t t rn t br dl n l t t br dl nd h ll l r n t n p t r r r n p x; nt r r t r th 1 t t r r n rr r t nt r r th h rt fr n f t St rn t fr G n t l 9th p l n l dr t t rn t Y- h p d th p l r n t rt l t d; ph ll - b b nt; p n l n t xt nd n nt r rl b nd Type data. Holotype: l [AK] nf ld r p n t n b b t t p ll t n r th n h lf l n th f r v b r 1975 G K h l ( AC t n Paratypes ( 1; AC U IC U C North I. ND. r n n r 19 n 19 3 G W l tt r (1 AK. Remarks. r h n n l r n f nd pr r l A l nd C x Cr n 19 7 E rb rn (1 ; 3 b n A tr l At l t t p fr Ch l pp r t 19 3 D. Spiller (1 ; 9 v 19 9 At n n h (1 b l n t th r p d ff r n nl b th x b n nf ld r p n r K h l M r 197 ( ; v l r l nt th th nt r r th rd f th r l n th 197 (1 ; 197 (1 ; 7 197 (1 ; 1 v 197 p r t d b th t rn h n l p f nd n pt p r p r ( n 1975 ( ; l nd pp r t b n ntr d t n fr 1975 (1 ; 13 M r 197 t l ht n b h (1 ; 13 M r 1979 S th W l h r th n t b n d r b d r h n n on Salix fragilis ( . I Ot t I.: 1- v 1977 p r t d fr C t b th h rt p ll tr n t t tz p t tr p (1 nd n r tt l tr p n b h (1 ; r n t t ll th pr n t l d th v 11-1 1977 W tt p t tr p (1 ; A —O t 197 p t tr p l n t d n l pr n nd th l t r l n tr t n f th ( ; 7-1 197 p t tr p (1 ; — b 1979 p t tr p ( pr n t n t nt n n r th d r I M t h r p p I 1977 — n 197 W tt M n p n A tr l r f nd n l f l tt r tz p t tr p (3 ; b—Apr 197 M M p t tr p (5 O r M r 19 3 Sp ll r [ ]x t pl t (5 ; 7 Apr 19 3 Ev n (1 rl h n n f l d n p South I. NN. l n b 17 A 19 3 A K 5; M p 3 W l r n t n r nd (1 FD. L. Te An 19 b 19 5 A K W l r l tt r (1 n th 3 3 -3 5 d h n n ; h d pr n t nd t t ll br n; l tr nd nd rb d Material examined. p r pl 1 n n-t p x - r n ; f r h t h; t b br n t l t r t pl fr A tr l (A IC U C v r b d AK/ d nd pr n t th t bd b nt; p n - C ll t d thr h t th r t r b l t d th v rt x br dl btr n t n rd d pr r l fr l v l ddl h lf h rt p t r r t ; d th pr n A t d th t rd n nd t n (G K h l n n r nn r r n f fl n d l t r ll b r d l tt nd v r r r n th n p n b h r t nf ld th xt nd n fr nn r r n t fr nt l p l (AK th n p n f nd n pt p r p
—32— r t th b h r n nd n th r n S l x fr l t th l nth l t n p th f W ttl tr (K h l 199 3 1 ; M p M tl ll t d n p t tr p h rl t ll t n r rd r n 19 3 fr th A l nd r n th - 7 d nd nt nn br n p pl r l f ld f l tr ll t p l br n; x nd l ll Remarks. S v r l p n fr r t p r t r n- t t n ll th p x f f r nd t f t b f r t n M n St t r t ( th t rn S th br n W l nd St x v r St t r t (n rth t rn d nd pr n t n rl p n t t th p l h d S th W l pp r t b b r f th h th rt d ; t t l t r t; t b r t d h rt p t r r nd r b d p App r ntl ntr d d nt - t ; r n thl r nd d t br dl btr n t l nd fr A tr l h l t n n pp r n f th b d b ; h ll l n t d n l pr n d l t h t d t n nd nt nn l b d f n d l t r ll b r n ; n t n tr t n r l p n t t r n t l d th th p r f n p n t r l nth n n p