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Remarkable Aquatic Predators in the Genus Ocyptamus (Diptera, Syrphidae)
ROTHERAY et al.: 385-389 Studia dipterologica 7 (2000) Heft 2 • ISSN 0945-3954 Remarkable aquatic predators in the genus Ocyptamus (Diptera, Syrphidae) [Bemerkenswerte aquatische Pradatoren in der Gattung Ocyptamus (Diptera, Syrphidae)] by Graham E. ROTHERAY, Manuel ZUMBADO, E. Geoffrey HANCOCK and F. Christian THOMPSON Edinburgh (United Kingdom) Santo Domingo (Costa Rica) Glasgow (United Kingdom) Washington (USA) Abstract Third-stage larvae, puparia and adults are described for two species of Ocyptamus MACQUART and new synonyms are proposed. The larvae were found in water pockets within epiphytic Bromeliaceae in Costa Rica. They attacked a wide taxonomic range of insect larvae that char- acteristically co-occur in these phytotelmata, apparently subduing prey with venom and suck- ing out the internal contents. They possess a number of morphological and behavioural fea- tures not known in other predatory syrphids. These features include an enlarged and flattened anal end bearing a sucker, elongate posterior breathing tubes with vertically inclined spiracu- lar plates, and patches of needle-like spicules on the underside of the thorax. Although only two species were reared, larvae of 6 other species were discovered, which suggest that many more species occur in bromeliads. Key words Predator, phytotelmata, bromeliads, anal sucker, venom Zusammenfassung Von zwei Arten der Gattung Ocyptamus MACQUART werden die dritten Larvenstadien, die Puparien und die Imagines beschrieben und charakterisiert sowie neue Synonyme festgelegt. Die Larven wurden in Phytothelmata von epiphytischen Bromelien in Kostarika gefunden. Sie attackieren ein weites Spektrum von Insektenlarven, die hier gemeinsam mit den Syrphiden- larven leben. Offensichtlich lahmen sie die Beute mittels eines Giftes und saugen dann den deren Korperinhalt aus. -
Is the Mega-Diverse Genus Ocyptamus (Diptera, Syrphidae) Monophyletic
Molecular Phylogenetics and Evolution 62 (2012) 191–205 Contents lists available at SciVerse ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev Is the mega-diverse genus Ocyptamus (Diptera, Syrphidae) monophyletic? Evidence from molecular characters including the secondary structure of 28S rRNA ⇑ Ximo Mengual a,c, , Gunilla Ståhls b, Santos Rojo c a Dept. of Entomology, National Museum of Natural History, Smithsonian Institution, PO Box 37012, MRC-0169, Washington, DC 20013-7012, USA b Zoological Unit, Finnish Museum of Natural History, PO Box 17, FIN-00014 University of Helsinki, Finland c Instituto Universitario CIBIO – Dpto. de Ciencias Ambientales y Recursos Naturales, Universidad de Alicante, Apdo. 99, E-03080 Alicante, Spain article info abstract Article history: Phylogenetic relationships between two New World Syrphinae taxa (Diptera, Syrphidae), i.e. the highly Received 17 March 2011 diverse genus Ocyptamus and the large genus Toxomerus, were analysed based on molecular characters. Revised 17 August 2011 The monophyly of both taxa was tested and the taxonomic status of included subgenera and species Accepted 23 September 2011 groups was examined. Toxomerus constitutes the monogeneric tribe Toxomerini with more than 140 Available online 29 September 2011 described species, while Ocyptamus (tribe Syrphini) is a very diverse genus (over 300 spp.) with multiple recognised subgenera and species groups. Sequence data from three gene regions were used: the mito- Keywords: chondrial protein-coding gene cytochrome c oxidase subunit I (COI) and the nuclear 28S and 18S ribo- Toxomerus somal RNA genes. The secondary structure of two expansion segments (D2, D3) of the ribosomal 28S Ocyptamus Monophyly RNA gene is presented for the family Syrphidae and used for the first time in a multiple sequence align- Syrphidae ment. -
Diptera: Syrphidae) with Notes on the Placement of the Subfamily by F
