University of Birmingham a Re-Evaluation of Goniopholidid

Total Page:16

File Type:pdf, Size:1020Kb

University of Birmingham a Re-Evaluation of Goniopholidid University of Birmingham A re-evaluation of goniopholidid crocodylomorph material from Central Asia Halliday, Thomas J.D.; Brandalise De Andrade, Marco; Benton, Michael J.; Efimov, Mikhail B. DOI: 10.4202/app.2013.0018 License: Creative Commons: Attribution (CC BY) Document Version Publisher's PDF, also known as Version of record Citation for published version (Harvard): Halliday, TJD, Brandalise De Andrade, M, Benton, MJ & Efimov, MB 2015, 'A re-evaluation of goniopholidid crocodylomorph material from Central Asia: Biogeographic and phylogenetic implications', Acta Palaeontologica Polonica, vol. 60, no. 2, pp. 291-312. https://doi.org/10.4202/app.2013.0018 Link to publication on Research at Birmingham portal Publisher Rights Statement: Checked for eligibility: 19/06/2018 General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. Where a licence is displayed above, please note the terms and conditions of the licence govern your use of this document. When citing, please reference the published version. Take down policy While the University of Birmingham exercises care and attention in making items available there are rare occasions when an item has been uploaded in error or has been deemed to be commercially or otherwise sensitive. If you believe that this is the case for this document, please contact [email protected] providing details and we will remove access to the work immediately and investigate. Download date: 23. Sep. 2021 http://app.pan.pl/SOM/app60-Halliday_etal_SOM.pdf SUPPLEMENTARY ONLINE MATERIAL FOR A re-evaluation of goniopholidid crocodylomorph material from Central Asia: Biogeographic and phylogenetic implications by Thomas J.D. Halliday, Marco Brandalise de Andrade, Michael J. Benton, and Mikhail B. Efimov Published in Acta Palaeontologica Polonica 2015 60 (2): 291-312. http://dx.doi.org/10.4202/app.2013.0018 Character coding for redescribed material, and changes to E. delfsi Changes in coding of Eutretauranosuchus delfsi Character coding for redescribed material, and changes to E. delfsi All characters and character states are as found in Andrade et al. (2011). The following are the states assigned to Kansajsuchus extensus, “Sunosuchus” cf thailandicus (formerly “Sunosuchus” shartegensis), PIN 2229 (formerly Turanosuchus aralensis) and some changes to Eutretauranosuchus delfsi. Kansajsuchus extensus: 1[1/2]?1[1/2]???00110?????10000????00?200??1??10?0???????????1 ?12011121?3??????110010??????1??????1?????????01001??00- 300010010????0???????11012011?0??100?10010?0??[1/2]0?????????? ??????????????????????00?11?????11???????1??????2????????????? ??????????????????????0?0?100?0?????00?000111?000??1??00001220 0000?000010?20??????????????1????????????????????????????????? 0201000100?????0000021011?????000??0[1/2]?1??0???????????????0 ??00??000??????1011????????????????01?????1????????0???2?????0 1011???????? PIN 4174-1 1[1/5]??[1/2]0[0/1]10010???111?0000000?0??2???????????1??1??0? 1??10120????10???????110010?111101?100111?0?????0000?????????? ??00????????00?????110010?01?1???????????????????????????????? ??????????????????????????????0[1/2]0??11??0??????0??????????? ???????????????????????????????????????0?????????????000?12?0? 000?001?????????02[1/2]0211100?1100100001100??0???11100?000??? 00???1????????0110000000011?0?00[0/1]0000011?2?02[0/1]000????? ??00?00???00?????????????????????????????????????????????????? ?????????????????? PIN 2229 ?????[0/1]??0010????1110?0???????????????????????????????10??? ??????????????[1/2]0?0??011?????????1????????????????????????? ?????????????????????????????????????????????????????????????? ?????????????????????????????????????????????????????????????? ????????????????????????????????????????????????????0??????00? ???????????????????11?0????00110021????110??????????02?101???? ???????????????????????????????0[1/2]0000[0/1]01200?????0????? ?????????????????????????????????????????????????