rt p p l nth l t n p h r d n t d El tr l n p tb l tr n v r pr n; h r pr n nt; pl t d t n t; r p n t r rr n d d h rt p t r r t th br d d t n t n ; r ll t b nf d n r p x; t t l t r t v rt x br dl btr n t ; fr nt l p l t r d t n t; r d t 7 ° 1 l n ; t b r t d r t d nt nn l b xp d; h ll nfl t d d pr n bl l l t r ll 7 l n ; nd r t bd b nt b t n h nd nt nn l b l n d l t r ll b t b r t 7 l n f r d th n d ff lt t p- r n E l r r nd l n bv t b t n r t fr t W n f ll d v l p d f t M x ll r p lp th nd nt bl ntl nd 3rd M l th l t t rn t br fl btr n t t p x nt t l n l t v ntr ll b r n r 3 t t l l l r b t th t rn t p x r r nd d t n n l nd th nt br dl r f r r n t th n b r n rr p l r nd th Type data. Holotype: f l [W W n t ] d p l t r l n tr t n j t p t r r t ddl th G ll n ll 1 b r 19 1 d n C ll t n n l d p t r d r ll nd nt n d tr ht r d 1 51 b ( M n r b n rr l nd n v ntr ll n d p p t Paratypes ( f l ; A IC U M AC b v pr x ; t r t rn l l br fl nd l North I. W p K r r t 11/1 19 3 d f n d b v pr x M n r [ r n] [ p n ] (1 K Str tf rd r n M t ll dr t t p x h ll l r n t C ll (1 WN. W ll n t n W lt n h 1 19 5 El tr th nd r t ; p n t r b r l t r l n- d n C ll 15 (1 f d p ll ; n rr r v dj nt t t r n d t l South I. SD. l r r d S n 3 t -th rd M t rn th l t r l r n tr ht 19 1 W E rl p n f rn n b h / p d rp / pr l n d p t r rl t t -th rd l n th f x br dl f f r t (1 M p t rn dr t th fr n f h rt t n p t r r r n M t t rn l htl n t p t r r t Material examined. p r pl p rt n f 1 t p - x t n t rn l pr j t n b l rf t p l ( n bd n nd l n t l ( M t p r t x St rn t fr M l n t l 9th t rn t K W /S - h p d th p l r n t d t h d; ph ll b b- C ll t d n b r nt; p n xt nd n nt r rl b nd b f t n rd d fr n r l v l p n r ll t d b p n n fr Remarks. l nth l t t r h n n nd f rn n b h/p d rp/br dl f f r t C t n t n rr pr n t l r h rt h d th l t r l bl nt r nt nn l pr n d l t th nd Remarks. Or n ll l p n f l t was nt nn l b nd l n t p n h r n t - t d th th h l t p f l b t nf rt n t l ll t ll h r d th r h n n , b t l nth is th t ld b f nd h n SC f n h d th r v r r d l r n d b th n rr b l tr n v r l f th l n t l nd t d bd n h t x n th pr n t nd th nv x pr n t l d n n th d n n h v n th h rpl d f n d h n r fl t th r tr t d d tr b t n f th nt b l l f th pr n t th t f nd n n l nd n ; nd r l n th r nth d r p ; th r fl t d n th p f n
—33— S bf l MAC A II AE pr n t th pr l n d v rt x nd th t r v nd f v f th f r t v bl bd n l t rn t rth A r - G n M r tr n n p h v h ll l nv x pr n t l d th n l r t r n t l t r l r n th h d b br dl M r tr n 1 3 377 p p M r tr btr n t nd th frr t bd n l t rn t l n l l n r n b r n l n t p r f v M r rthr rté-Sén tèr 1 1 Unn r h nth d b t n n r p r n v - r pl nt n f r M r tr . t t n nd r t v d r n th d d b h rpl n tr t d t f r l nd r n ; fr nt l p l t r n t v bl ; fr n l