A NEW GENUS OF MICRODONTINE FLIES (DIPTERA: SYRPHIDAE) WITH NOTES ON THE PLACEMENT OF THE SUBFAMILY BY F. CHRISTIAN THOMPSON I discovered the following new genus of Syrphidae while reviewing the Neotropical Microdontinae. I had planned to put aside its de- scription until my study of the subfamily was finished. However, publication of a second species, assignable to it by van Doesburg (1966) has necessitated publishing a name now for this genus so that it can be included in the Catalog of South American Diptera. Paragodon, new genus Very small (4-5 mm.) microdontine flies. Face simple (slightly produced in paragoides); cheeks absent, eyes bordering on the oral opening; eyes dichoptic in both sexes; occiput evenly developed. An- tennae short, about one-half as long as face; aristae short and thick- ened. Thorax about as long as broad; pleura bare except mesopleura pilose and meta.pleura with microscopic, pile; scutellum without apical spines and fringe; metasterna undeveloped and bare; metathoracic spiracles without hair fringes. Legs simple, with no basal setal patches on the femora and with cicatrices only on the hind femora. Vings without the spurious vein, with all apical crossveins straight. Abdomen oval, lateral margins slightly emarginate and rolled under ventrally, with ISt sternite bare and st spiracles without hair fringes. Genitalia simple; ejaculatory apodeme simple, apical por- tion not triangularly flared; ejaculato.ry sac not sclerotized; ejacula- tory process single, short, not posteriorly fused to ejaculatory hood; ejaculatory hood with anterior ventral portion elongate; sustentacu- lar apodeme present, double, fused anteriorly to base of ejaculatory Contribution no. 1392 from the Systematics and Morphology Research Laboratory, Department of Entomology, University of Massachusetts. -
The Ecology of Mitcham Common 1984 Report
THE ECOLOGY OF MITCHAM COMMON THE(A ECOLOGY report on the statusOF MITCHAM of the flora and COMMON fauna) The final report of the "Ecological Survey of Mitcham Common" Supervised by: R.K.A. Morris BSc. FRES Participating authors: R.D. Dunn BSc. A.M. Harvey BSc. J.A. Hollier BSc. ARCS. FRES. C.M. Johnstone Cert. Ecol. Cons. A.D. Sclater BSc. FRES. C. Wilson BSc. Funded by: The Manpower Services Commission Administered by: Merton Community Programme Agency Sponsored by: The Mitcham Common Conservators and the London Borough of Merton Department of Recreation and Arts Report completed and submitted: September 1984. Crown Copyright. Cover photograph: Seven Islands Pond from Mill Hill, September 1974 (Photo Dr P.G. Morris) iv 2016 version This report was produced by a team of recent graduates, employed under the 'Community Programme' and funded by the Manpower Services Commission. The objectives of the Programme were to provide the long-term unemployed with opportunities to train or re- train, so that they might get more permanent work. This Programme funded a considerable number of environmental jobs, and provided the stepping stone for many ecologists to move into mainstream jobs. I have lost contact with most of the team members of this project, but am aware that at least one (apart from me) went onto a successful career in an ecological discipline. Looking back to the year of 1983-84, it is difficult to appreciate the achievement of the team. We commenced work in September 1983 and were due to report in late August 1984. The timing was unfortunate because we were unable to make best use of the year, with the winter occupying most of the project. -
Diptera: Syrphidae)
MEMOIRS of THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Number 9 THE FLOWER FLIES OF THE WEST INDIES (DIPTERA: SYRPHIDAE) by F. CHRISTIAN THOMPSON Agricultural Research Service Agricultural Research, Sci. and Educ. Admin. U.S. Department of Agriculture, Washington, D.C. Published by THE ENTOMOLOGICAL SOCIETY OF WASHINGTON Washington, D.C. 1981 PUBLICATIONS COMMITTEE of THE ENTOMOLOGICAL SOCIETY OF WASHINGTON 1981 E. Eric Grissell John M. Kingsolver Wayne N. Mathis George C. Steyskal Thomas E. Wallenmaier David R. Smith, Editor Printed by Allen Press, Inc. Lawrence, Kansas 66044 Date issued: 2 September 1981 TABLE OF CONTENTS Abstract ...................................................................... 4 Acknowledgments .......................... " ................ ,................. 5 Introduction .................. , ........................... ,.................... 7 Economic Importance ........ , ........................................ ,........ 7 Distribution .................................................... ,.............. 9 Taxonomy .............................................................. ,..... 13 Key to Genera of West Indian Syrphidae ......................................... 17 Syrphus Fabricius .............................................................. 20 Allograpta Osten Sacken .............................................. ,........ 23 Pseudodoros Becker .................................. , . 33 Ocyptamus Macquart ........................................................... 34 Salpingogaster Schiner ..................................... -
