VOL. 104, NO. 3 S VOL. 104 JUNE 2020 Special Issue NO. 3 Instructions to Contributors

1. Send manuscript to: Wanda I. Lugo Technical Editor Agricultural Experiment Station

Jardín Botánico Sur pecia l 1193 Calle Guayacán San Juan, PR 00926-1118 I

2. Submit only manuscripts not previously published or submitted for publication ss ue elsewhere. Manuscripts must convey only unpublished data. TH E JO U RN A L OF 3. Type, double spacing on 8 1/2 × 11 inch paper. Set 1-inch margins. 4. Include an original and three copies of the text and all exhibits. 5. Include an abstract no longer than 200 words. After the abstract, list no more than five key words. Also write a “resumen” in Spanish no longer than 250 words. Pro- vide a Spanish version of the title for the “resumen.” 6. Number illustrations in consecutive Arabic numbers. Set captions above the tables. Set captions for figures (photographs, drawings, diagrams) and footnotes on a sepa- rate sheet. Photographs must be large glossy prints with good contrast. Drawings must be in india ink. Identify figures on the back with author’s name and illustra- Biodiversity of Heteroptera in Puerto Rico:

tion number. A GR ICU LT U R E Part II. Annotated Checklist and Keys of 7. List references alphabetically. Use the Harvard system (last name, year) to refer to them in the text. When there are three or more authors, mention only the first Lygaeoidea () in the text, substituting et al. for the other names. Include all authors in the refer- ences. Include titles of references. Date must follow the author’s name. A.E. Segarra-Carmona, R.A. Franqui and H. Pérez-Martínez 8. The Agricultural Experiment Station will charge all authors who are not staff members for all the costs incurred by the publication of their articles at the current Abstract...... 1 of

costs as billed to us by the printer. Paying contributors will receive a free copy of the number in which their article is included. th e Resumen...... 2 U N I V E RS TY OF PUE RTO R IC O J 9. The Journal does not issue reprints of articles published. Authors who want re- Introduction...... 2 prints must tell us beforehand and state number of reprints and address to which Key to Superfamilies of Pentatomomorpha ...... 3 they will be mailed. We will forward their orders to the printer, who will mail the reprints and bill them directly to the stated address. Superfamily Lygaeoidea...... 4 —–——— Biology and Economic Importance...... 5 State of Knowledge and Objectives ...... 5 Annual subscription: $25.00 in U.S. and Canada; other countries add $1.00 per issue for postage. Single issue, current or back (if available): $5.00 in U.S. and Taxonomical Accounts...... 6 Canada; other countries add $1.00 for postage. Key to Lygaeoidea Families in Puerto Rico...... 7 Family BERYTIDAE (Fieber)...... 8

POSTMASTER: Family LYGAEIDAE (Schilling)...... 11 Send address changes to: Publication Section Subfamily Ischnorhynchinae Stål ...... 14 Agricultural Experiment Station Subfamily Lygaeinae Stål...... 15 Jardín Botánico Sur 1193 Calle Guayacán Family CYMIDAE Baerensprung...... 25 San Juan, PR 00926-1118 Family NINIDAE Barber...... 26

(Continued Inside Back Cover) Statement of Circulation Policy u n e The Agricultural Experiment Station of the University of Puerto Rico does 2020 not discriminate because of sex, race, national origin or religious and political UNIVERSITY OF PUERTO RICO affiliation in the distribution of its publications. Mayagüez Campus College of Agricultural Sciences AGRICULTURAL EXPERIMENT STATION San Juan, P.R. UNIVERSITY OF PUERTO RICO Mayagüez Campus Family BLISSIDAE Stål...... 27 AGRICULTURAL EXPERIMENT STATION Subfamily Blissinae Stål...... 28 San Juan, Puerto Rico Family GEOCORIDAE Baerensprung ...... 30 Jorge Haddock, Ph.D., President Subfamily Pamphantinae Barber & Bruner...... 31 Agustín Rullán-Toro, Ph.D., Chancellor Subfamily Geocorinae Stål...... 31 OFFICE OF THE DIRECTOR Family OXYCARENIDAE Stål ...... 34