[1/2]??????1? ?1???????? Changes in coding of Eutretauranosuchus delfsi 2 – Skull geometry: relative position of tooth row, quadrate articular facet and occipital condyle – (? Unknown to 1 tooth row at a lower level than quadrate, aligned to occipital condyle) 3 – Skull geometry: relative position of tooth row and occipital condyle – (? Unknown to 0 unaligned) 4 – Skull geometry: relative position of quadrate and occipital condyles (? Unknown to 1 aligned) 8 – Rostrum: relation with the skull at maturity in lateral view (? Unknown to ½ Smooth decrease in height from skull/a poor fit) 28 – Foramen at premaxilla-maxilla suture (? Unknown to 0 absent) 29 – Perinarial crests: presence and morphology (? Unknown to 0 absent, surface even) 31 – Naris: orientation in the sagittal plane (? Unknown to 2 dorsal) 32 – Shape of naris (? Unknown to 0 subcircular) 33 – Naris distance to snout (? Unknown to 0 close) 34 – Presence of anterior narial notch (? Unknown to 0 absent) 35 – Presence of lateral narial notch (? Unknown to 0 absent) 36 – Composition of naris posterior border (? Unknown to 1 premaxilla) 37 – Presence and morphology of narial bar (? Unknown to 0 absent) 38 – Projection of internarial bar relative to premaxilla (? Unknown to – inapplicable) 39 – Presence of dorsal projection of anterior rami of premaxilla (? Unknown to 0 does not project) 59 – Presence of posterodorsal process of premaxilla at contact with maxilla and nasal (? Unknown to 1 present) 60 – Morphology of premaxilla-maxilla suture (? Unknown to 1 complex with anterior directed process from maxilla) 61 – Rostrum: presence of wedge like maxillary process (? Unknown to 1 present) 62 – Premaxilla morphology anterior to naris (? Unknown to 2 broadly meet at vertical wall) 65 – Shape of premaxilla in dorsal view (? Unknown to 3 paddle shaped) 95 – Lacrimal proportional length (? Unknown to 0 AP/ML closer to 2 or smaller) 97 – Posteroventral process of the lacrimal (? Unknown to 0 absent) 99 – Prefrontal-lacrimal crest dorsal to orbit (? Unknown to 0 absent) 100 – Transverse rostral crest: presence (? Unknown to 0 absent) 101 – Transverse frontal crest: presence (? Unknown to 0 absent) 104 – Crests margining the border of the sup-temp-fen (0 absent to 1 present) 160 – Orbit composition of anterior border (? Unknown to 2 lacrimal and prefrontal) 184 – Jugal fit to maxilla (? Unknown to 0 continuous) 196 – Postorbital bar: number of projections (? Unknown to [1/2] one/two projections) 204 – Quadratojugal participation in intertemporal foramen (? Unknown to 0 participates) 257 – Ectopterygoid posterior ramus (? Unknown to 0 reaches posterior tip) 335 – Participation of splenial in symphysis (? Unknown to 1 present) 340 – Dentary distal end (? Unknown to 1 long, reaches posteriorly to the orbit) 345 – Extension of surangular anterior lateral ramus (? Unknown to 1 long) 368/9 – Tooth cross-section (? Unknown to 1 circular) 370/1 – Faceted/laminar teeth (? Unknown to 0 absent) 378 – Carinae (1 present to 0 absent) .
Recommended publications
  • 8. Archosaur Phylogeny and the Relationships of the Crocodylia
    8. Archosaur phylogeny and the relationships of the Crocodylia MICHAEL J. BENTON Department of Geology, The Queen's University of Belfast, Belfast, UK JAMES M. CLARK* Department of Anatomy, University of Chicago, Chicago, Illinois, USA Abstract The Archosauria include the living crocodilians and birds, as well as the fossil dinosaurs, pterosaurs, and basal 'thecodontians'. Cladograms of the basal archosaurs and of the crocodylomorphs are given in this paper. There are three primitive archosaur groups, the Proterosuchidae, the Erythrosuchidae, and the Proterochampsidae, which fall outside the crown-group (crocodilian line plus bird line), and these have been defined as plesions to a restricted Archosauria by Gauthier. The Early Triassic Euparkeria may also fall outside this crown-group, or it may lie on the bird line. The crown-group of archosaurs divides into the Ornithosuchia (the 'bird line': Orn- ithosuchidae, Lagosuchidae, Pterosauria, Dinosauria) and the Croco- dylotarsi nov. (the 'crocodilian line': Phytosauridae, Crocodylo- morpha, Stagonolepididae, Rauisuchidae, and Poposauridae). The latter three families may form a clade (Pseudosuchia s.str.), or the Poposauridae may pair off with Crocodylomorpha. The Crocodylomorpha includes all crocodilians, as well as crocodi- lian-like Triassic and Jurassic terrestrial forms. The Crocodyliformes include the traditional 'Protosuchia', 'Mesosuchia', and Eusuchia, and they are defined by a large number of synapomorphies, particularly of the braincase and occipital regions. The 'protosuchians' (mainly Early *Present address: Department of Zoology, Storer Hall, University of California, Davis, Cali- fornia, USA. The Phylogeny and Classification of the Tetrapods, Volume 1: Amphibians, Reptiles, Birds (ed. M.J. Benton), Systematics Association Special Volume 35A . pp. 295-338. Clarendon Press, Oxford, 1988.