Chapter 7 – Associations Between Microdontinae and Ants
Cover Page The handle http://hdl.handle.net/1887/18582 holds various files of this Leiden University dissertation. Author: Reemer, Menno Title: Unravelling a hotchpotch : phylogeny and classification of the Microdontinae (Diptera: Syrphidae) Issue Date: 2012-03-13 7 Review and phylogenetic evaluation of associations between Microdontinae (Diptera: Syrphidae) and ants (Hymeno- ptera: Formicidae) Abstract. The immature stages of hoverflies of the subfamily Microdontinae (Diptera: Syrphidae) are known to develop in ants nests, as predators of the ant brood. The present paper reviews published and unpublished records of associations of Microdontinae with ants, in order to discuss the following questions: 1. are alle Microdontinae associated with ants?; 2. are Microdontinae associated with all ants?; 3. are particular clades of Microdontinae associated with particular clades of ants? A total number of 103 records of associations between the groups are evaluated, relating to 42 species of Microdontinae belonging to 14 (sub)genera, and to 58 species of ants belonging to 23 genera and four subfamilies. Known associations are mapped onto the most recent phylogenetic hypotheses of both ants and Microdontinae. The taxa of Microdontinae found in association with ants appear to occur scattered throughout their phylogenetic tree, and one of the supposedly most basal taxa (Mixogaster) is known to be associated with ants. This suggests that associations with ants evolved early in the history of the subfamily, and have remained a predominant feature of their lifestyle. When considering the phylogeny of ants, associations with Microdontinae are only known from the subfamilies Dolichoderinae, Formicinae, Myrmicinae and Pseudomyrmecinae, which are all part of the the so-called ‘formicoid’ clade. -
HOVERFLY NEWSLETTER Dipterists
HOVERFLY NUMBER 41 NEWSLETTER SPRING 2006 Dipterists Forum ISSN 1358-5029 As a new season begins, no doubt we are all hoping for a more productive recording year than we have had in the last three or so. Despite the frustration of recent seasons it is clear that national and international study of hoverflies is in good health, as witnessed by the success of the Leiden symposium and the Recording Scheme’s report (though the conundrum of the decline in UK records of difficult species is mystifying). New readers may wonder why the list of literature references from page 15 onwards covers publications for the year 2000 only. The reason for this is that for several issues nobody was available to compile these lists. Roger Morris kindly agreed to take on this task and to catch up for the missing years. Each newsletter for the present will include a list covering one complete year of the backlog, and since there are two newsletters per year the backlog will gradually be eliminated. Once again I thank all contributors and I welcome articles for future newsletters; these may be sent as email attachments, typed hard copy, manuscript or even dictated by phone, if you wish. Please do not forget the “Interesting Recent Records” feature, which is rather sparse in this issue. Copy for Hoverfly Newsletter No. 42 (which is expected to be issued with the Autumn 2006 Dipterists Forum Bulletin) should be sent to me: David Iliff, Green Willows, Station Road, Woodmancote, Cheltenham, Glos, GL52 9HN, (telephone 01242 674398), email: [email protected], to reach me by 20 June 2006. -
Diptera: Syrphidae)