v t d v r nt nn l b r n t h lf E l r b l n th t M r tr t r n p r n t n b t n f t Ant nn l n t th - 7; M p 5 nt - t l t t l n d nd f v n dth x l ( d nt f t n K h l 199 nd nt 9-11 n t bl d r M x ll r p lp th nd nd 3rd nt b r n tr n l l b th t n n th 3 - 3 d d r br n; nt nn r 3rd nt l n r th n d ; th nt l r nd f r ll b ll t br n n d t l h lf; t t l r f r t r t v r b d r th r x l n t ; d nv x th r nd d l t r l d nd pr n t th t bd b nt d r n l htl n tr t d n r b ; p x th d t n t b l n t bl ntl tr n l r t n rr l btr n t n rr ll r; b th th n nt b l tr n v r l b h nd ; p n t r h rpl d f n d nd d n t b - xt nd n nt r v ntr ll t l t r l r n t p t j t t n 3 fr ntr t ntr b n d r l f pr x th p t b r d b t r x tt r d nd ll r t rd b ; t r n t n b - nt pr n nt; v t p n xt rn ll l d t n l r f t 3 l n nt rn ll l n t r t rn l l r n t l n t th d p r p n t r th n n h d El tr l n pl t r tr n v r pr n; l t- 3 p rt r l r n tr ht l htl nv r n t p x; th n El tr th p n t r l r r n rl d t n t th l n t d n l l r n p t r r t h r nd xt nd- n pr n t d n nd b r l n r b r l v r n p t r rl t n r l tr l p x; p n t r ll t f d ; t t l t r t 1 n n l n ; t r l; nd r t b nt M t ll tr n t b r t t b t 3 - ° 9 l n n l d bl l M t rn nd p t rn f d th l n f f n l t r ll v r d n l d p t r rl r bl l d r ll nd t d t t b v l d f n d th r d b th l n fl n W n f ll d v l p d r l p n t t n l nd p M p t rn M l th th t rn t n rr l n t h d t p x n t xt nd n l t r ll t l tr nd p t r rl t t p t rn l t r ll G n t l p n th ddl p rt br dl r nd d b rd r d th p b n l t r ll M p r r d n rr l n t h d t ddl b r l v bl n rr r n p t r r t p t rn M t rn n rr l p r t n x t n p d t l t p l [AK] 3 n r 1975 t t rn bl nt p nt St rn t fr ; 1 t v bl M M b t n h ll l d l ( AC t rn th n rr h ll r v j t p t r r t r t p (33; CM C U AC U C rth h t x n l d b d n l n f t h I K h h 15 -1 [ 5 - ] 3 b nd r v th p r t f v pr j t n n rd 3 19 G v n ( K n r 1 v 1917 ( th 1 t t r r nl r d d n l t v ntt ll ttl M n G r 1 v 19 r (1 ; G n t l 9th t rn t Y- h p d th t h rt nd p l M n G r 1951 r (1 W p r l n xp nd d t t p; ph ll b nd p r r St t r t M t h r b 1975 W l r d t n t; p n tr p rt t t p x p n (1 ; r t S n t r 11-1 19 3 p- n n r f r t (1 ; 11-1 19 3 M n r r n r M r tr l r p l t n n h p (1 Wh n r d 7 1977 K h l (1 r l t n h p h v n v r b n th r hl v l t d M - AK S d t h l t p (1 nf ld r p n r v b r f th M r tr n r d r b d pr r l b K h l 1 v 197 (1 ; 197 d d x l M r thr h lt t d f h rt d r pt v p p r (1 ; 15 b 1977 n tr (1 Or t 15 b 195 v r n r h l nd p d ff r fr M M nd r b r f r tt n l (1 t r n th f rth A r h h n l d th t p p 191 ( ; 9 19 M W C rt r (1 ; 197 A l n r n n th r nd d l t r l r n f th M dd n n fl ht t d (1 W t r n n