    [Show full text]
  • SUPPLEMENTARY ONLINE MATERIAL for a Re-Evaluation Of
    http://app.pan.pl/SOM/appXX-Halliday_etal_SOM.pdf SUPPLEMENTARY ONLINE MATERIAL FOR A re-evaluation of goniopholidid crocodylomorph material from Central Asia: Biogeographic and phylogenetic implications by Thomas J.D. Halliday, Marco Brandalise De Andrade, Michael J. Benton, and Mikhail B. Efimov Published in Acta Palaeontologica Polonica 201X 5X (X): xxx-xxx. http://dx.doi.org/10.4202/app.2013.0018 Character coding for redescribed material, and changes to E. delfsi Changes in coding of Eutretauranosuchus delfsi Character coding for redescribed material, and changes to E. delfsi All characters and character states are as found in Andrade et al. (2011). The following are the states assigned to Kansajsuchus extensus, “Sunosuchus” cf thailandicus (formerly “Sunosuchus” shartegensis), PIN 2229 (formerly Turanosuchus aralensis) and some changes to Eutretauranosuchus delfsi. Kansajsuchus extensus: 1[1/2]?1[1/2]???00110?????10000????00?200??1??10?0???????????1 ?12011121?3??????110010??????1??????1?????????01001??00- 300010010????0???????11012011?0??100?10010?0??[1/2]0?????????? ??????????????????????00?11?????11???????1??????2????????????? ??????????????????????0?0?100?0?????00?000111?000??1??00001220 0000?000010?20??????????????1????????????????????????????????? 0201000100?????0000021011?????000??0[1/2]?1??0???????????????0 ??00??000??????1011????????????????01?????1????????0???2?????0 1011???????? PIN 4174-1 1[1/5]??[1/2]0[0/1]10010???111?0000000?0??2???????????1??1??0? 1??10120????10???????110010?111101?100111?0?????0000?????????? ??00????????00?????110010?01?1????????????????????????????????
    [Show full text]
  • Genus/Species Skull Ht Lt Wt Time Range Adzhosuchus U.Jurassic Mongolia A. Fuscus U.Jurassic Mongolia Aegyptosuchus U.Cretaceous Egypt A
    Genus/Species Skull Ht Lt Wt Time Range Adzhosuchus U.Jurassic Mongolia A. fuscus U.Jurassic Mongolia Aegyptosuchus U.Cretaceous Egypt A. peyeri Cenomanian Egypt Aelodon see Aeolodon Aeollodon see Aeolodon Aeolodon U.Jurassic Germany A. priscus 16 cm 1.2 m? Kimmeridgian Germany Aggiosaurus U.Jurassic France A. nicaeensis U.Jurassic France Aigialosuchus U.Cretaceous Sweden A. villandensis Campanian Sweden Akanthosuchus Paleocene W USA A. langstoni Torrejonian New Mexico(US) Akantosuchus see Akanthosuchus A. langstoni see Akanthosuchus langstoni Albertochampsa 20 cm 1.6 m? U.Cretaceous Canada A. langstoni 20 cm 1.6 m? Campanian Alberta(Cnda) Aligator see Alligator Alligator 5.8 m Oligocene-Recent N America,China A. ameghinoi A. australis see Proalligator paranensis? A. cuvieri see Alligator mississippiensis A. darwini see Diplocynodon darwini A. gaudryi see Arambourgia gaudryi A. hantoniensis see Diplocynodon hantoniensis A. helois see Alligator mississippiensis A. heterodon see Crocodylus heterodon & Allognathosuchus heterodon A. lacordairei see Crocodylus acutus A. lucius see Alligator mississippiensis A. lutescens see Caiman lutescens A. mcgrewi 2 m Barstovian Nebraska(US) A. mefferdi Clarendonian Nebraska(US) A. mississipiensis living American Alligator M.Miocene-Recent Florida,Nebraska,Missouri,Georgia(US) A. olseni 25 cm 2.5 m? Hemingfordian Florida(US) A. parahybensis Pliocene Sao Paulo(Brazil) A. prenasalis 76 cm Chadronian S Dakota(US) A. sp. Arikareean Texas(US) A. sp. Barstovian Texas(US) A. sp. Duchesnean Texas(US) A. sp. Miocene Nebraska(US) A. styriacus see Crocodylus styriacus A. thompsoni(thomsoni) 36 cm 2.15 m Barstovian Nebraska(US) A. visheri 2 m Chadronian S Dakota(US) Alligatorellus 30 cm U.Jurassic Germany A.