A revision of Nearctic Dasysyrphus (Diptera: Syrphidae) Michelle Mary Locke A thesis submitted to the Faculty of Graduate Studies and Research in partial fulfillment of the requirements for the degree of Master of Science in Biology Carleton University Ottawa, Ontario ©2012 Michelle Mary Locke Library and Archives Bibliotheque et Canada Archives Canada Published Heritage Direction du Branch Patrimoine de I'edition 395 Wellington Street 395, rue Wellington Ottawa ON K1A0N4 Ottawa ON K1A 0N4 Canada Canada Your file Votre reference ISBN: 978-0-494-91543-1 Our file Notre reference ISBN: 978-0-494-91543-1 NOTICE: AVIS: The author has granted a non L'auteur a accorde une licence non exclusive exclusive license allowing Library and permettant a la Bibliotheque et Archives Archives Canada to reproduce, Canada de reproduire, publier, archiver, publish, archive, preserve, conserve, sauvegarder, conserver, transmettre au public communicate to the public by par telecommunication ou par I'lnternet, preter, telecommunication or on the Internet, distribuer et vendre des theses partout dans le loan, distrbute and sell theses monde, a des fins commerciales ou autres, sur worldwide, for commercial or non support microforme, papier, electronique et/ou commercial purposes, in microform, autres formats. paper, electronic and/or any other formats. The author retains copyright L'auteur conserve la propriete du droit d'auteur ownership and moral rights in this et des droits moraux qui protege cette these. Ni thesis. Neither the thesis nor la these ni des extraits substantiels de celle-ci substantial extracts from it may be ne doivent etre imprimes ou autrement printed or otherwise reproduced reproduits sans son autorisation. -
Baccha (Ocyptamus) Medina, B
The Syrphidae of Puerto Rico1'2 H. S. Telford3-* One cannot state with certainty when the first syrphid was collected from Puerto Rico and adjacent islands. Fabricius described a number of 1 Manuscript submitted to Editorial Board October 30, 1972. 2 Scientific paper number 3914. College of Agriculture Research Center, Washing ton State University, Pullman, Washington. Work was conducted under Project No. 0046. 3 Professor and Entomologist, Department of Entomology, Washington State University; Visiting Scientist, Department of Entomology, Agricultural Experiment Station, Mayagiiez Campus, Uío Piedras, Puerto Rico, September 1968-March 1969. This study was made possible by financial support from the Department of En tomology, Agricultural Experiment Station, Mayagiiez Campus, University of Puerto Rico, Río Piedras. I wish to thank Dr. L. P. R. F. Martorell, formerly Chairman, Department of Entomology, Agricultural Experiment Station, especially for his support and aid in all aspects of the project. Mr. Silverio Medina Gaud, Associate Entomologist, Agricultural Experiment Station, was of considerable help. He ac companied me on almost all field trips, assisted in sorting and preparing the material and made valuable field trips on his own. Dr. J. R. Vockeroth, Entomology Research Institute, Ottawa, Canada, verified many determinations and offered advice on nomenclatural problems. Others who materially aided in the loan of specimens, verified determinations or in other ways were: Dr. George Drury, U.S. Atomic Energy Commission, El Verde-Caribbean National Forest, Puerto Rico; Dr. Y. S. Sedman, Western Illinois University; Dr. L. V. Knutson, Systematic Entomology Laboratory, Agricultural Research Service, United States Department of Agriculture; Dr. P. W. Wygodzinsky, American Museum of Natural History; Dr. -
Adult Behaviour in Two Species of Cerioicine Flies, Primoerioides Petri (Hervé-Bazin) and Ceriana Japonica (Shiraki) (Diptera S
1 Adult behaviour in two species of cerioidine flies, Primocerioides petri (Hervé-Bazin) and Ceriana japonica (Shiraki) (Diptera Syrphidae) Toshihide ICHIKAWA & Kenji ŌHARA [Abstract, captions of figs all in English] Introduction Most adult Diptera belonging to the Syrphidae are known to be diurnal pollinators which fly to the flowers of various seed plants and feed on nectar and pollen (1, 2, 3) Many different species have black-and-yellow or black-and-orange stripes and are thought to be simulating female adult Apoidea, which are the most important pollinators and have poisonous stings and similar body stripes (4, 5, 6). A tribe of the Syrphidae, Cerioidini, which we study in this paper, has body stripes similar to none of the Apoidea, but instead simulate Eumenidae or Polistinae, both belonging to the Vespoidea, which are generally called hunter wasps. The Cerioidini includes five genera (Ceriana Rafinesque 1815, Monoceromyia Shannon 1925, Polybiomyia Shannon 1925, Sphiximorpha Rondani 1850 