-3 - 1951 Sp ll r (1 C Gr t rr r I K t 5 v r n t th p n t r l r r nd d p r th n n 1957 C W tt, pt p r [K nz ] r d f r t h d d n nd h rpl d f n d b t 3 p rt (1 ; n h 3 1957 W tt (3 p – El tr th p n t r th z th n pr n t C r l n d 5 M r 1977 tz p n (1 TK. b t l d t n t d n nd b r l t b r l v r r t Mt M n r (b l 15 19 3 M n r r n f d ; t t l t 1 l n ; t bd b nt t p (1 r S n 1 19 3 W E rl b t ° l n n l d bl l l t r ll v r (1 RI. Oh n M n r n [r l ] 3 Apr 19 d bl l d r ll l n fl n W n f ll d v l- rr (1 WI. ld n K t h n r r 1 Apr 193 p d R.M. Bull (1). M l th th t rn t pr l n d; r n r d ll South I. NN. C thr n r (G bb 3 v 19 5 nv r nt t d pl n t h d p x G n t l p n th G rl (1 ddl p rt n n rr l r nd d t p x
Material examined. p r pl 3 n n-t p x - Type data. M r tr x l : l t t p l nd n pl (AMΝΖ M AC p r l t t p l b ll d " [C ] r " fr th AK WO C K I WI / ΝΝ C ll t n ( ΜΝΗ th l ft nd r ht p n n C ll t d v b r– n d bl nt h r d n t d b SC rd d fr l v l t 5 M r tr v rt l : l t t p l l b ll d "[AK] rn ll t v n v t t n n r tr n b h A l nd " fr th Sh rp C ll t n ( M nd r t nfr ld AK ( M r tr x l , K h l 199 p r l t t p th d nt l l l t l b l h r d - 3 nd pp r ntl t d th d (C r x, n t d b SC S rp , C p r p ll n n p fl t M r tr fl v p : h l t p f l I b ll d "[ ] th d Al ll t d p n h ll l d l r " n th r n C ll t n ( M h p n nd K nz r d . t ll l b ll d "M r tr x l " b t th nl p n n th Βr n C ll t n th th rr t l l t d t nd f t th d r pt n p rf tl It h b n l b ll d Remarks. M. t r p r t d fr M. x l b th th h l t p b SC b r t l tr l t nd th r d t n t p n t t n f th h d nd l tr h p f p th t th M r n Material examined. p p n pl 39 n n-t p f r l nd x pl (CM C U M AC U C AK C K O WI W /— C ll t d O t b r– n r nd n n M r tr x l rd d fr n r l v l 5 ; M p C ll t d pr r l b p n v t t n n r p r n r t r ; r rd d fr l hn nd C r x p x l 1 77 1 7 (M r tr . r n 1 9 1911 19 d n 19 3 3 5 t K h l 199 3 Remarks. M. x l p r t d fr M. t r b th ( d nt f t n bd b nt t n th l tr th l d t n t p n t t n v rt l Sh rp 1 77 9 (M r tr . r n 1 9 n th h d nd l tr p rt l rl b t n th nd 1911 19 ( n n f x l . d n 19 3 th r br dl r nd d t btr n t h d b 3 5 d r pt n f x l p bl h d n br r 1 77; fl v p r n 1 93 11 (M r tr . 1911 19 Sh rp d r pt n f v rt l p bl h d n n n n 1 77 n th 3 -3 d d r br n; nt nn r nd f r ll b ll t br n n d t l h lf; t t l t r t v r b d E E E CES d nd pr n t th t bd b nt t n rl ppr d d l n t th b r l rl r nd d Abd ll h Μ. 19 9 h n t r l l f t n f th f l t n rr l btr n t ; p n t r v n fr n pr - Anth d th l l nd th l l b- r v l d n r t h rpl d f n d th h ll b t n rv t n t h nt l h t hr ft I6: th n p r r b t t ll d t n t t n r h d b b t 3 3-3 p rt n d b t n ; t r n t n Abd ll h Μ ; Abd ll h A 19 h t x n p t n b t n l r f t l n f r d th pr p d n tr b f th