    [Show full text]
  • Revisão Filogenética De Mesoeucrocodylia: Irradiação Basal E
    UNIVERSIDADE DE SÃO PAULO FFCLRP - DEPARTAMENTO DE BIOLOGIA PROGRAMA DE PÓS-GRADUAÇÃO EM BIOLOGIA COMPARADA Revisão filogenética de Mesoeucrocodylia: irradiação basal e principais controvérsias Felipe Chinaglia Montefeltro Tese apresentada à Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto da USP, como parte das exigências para a obtenção do título de Doutor em Ciências, Área: Biologia Comparada. RIBEIRÃO PRETO - SP 2013 UNIVERSIDADE DE SÃO PAULO FFCLRP - DEPARTAMENTO DE BIOLOGIA PROGRAMA DE PÓS-GRADUAÇÃO EM BIOLOGIA COMPARADA Revisão filogenética de Mesoeucrocodylia: irradiação basal e principais controvérsias Felipe Chinaglia Montefeltro Orientador: Max Cardoso Langer Tese apresentada à Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto da USP, como parte das exigências para a obtenção do título de Doutor em Ciências, Área: Biologia Comparada. RIBEIRÃO PRETO - SP 2013 FICHA CATALOGRÁFICA Montefeltro, Felipe Chinaglia Revisão filogenética de Mesoeucrocodylia: irradiação basal e principais controvérsias. Ribeirão Preto, 2013. 285 p. : il. ; 30 cm Tese de Doutorado, apresentada à Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto/USP. Área de concentração: Biologia Comparada. Orientador: Langer, Max Cardoso. 1. Crocodyliformes. 2. Mesoeucrocodylia 3. Metasuchia. 4. Notosuchia. 5. Pissarrachampsa . 6. Filogenia AGRADECIMENTOS Agradeço ao orientador Max Cardoso Langer pelo auxilio e oportunidade de desenvolver o projeto de doutorado sob sua tutela no Laboratório de Paleontologia da FFCLRP. Agradeço também ao orientador Hans C. E. Larsson pela oportunidade e auxilio durante o tempo desenvolvido no Redpath Museum da McGill University. Agradeço o suporte financeiro deste projeto às instituições: Fundação de Amparo à Pesquisa do Estado de São Paulo (FAPESP), Programa de Pós- Graduação em Biologia Comparada da FFCLRP e Laboratório de Paleontologia da FFCLRP.