and Primocerioides Shannon 1927) and 197 recorded species worldwide, but for only 19 of which are the life-style and development times known (7). In Japan three species of the Cerioidini are known: recent records of the capture of adult insects show that they occur in Honshū, Shikoku and Kyūshū. Observations of adult Monoceromyia pleuralis (Coquillett) are relatively frequent so it became clear that they visit Quercus acutissima and Ulmus, which exude tree-sap from May to September (8, 9, 10, 11). The lifestyle and life history of the other two species, Primocerioides petri (Hervè-Bazin) and Ceriana japonica (Shiraki), are not known, and so accidental encounters remain the only possibility. -
Review and Phylogenetic Evaluation of Associations Between Microdontinae (Diptera: Syrphidae) and Ants (Hymenoptera: Formicidae)
Hindawi Publishing Corporation Psyche Volume 2013, Article ID 538316, 9 pages http://dx.doi.org/10.1155/2013/538316 Review Article Review and Phylogenetic Evaluation of Associations between Microdontinae (Diptera: Syrphidae) and Ants (Hymenoptera: Formicidae) Menno Reemer Naturalis Biodiversity Center, European Invertebrate Survey, P.O. Box 9517, 2300 RA Leiden, The Netherlands Correspondence should be addressed to Menno Reemer; [email protected] Received 11 February 2013; Accepted 21 March 2013 Academic Editor: Jean-Paul Lachaud Copyright © 2013 Menno Reemer. This is an open access article distributed under the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited. The immature stages of hoverflies of the subfamily Microdontinae (Diptera: Syrphidae) develop in ant nests, as predators ofthe ant brood. The present paper reviews published and unpublished records of associations of Microdontinae with ants, in order to discuss the following questions. (1) Are all Microdontinae associated with ants? (2) Are Microdontinae associated with all ants? (3) Are particular clades of Microdontinae associated with particular clades of ants? (4) Are Microdontinae associated with other insects? A total number of 109 associations between the groups are evaluated, relating to 43 species of Microdontinae belonging to 14 genera, and to at least 69 species of ants belonging to 24 genera and five subfamilies. The taxa of Microdontinae found in association with ants occur scattered throughout their phylogenetic tree. One of the supposedly most basal taxa (Mixogaster)isassociatedwith ants, suggesting that associations with ants evolved early in the history of the subfamily and have remained a predominant feature of their lifestyle. -
Man Ual Ofnearctic Diptera Volume 2
Man ual ofNearctic Diptera volume 2 Coordinated by J. F. McAlpine (Editor) B. V. Peterson G. E. Shewell H. J. Teskey J. R. Vockeroth D„ M. Wood Biosystematics Research Centre (formerly Institute) Ottawa, Ontario Research Branch Agriculture Canada Monograph No. 28 1987 M M \ SYRPHIDAE 52 J. R. VOCKEROTH AND F. C. THOMPSON Fig. 52.1. Adult male of Syrphus torvus Osten Sacken. Small to large, very slender to robust flies (Figs. 1-3), Mouthparts variable in length, usually correlated with length 4-25 mm long. Body usually black, very often with yellow of subcranial cavity; taxonomic significance of variation or orange markings on head and thorax and particularly on unknown. Antenna sometimes borne on a short or long fron- abdomen, more rarely predominantly brown, yellow, tal prominence (Figs. 12, 13); scape and pedicel subcylin- metallic green, or blue, or with various combinations of these drical but varying greatly in length, with hairs or setae; first or other colors. Integument usually smooth but sometimes flagellomere varying greatly in shape, and often with a partly or totally punctate, sculptured, or rugose, usually near- distinct sensory pit on lower part of inner surface; arista ly covered with dense short hairs, rarely with long hairs or usually with two aristomeres but sometimes with three, usual- nearly bare; some hairs sometimes flattened or scale-like and ly dorsal but sometimes subapical or apical, usually longer forming dense tomentum, or on thorax strong and bristle- than first flagellomere but very short in some Microdontinae like; both haired and bare portions shining, slightly to densely and in some groups with apical arista, usually bare or with pruinose, or with very short dense pile.