–35– E r n n (C l Anth d Ent l t 1912: Descriptions of new genera and species of nthl z n 04: 73-7 C l pt r r n t n nd pr d n f th . . Arn tt r 19 3 d l d ( rd r 1 59 th In t t t ( (n 44: 379- f l nt-l fl r b tl l 3 p 7 3-7 1914: Descriptions of new genera and species of n: h b tl f th Un t d St t ( n l f r C l pt r p rt II ll t n f th Ν l nd In t d nt f t n W h n t n C C th l Un v r t t t I(2 : 79-1 r x + 111 pp 1923: Descriptions of new species of Coleoptera, Arn tt r; S l n G A ; h d G M 1993 p rt IlI ll t n f th l nd In t t t (8 : h n t nd p d r ll t n f th rld ( nd 7-7 dn l r & f n h ndb n . II. G n v ll r ll Ι 19 C l pt r Anth d n d It l , ( S ndh ll Cr n r v + 31 pp v l. II. l n Ed z n C ld r n v + pp l b rn Τ 1 91 rth r n t n A tr l n C l pt r 195 Anth f ll x r n n n f A th d r pt n f n n r nd p r n fl r l (C l Anth d Ent l t nthl t n f th l S t f S th A tr l I4: z n 86: 9 9 -3 5 tl r G r 19 In t pr d t r f b ll r l r K G 1913 On th t t p t n f th r r v r lt r n Ar z n 8: - 7 l pt r n d ( t r r th n t C rr l E ; E n r Τ 197 C nth r d n p t nt f d n n ll d n r r n t n f th Ent d t rr nt t n t S n 8 : 755-757 l l S t f nd n (8th r : 7- 9 C 1 95 C l pt r l l n t I.Ann l f 19 th d ; r 99 th d r hr d ; th Y r A d f S n 8: 35- 3 n S h n l n S ( d. , C l pt r r t l r -l n W n 1 pp Ch ndl r S 1975 A r v n f n rthr C nt 19 b r hr d ; r 99 th d r - th pr nt t n f t d-C n z p t n hr d ; n S h n l n S ( d. , C l pt r r (C l pt r Anth d r n t n f th A r t l rl n W n 1 pp n Ent l l S t 0 : 319-35 n d n 1953 Anth fr nç d r p d 1976: Use of cantharidin and meloid beetles to fl r l t d r p v n v fr nç ttr t Anth d (C l pt r n f nt l t 2: d nt l 20: 9-1 3 179-1 195 In t C l ptèr Anth d n d 1982: A revision of North American Notoxus with M d r, v l. 6. In t t t d h r h S nt - l d t n l f th W rld p f n n r v - b z z 153 pp (C l pt r Anth d Ent r ph : 333- 3 1971-9 Anth d d l f n d r n ( C l b n I ; Ch l zz ; n h tt 19 5 C l pt r p rt nt l t I I,2 : 13 -1 ; I 4, 0: Anth d f th n l tt r l th f th O br n 1 1-111; 6, 3 1 5-1 ; I 8 , : 1 1-11 ; v r nd d r pt n f pr n l f r f th 8 , 4 : 5- ll t n n l d l S été Anth ( ll 179 nd C l d n (M l nt l nné nn d n 8 , 8: 5- ; 0, 59 9- ; 1 n [ ] d 68: 93-511 0, : 3 3-3 C l b n I ; Ch l zz ; ll M ; r ll ; r tt n E 197 C l pt r (b tl p 95- 1 n M n A 1991 l tt r f n d l t b l h n t f A tr l t xtb f r t d nt nd nt t nt l n d r n (G d n d r r h r r CSI O/M lb rn Un v r t r p p l z n z n z n d ll n f l p ό x + 1 9 pp n r d 4: 7-1 9 r n Τ 1 M n l f th l nd C l pt r Cr b Τ K ; d l S ; W tt C 197 rd n W ll n t n C l n l M nd G l l S rv p n l l t n l nd n rb tr r - p rt nt x x + 51 pp t f r nd d d f n d l nd j rn l 1 1 M n l f th l nd C l pt r p rt f z l 3 9 + p II W ll n t n C l n l M nd G l l S r- Cr n A 1955 h n t r l l f t n f th v p rt nt p xx + 53-7 f l fC l pt r nd n th n l l d (S n 1886: Manual of the New Zealand Coleoptera, part 19 7 r pr nt pt n E W Cl 1 7 pp I W ll n t n G r d b r p 17-973 C thb rt r 19 7 In t ff t n t p t t 1893: Manual of the New Zealand Coleoptera, Part A r lt r h ndb n . 3 9 A r lt r r h V. W ll n t n S l C t ll p xv + 975-13 S rv U S A pp