    [Show full text]
  • A New Skeleton of the Neosuchian Crocodyliform Goniopholis with New Material from the Morrison Formation of Wyoming
    A NEW SKELETON OF THE NEOSUCHIAN CROCODYLIFORM GONIOPHOLIS WITH NEW MATERIAL FROM THE MORRISON FORMATION OF WYOMING by Bruce R. Erickson MONOGRAPH VOLUME 10: PALEONTOLOGY Published by THE SCIENCE MUSEUM OF MINNESOTA SAINT PAUL, MINNESOTA 55102 Frontispiece: Flesh restoration of Goniopholis. CONTENTS ABSTRACT ..................................................................7 INTRODUCTION .............................................................7 SYSTEMATIC PALEONTOLOGY ................................................7 DESCRIPTION ................................................................7 SCALATION ................................................................14 TAPHONOMY ...............................................................16 ACKNOWLEDGMENTS .......................................................19 TABLE 1 ....................................................................25 TABLE 2 ....................................................................25 TABLE 3 ....................................................................26 REFERENCES ...............................................................27 MONOGRAPH VOLUME 10: PALEONTOLOGY International Standard Book Number: 911338-90-X A NEW SKELETON of THE NEOSUCHIAN CROCODYLIFORM GONIOPHOLIS WITH NEW MATERIAL FROM THE MORRISON FORMATION OF WYOMING Bruce R. Erickson Fitzpatrick Chair of Paleontology MONOGRAPH VOLUME 10: PALEONTOLOGY The Science Museum of Minnesota 120 West Kellogg Blvd. Saint Paul, Minnesota 55102 USA. December 1, 2016 SCIENCE MUSEUM OF MINNESOTA
    [Show full text]
  • The Jurassic/Cretaceous Boundary: a Hidden Mass Extinction in Tetrapods?
    The Jurassic/Cretaceous boundary: a hidden mass extinction in tetrapods? Jonathan P. Tennant CID: 00661116 Imperial College London Department of Earth Science and Engineering Thesis submitted to fulfil the requirements for the degree of Doctor of Philosophy and Diploma of Imperial College Image credit: Robert Nicholls (CC BY 4.0). Depicts Sarcosuchus imperator, a giant predatory crocodyliform from the Cretaceous of North Africa. 1 Declaration of originality I declare that the works presented within this thesis are my own, and that all other work is appropriately acknowledged and referenced within. Copyright declaration The copyright of this thesis rests with the author, and it is made available under a Creative Commons Attribution (CC BY 4.0) license. Researchers are free to copy, distribute and transmit this thesis on the condition that it is appropriately attributed. Jonathan Peter Tennant Supervisors: Dr. Philip Mannion (Imperial College London); Prof. Paul Upchurch (University College London); Dr. Mark Sutton (Imperial College London). 2 Acknowledgements First and definitely foremost, I want to extend my greatest thanks to Phil Mannion. As his first PhD student, I am sure he regretted his decision after day one, but stuck with it until the end, and has been a stoic mentor throughout. This project would have been a shadow of what it came to be without his guidance. I am still yet to get him on Twitter though. I am also hugely grateful to Paul Upchurch and Mark Sutton for their input and experience throughout this project. I also could not have completed this project without the encouragement and support from my girlfriend, friends, and family, and am hugely grateful to them.
    [Show full text]
  • Choristoderes and the Freshwater Assemblages of Laurasia
    ISSN (print): 1698-6180. ISSN (online): 1886-7995 www.ucm.es/info/estratig/journal.htm Journal of Iberian Geology 36 (2) 2010: 253-274 doi:10.5209/rev_JIGE.2010.v36.n2.11 Choristoderes and the freshwater assemblages of Laurasia Coristoderos y las asociaciones de agua dulce de Laurasia R. Matsumoto, S. E. Evans Department of Cell and Developmental Biology, University College London, Gower Street, London, WC1E 6BT, England [email protected]; [email protected] Received: 11/11/09 / Accepted: 30/06/10 Abstract Choristoderes are freshwater aquatic diapsid reptiles of uncertain phylogenetic position. Although the lineage probably diverged from other diapsids in the Permo-Triassic, choristoderes apparently never reached Gondwana. Within Laurasia, however, they have been recovered from Japan, China, Mongolia, Trans-Baikal Russia, Central Asia, Western Europe, and North America, reaching ex- treme northern latitudes. Setting aside controversial Triassic records, their known temporal range currently extends from the Middle Jurassic (Britain, Kyrgyzstan) to the Miocene (Czech Republic). However, although small choristoderes are known to span the entirety of this period, the larger, more derived neochoristoderes are recorded only from the Early Cretaceous through to the earliest Eocene. The gavial-like neochoristodere Champsosaurus is the most familiar taxon, characterised by large size, a long rostrum and flared temporal fenestrae, but research over the last three decades has revealed many new genera and exposed an unexpected diver- sity in terms of body size (small to relatively large), neck length (long v. short) and skull morphology (longirostrine v. brevirostrine, open v. closed lower temporal fenestrae). Typically choristoderes occur as part of a mesic assemblage that includes fish, lizards, mammals, turtles, frogs, salamanders, small dinosaurs and, usually, crocodiles.
