The University of Chicago 8VHRI$OSKD%HWDDQG*DPPD'LYHUVLW\0HDVXUHVWR&KDUDFWHUL]H6HHG'LVSHUVDOE\$QLPDOV $XWKRU V 'RXJODV*6FRILHOG3HWHU(6PRXVH-RUGDQ.DUXELDQDQG9LFWRULD/6RUN 6RXUFH7KH$PHULFDQ1DWXUDOLVW9RO1R 'HFHPEHU SS 3XEOLVKHGE\The University of Chicago PressIRUThe American Society of Naturalists 6WDEOH85/http://www.jstor.org/stable/10.1086/668202 . $FFHVVHG Your use of the JSTOR archive indicates your acceptance of the Terms & Conditions of Use, available at . http://www.jstor.org/page/info/about/policies/terms.jsp . JSTOR is a not-for-profit service that helps scholars, researchers, and students discover, use, and build upon a wide range of content in a trusted digital archive. We use information technology and tools to increase productivity and facilitate new forms of scholarship. For more information about JSTOR, please contact
[email protected]. The University of Chicago Press, The American Society of Naturalists, The University of Chicago are collaborating with JSTOR to digitize, preserve and extend access to The American Naturalist. http://www.jstor.org This content downloaded from 128.97.244.148 on Mon, 5 Aug 2013 10:15:52 AM All use subject to JSTOR Terms and Conditions vol. 180, no. 6 the american naturalist december 2012 Use of Alpha, Beta, and Gamma Diversity Measures to Characterize Seed Dispersal by Animals Douglas G. Scofield,1,2 Peter E. Smouse,3 Jordan Karubian,4 and Victoria L. Sork1,5,* 1. Department of Ecology and Evolutionary Biology, University of California, Los Angeles, California 90095; 2. Umea˚ Plant Science Centre, Department of Plant Physiology, Umea˚ University, 901 87 Umea˚, Sweden; 3. Department of Ecology, Evolution, and Natural Resources, School of Environmental and Biological Sciences, Rutgers University, New Brunswick, New Jersey 08901; 4.