-3 - v d n A ; W d E 19 9 r pt n nd b l - 1929: Anthicidae of the Philippines, II. h l pp n l n t n th l rv f Anth h r C j rn l f n 40: 53- 3 (C l pt r Anth d C l pt r t b ll t n 3 K h l G 199 tl n b rb n nv r n nt 5- l nd t d h d nt t nd t t f n 1979 h l rv nd r l t n h p f C n n t C l pt r n th n t r l nd d f d h b t t f C nt (C l pt r t r r C l pt r t b l nf ld A l nd (197 -19 9 SI l nt r t l t n : 33-39 t n r p rt n . 3 11 pp E n r Τ 19 In t n l f r r l h Elt rn rté-Sén tèr d 1 M r rthr vr n rh ndl n n d r d t h n z l h n G ll n 1 pp Ιn G ér n-Mén v ll M E ( d. , h ft 8 : 9-17 Sp t n r ph énér d n x rt lé r r ; G r n 1 C l pt r h l n rt r pré nt t n d nr v l r d r pt n t 1 Extr t d Ann l d l S été Ent l d ll d t t l pè d tt r nd d v n d r n r M lt t pp rè n n l r èr p rt In t C lé ptèr r n t h n t h r 191 Appl p d r ld nd t ntr l 1 M n r ph d Anth t nr v n n th r d r n f th f rth t U.S. C lé ptèr Ηétér èr d l tr b d r hél d p rt nt f A r lt r b ll t n n . 2. pp r S p xx + 3 pp r hh r 19 S n l nd d l v r t n n L nd hr 1977 I h r p lp n t d l (C l pt r : t v t f nt-l fl r-b tl (Anth d l- Anth d pr d t r f t t d th th l t d n S r t n 19 - 3 t n n t M nt r p n Ann l f th Ent l l S t f nd r rd : 1 3-17 A r 0: 1- 3 G z A 1777 Ent l h Β trä z d tt r r n 19 C l pt r p -553 n r r nné z ölft n A b d t r t v l 1 S ( d. , S n p nd l f t n f l v n r n- pz W d nn 73 pp v l Y r M Gr ll 1 3 pp Görn tz Κ 1937 C nth r d n l G ft nd Anl n tt l r n ; r tt n E 1991 C l pt r (b tl für In t n Arb t n über ph l h und ange p 5 3- 3 n h n t f A tr l t xtb f r Entomologie rl n hl 4: 11 -157 ndt t d nt nd r r h r r v l ( nd dn Ith b rd 193 Ent l l nv t t n n th ( Y C rn ll Un v r t r v + 1137 pp p d f nd l ( 3 Anth d (C l h r n ; t n A r 1995 ( n pr l Ind n f r t r rd ( nt l r 20(6 : 1-1 nd bf l f C l pt r ( th l t d n r nt n Η E 19 5 A n r ph f th b tl t d n t r f r n nd d t n f l - r p n In th t r d pr d t ; v l 1 nd n r t h M l S A p n ( p. , l ph l - ( t t v + 3 pp n nd l f t n f C l pt r p p r l - br t n th th b rthd f A Cr n tt n W 19 Ind x f n v l nd n- d n l v + 37 pp A M 1 95 r pt n f n p f A tr l- n C l pt r r d n f th nn n S t f d n G 19 3 An nd x f l nd b tl Ν S th W l (I8 4 (2nd r : 5 9- 3 r n t n f th l nd In t t t 4: 353- 399 1 95b r pt n f n p f A tr l n C l pt r r d n f th nn n S t f K n 1 9 r pt n f th Anth d f A - S th W l (2nd r I0: -319 tr l r n t n f th Ent l l S t f 1922: On Australian Anthicidae (Coleoptera). Ν S th W l 1- r d n f th nn n S t f S th K tn r D.H. 19 h l n t b t r p 1- W l 4 : 71-51 n r nn ( d. , S l n t v l 3 v B. 19 Afr n n bf l f Anth d Y r A d r (C l pt r fr th rn Afr Ent l K t C Y 19 t t f nth d l rv S nd n v I : 19- (C l pt r Anth d C l pt r t b ll t n 6: nn C 175 S t t r p r r n tr n t r 7 -95 nd l rd n n r p Kr h-Str ld Η 19 5 Anth d f th h l p- h r t r b d ff r nt n n l ; v l 1 p n I. h l pp n j rn l f n 2 : 515-535 (1 th dn l r nt S lv pp