    [Show full text]
  • (Crocodylomorpha: Neosuchia): Implications for the Rise of Eusuchia
    Zoological Journal of the Linnean Society, 2016, 177, 854–936. With 11 figures Evolutionary relationships and systematics of Atoposauridae (Crocodylomorpha: Neosuchia): implications for the rise of Eusuchia JONATHAN P. TENNANT1*, PHILIP D. MANNION1 and PAUL UPCHURCH2 1Department of Earth Science and Engineering, Imperial College London, South Kensington, London SW7 2AZ, UK 2Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, UK Received 18 August 2015; revised 5 January 2016; accepted for publication 19 January 2016 Atoposaurids are a group of small-bodied, extinct crocodyliforms, regarded as an important component of Jurassic and Cretaceous Laurasian semi-aquatic ecosystems. Despite the group being known for over 150 years, the taxonomic composition of Atoposauridae and its position within Crocodyliformes are unresolved. Uncertainty revolves around their placement within Neosuchia, in which they have been found to occupy a range of positions from the most basal neosuchian clade to more crownward eusuchians. This problem stems from a lack of adequate taxonomic treatment of specimens assigned to Atoposauridae, and key taxa such as Theriosuchus have become taxonomic ‘waste baskets’. Here, we incorporate all putative atoposaurid species into a new phylogenetic data matrix comprising 24 taxa scored for 329 characters. Many of our characters are heavily revised or novel to this study, and several ingroup taxa have never previously been included in a phylogenetic analysis. Parsimony and Bayesian approaches both recover Atoposauridae as a basal clade within Neosuchia, more stemward than coelognathosuchians, bernissartiids, and paralligatorids. Atoposauridae is a much more exclusive clade than previously recognized, comprising just three genera (Alligatorellus, Alligatorium, and Atoposaurus) that were restricted to the Late Jurassic of western Europe, and went extinct at the Jurassic/Cretaceous boundary.
    [Show full text]
  • Calsoyasuchus Valliceps, a New Crocodyliform from the Early Jurassic Kayenta Formation of Arizona
    Journal of Vertebrate Paleontology 22(3):593±611, September 2002 q 2002 by the Society of Vertebrate Paleontology CALSOYASUCHUS VALLICEPS, A NEW CROCODYLIFORM FROM THE EARLY JURASSIC KAYENTA FORMATION OF ARIZONA RONALD S. TYKOSKI1,2, TIMOTHY B. ROWE1,2,3, RICHARD A. KETCHAM1,3, and MATTHEW W. COLBERT1,2,3 1Jackson School of Geosciences, [email protected]; 2Texas Memorial Museum Vertebrate Paleontology Laboratory; 3High Resolution X-ray CT Facility, The University of Texas at Austin, Austin, Texas 78712 ABSTRACTÐWe describe a new fossil crocodyliform archosaur from the Early Jurassic Kayenta Formation of the Navajo Nation that is surprisingly derived for so ancient a specimen. High-resolution X-ray CT analysis reveals that its long snout houses an extensive system of pneumatic paranasal cavities. These are among the most distinctive features of modern crocodylians, yet the evolutionary history of this unique system has been obscured by the inaccessibility of internal structures in most fossil crania. Preliminary phylogenetic analysis indicates that the new species is the oldest known member of a monophyletic Goniopholididae, and within this lineage to be the sister taxon of Eutretauranosu- chus, from the Late Jurassic Morrison formation of Colorado. Goniopholididae became extinct at the end of the Cretaceous, but it is more closely related to living crocodylians than are several lineages known only from Cretaceous and younger fossils. The new taxon nearly doubles the known length of goniopholid history and implies a deep, as yet undiscovered, Mesozoic history for several crocodyliform lineages that were once thought to have relatively com- plete fossil records. INTRODUCTION 7) for this report.