Raúl Macchiavelli, Ph.D., Professor...... Dean and Director Family PACHYGRONTHIDAE Stål...... 35 L. N. Avilés, Professor ...... Acting Associate Dean and Deputy Director Subfamily Pachygronthinae Stål...... 36 V. Carro-Figueroa, M.A., Sociologist...... Assistant Dean for Research Y. Montes-Arroyo, J.D...... Legal Assistant to the Dean and Director Family Amyot & Serville . . . . . 37 J. Hernández-Miró, M.B.A...... Human Resources Officer Subfamily Amyot & Serville ...... 40 EDITORIAL BOARD Tribe Antillocorini Ashlock ...... 40 Tribe Cleradini Stål...... 42 L.N. Avilés, M.S., Professor Ricardo Goenaga, Ph.D., Plant Physiologist, USDA-ARS, V. Carro-Figueroa, M.A., Sociologist Tropical Agriculture Research Station Tribe Lethaeini Stål...... 42 Wanda I. Lugo, M.S., Professor and Technical Editor María de L. Lugo-Torres, Ph.D., Weed Scientist Arístides Armstrong, M.S., Entomologist Lydia I. Rivera-Vargas, Ph.D., Phytopathologist Tribe Myodochini Stål...... 45 Tribe Rhyparochromini Stål...... 58 SCIENTIFIC STAFF1 Tribe Ozophorini Sweet...... 59 Tribe Udeocorini Sweet ...... 63 Agricultural Economics and Rural Sociology A. Segarra-Carmona, Ph.D., Entomologist V. Snyder-Sevits, Ph.D., Soil Physicist V. Carro-Figueroa, M.A., Sociologist Discussion...... 63 R. Tirado-Corbalá, Ph.D., Associate Professor M. Cortés, M.A., Economist E. Valencia-Chin, Ph.D., Professor J.C. Hernández-Correa, Ph.D., Associate Professor Checklist of Lygaeoidea in Puerto Rico...... 66 R. Vargas, Ph.D., Nematologist, Acting Director of A. Jiménez-Maldonado, Ph.D., Director of Department Department H.S. Tavárez, Ph.D., Associate Professor Acknowledgements...... 68 D. Viteri, Ph.D., Associate Professor Literature Cited ...... 69 Agricultural Engineering and Biosystems Science J.A. Huertas-Miranda, Ph.D., Associate Professor Plates I-VII...... 74 J.A. Arroyo-Aguilú, Ph.D., Animal Nutritionist, Professor F. Monroig, Ph.D., Professor and Acting Director of Emeritus Department A. Casas-Guérnica, M.S., Animal Nutritionist K. Domenech, Ph.D., Assistant Researcher Agroenvironmental Sciences E. Jiménez-Cabán, Ph.D., Professor A. Armstrong, M.S., Entomologist J.R. Latorre-Acevedo, Ph.D., Animal Physiologist and I. Cabrera-Asencio, M.S., Entomologist Director of Department R.N. Chávez-Jauregui, Ph.D., Professor V.M. Negrón-Pérez, Ph.D., Assistant Researcher J.A. Chong-Nuñez, Ph.D., Associate Horticulturist M. Pagán-Morales, Ph.D., Professor M.E. Cintrón-Muñoz, M.S., Assistant Chemist T. Ruiz-López, Ph.D., Animal Husbandman E.C. Estévez-Salazar, Ph.D., Phytopathologist C.A. Flores-Ortega, M.S., Associate Horticulturist2 Central Analytical Laboratory R.A. Franqui-Rivera, Ph.D., Entomologist A. Ramos-Pérez, M.S., Assistant for Research in Charge F. Gallardo-Covas, Ph.D., Entomologist and Administrator of Adjuntas Substation Agroenvironmental Chemistry Laboratory M.C. Giraldo-Zapata, Ph.D., Associate Professor Á.M. Linares-Ramírez, Ph.D., Associate Professor J.A. Dumas-Rodríguez, Ph.D., Chemist M. de L. Lugo, Ph.D., Weed Scientist G.A. Martínez, Ph.D., Soil Chemist Publications Section S. Medina-Gaud, Ph.D., Entomologist, Professor Emeritus W.I. Lugo, M.S., Professor and Technical Editor in Charge M.A. Muñoz-Muñoz, Ph.D., Soil Chemist L. Albanese, B.A., English Consultant J.M. O’Hallorans-Castillo, Ph.D., Associate Agronomist M. de C. Plaza-Delestre, Ph.D., Horticulturist Library C. Rivera-Goyco, Ph.D., Research Assistant L.E. Méndez-Márquez, M.I.S., Librarian I in Charge and W. Robles-Vázquez, Ph.D., Professor Administrator of Río Piedras Research Center J.C.V. Rodrigues, Ph.D., Virologist E. Román-Paoli, Ph.D., Agronomist ______1June 30, 2020 J. Román-Toro, Ph.D., Nematologist, Professor Emeritus 2On leave C. Ruiz-Díaz, M.S., Research Assistant