-37- M W 1 7 t n ll t n f n t fr St v n C 1 C l pt r r r d n l G nd h r n t n f th Ent l l S t n nd d r pt r M r d l S été f S th W l -31 I pér l d t r l t d M I: 155-1 7 [n t M l nt Ε ; C 1 t r n t r ll d C lé ptèr n] r t v l r pr nt d 1 11 th t tl n r n h d r n C ll èr ; v l 17 r S v 1 pp nd d ff r nt p n t n r pt n d d x In t d l ér d n l pp 11 -13 [ n] n Ε 1 3 Ent l l n t Ent l l z n 5 37 - Uh nn G 197 G tt n n d r Anth d nd hr t t h An rdn n Ent l h lätt r 7 Orph n d G M ; G nz l z ; rtl tt 1971 1 -1 Id nt f t n nd v l t n f p n b ll r pr d - t r n th rn C l f rn rn l f n nt 197 G tt n n d r Anth d nd hr l 64: 1- t t h An rdn n l S t t h 1 7 r pt n f n n r nd p An rdn n Ent l h lätt r 7 75- f l nd C l pt r rt lII. Ann l nd 199 Anth d d r b r h n lb n l 22. z n f n t r l h t r (4th r I8: 57- 7 tr z r K nntn d r Anth d (C l pt r 1 77 r pt n f n n r nd p f Anth d M tt l n n d r Mün hn r nt l l nd C l pt r rt I Ann l nd h n G ll h ft 82: 7-1 z n f n t r l h t r (4th r : 1 -1 7 v n ll C 195 C nth r d n nd Ant d [ ] ll G v n 179 n S In t ; v l 1 S th Afr n j rn l f n I: 15 -155 Upp l Ed n 35 pp W nn Ε 1 9 Kr t h rz hn d r r - t r n M 1973 bl b h v r l th r -r l - ph l n nd t r t ph l n Arthr p d n M t An b t n n n rthr l n nd . n (C l pt r d r b n nd t hr b n n r Art n Anth d n f nt l t 4 : 7-73 rl n xY + 31 pp M 191 l d l d ; r S r pt d d l d W tt C 1979 Abbr v t n f r nt l l ll - p 11- 7 n S S h n l n ( d. , C l pt r r t n Ν l nd j rn l f z l 6: 519-5 C t l rl n W n W rn r G 19 A r v n f th r t p f 1911b Anth d ; r 3 In S S h n l n ( d. , S p nt (C l pt r : Anth d . Ann l f th Ent C l pt r r C t l rl n W n 1 pp l l S t f A r : 9 - 9 ll A 199 S t t p t n f l p lp n 19 A r v n f th rth A r n p f (C l pt r n br n d nd p t n f th Anth . tr (C l pt r Anth d M ll n r hr d C n d n nt l t 26: 515-53 p bl t n f th Ent l l S t fA r 195- ll A ; Y n K 1991 r pt n f th 4: t r l rv f d l fl b ll t (C l pt r Y n K 19 C nth r d n nd n t n h t r l r hr d d l n th ph l n t pl - r v Gr t nt l t I : 1 7-19 t n f th d v r C n d n j rn l f z l 6 : 19 b ld r rd nd b rv t n f n t 3 - 3 t d th nth r d n Gr t nt l t n h tt ; C l b n I ; Ch l zz 19 r h : 195-199 n th t f S l h h r nd th d n f S r 19 5 r pt n f th l rv f I schalia U nl Anth d (C l pt r . M n t r z l vancouverensis rr n t n (C l pt r Anth d t l n (n r , ppl nt I 0: 1 9-1 I h l n th b rv t n n th t t p - S hätz C ; ttn r K 199 C nth r d n- r t n b t n f th n Coleopterist's bulletin 39 1- l tr l n t h f l nth d p (C l Anth - 1991 d l d ( n br n d l nt-l d t hr ft für t rf r h n 4 : 9 - 99 fl r b tl p 5 -5 in W Stehr (ed.), I - Sh rp 1 77 r pt n f n p nd t r n t v l b (I K nd ll/ nt xv nd t n f n n r f C l pt r fr + 975 pp l nd Ent l t nthl z n 4: 0. 1991b Anth d ( n br n d Ant-l S th G 19 C l f rn tt n n t ll t n 660 f fl r b tl p 55 -55 n W St hr (ed.), I - th C l f rn A r lt r l Exp r nt St t n, t r n t v l b (I K nd ll/ nt xv r l , CA pp + 975 pp
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6 O ICES GA AA UI h r f r f r d n l p bl t n h K h t r t n r r n p p h b n t bl h d t n r th th xp rt h h h n t h n h t r n n l d t p bl h n t pr h n v h t n p n n r r h nt f l nt n th l nd f n . r n . h t h n t n r r n h r pr f n l n t n pt n nd p p , n r , t h h r t pr nt t n, t v r ff rt d t pr v d t h t h , h t r n t h r r f r d nt f t n nd nf r t n th t n r r , n r r , , h r t r bl t th n n p l t. t r n p n r t . n f . . d l th n n r n nv rt K n p p n f l nd br t nl , n th v rt br t r ll d r p n n r r h , n , n nt d, nd r n f r r v r d b th t t r n . E t n h t n n r M r n n f . . . r r h . K r t n p p n M r n n f l nd n t h t h p n n n t t . C ntr b t n r nv t d fr n p r n th th r t p l t ll nd r r . M t r l fr th . . Arthr p d C ll t n v l bl h t h n . K t t n n t h n t f r t d . t p p . E t n t h h n r r h r t C ntr b t r h ld d th r nt nt n th l nd Arthr p d C ll t n h t t . n ppr pr t b r f th n Adv r Gr p r th th S r Ed t r b f r n M h t h r t t n r ll n r d n ll b v n. h t h t h n t n , t Et t r n , t t t n t h t h .
S b r b r h ld ddr n r t n f . . , br r , Mt Alb rt r h C ntr , r v t h p p . M t h t n f 2 6 , A l nd, l nd. . . t br r , Mt Alb rt r h C ntr , r v t 2 6 , A l nd, l nd. S b r pt n t r : A t nd n rd r n nv ll b nt th h n , n E r n t h : A K h t ft r p bl t n p bl pr t n l t p p , p p , t fl r th rd r f r ll b nt fr t t t . t , r t n t t n t p p . t n p n n , t n , t n t l pr ( tl n pr nt , p 62 n l d . p n nd rf p t . S b r b r n l nd nd A tr l p th nd t d t (t r h t h r n 62 : K t p nt n $ GS n l d d n th pr . Oth r t n t p n n r t t t . M t b r b r p th l t d pr n $US, r v t n h r r n A h t r r l nt. n t r r n . K t h t t t h r t n n t r M r n . f ll n b r r v l bl , nd n b r b r h n t bt n f ll t r l Ε t r n p p ο . M h h h t n r t d nt. ll r nd n t t n p p , t n b r pt n nt r ff r d tr d d nt f h h t t . E r p h n t t h h ο t t 20 . n t h p p .
64 n f l nd K te A t n p A t r
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l nd Anth d S nn d pr v d d b St ph n n (www.bugz.org.nz). OC d t xt rr t d f r th r h bl b r v r Cr b FNZ r d t r 1 l 1 U r xtr t t xt fr th f r th r n b t t h t n t th r n l d nt f r t xt n r nd f r tt n