    [Show full text]
  • Cranial Osteology and Phylogenetic Relationships of Hamadasuchus Rebouli (Crocodyliformes: Mesoeucrocodylia) from the Cretaceous of Morocco
    Blackwell Publishing LtdOxford, UKZOJZoological Journal of the Linnean Society0024-4082© 2007 The Linnean Society of London? 2007 1494 533567 Original Articles HAMADASUCHUS REBOULIH. C. E. LARSSON and H.-D. SUES Zoological Journal of the Linnean Society, 2007, 149, 533–567. With 9 figures Cranial osteology and phylogenetic relationships of Hamadasuchus rebouli (Crocodyliformes: Mesoeucrocodylia) from the Cretaceous of Morocco HANS C. E. LARSSON1* and HANS-DIETER SUES2 FLS 1Redpath Museum, McGill University, 859 Sherbrooke Street W., Montréal, QC H3A 2K6, Canada 2National Museum of Natural History, Smithsonian Institution, NHB MRC 106, PO Box 37012, Washington, DC 20013–7012, USA Received February 2005; accepted for publication June 2006 This paper presents a detailed description of the skull and part of the mandible of the crocodyliform reptile Hama- dasuchus rebouli from the Kem Kem beds (Upper Cretaceous: Albian–Cenomanian) of south-eastern Morocco. This taxon of deep-snouted ziphodont crocodyliform can be diagnosed by a number of autapomorphies. Phylogenetic anal- ysis of a diverse array of crocodylomorph taxa found strong support for a clade comprising H. rebouli, Peirosauridae, and Sebecus. The name Sebecia nom. nov. is proposed for this grouping, which is diagnosed by numerous charac- ters, including the participation of the quadratojugal in the mandibular condyle. The distribution of this diverse and long-lived clade lends further support to the biogeographical hypothesis that faunal connections existed between Africa and South America well into mid-Cretaceous times. © 2007 The Linnean Society of London, Zoological Journal of the Linnean Society, 2007, 149, 533–567. ADDITIONAL KEYWORDS: Crocodylomorpha – Hamadasuchus – skull. INTRODUCTION Araripesuchus wegeneri (Buffetaut & Taquet, 1979; Buffetaut, 1981a; Ortega et al., 2000; referred to The evolutionary history of Mesozoic crocodyliform Hamadasuchus by Prasad & Lapparent de Broin, reptiles from Africa is still poorly understood.
    [Show full text]
  • A NEW ATOPOSAURID CROCODYLOMORPH from the MORRISON FORMATION (UPPER JURASSIC) of WYOMING, USA John R
    GEOLOGY OF THE INTERMOUNTAIN WEST an open-access journal of the Utah Geological Association ISSN 2380-7601 Volume 5 2018 A NEW ATOPOSAURID CROCODYLOMORPH FROM THE MORRISON FORMATION (UPPER JURASSIC) OF WYOMING, USA John R. Foster Theme Issue An Ecosystem We Thought We Knew— The Emerging Complexities of the Morrison Formation SOCIETY OF VERTEBRATE PALEONTOLOGY Annual Meeting, October 26 – 29, 2016 Grand America Hotel Salt Lake City, Utah, USA © 2018 Utah Geological Association. All rights reserved. For permission to copy and distribute, see the following page or visit the UGA website at www.utahgeology.org for information. Email inquiries to [email protected]. GEOLOGY OF THE INTERMOUNTAIN WEST an open-access journal of the Utah Geological Association ISSN 2380-7601 Volume 5 2018 Editors UGA Board Douglas A. Sprinkel Thomas C. Chidsey, Jr. October 2018 – September 2019 Utah Geological Survey Utah Geological Survey President Peter Nielsen [email protected] 801.537.3359 801.391.1977 801.537.3364 President-Elect Leslie Heppler [email protected] 801.538.5257 [email protected] [email protected] Program Chair Gregory Schlenker [email protected] 801.745.0262 Treasurer Dave Garbrecht [email protected] 801.916.1911 Bart J. Kowallis Steven Schamel Secretary George Condrat [email protected] 435.649.4005 Past President Paul Inkenbrandt [email protected] 801.537.3361 Brigham Young University GeoX Consulting, Inc. 801.422.2467 801.583-1146 [email protected] [email protected] UGA Committees Environmental Affairs
    [Show full text]
  • (Crocodyliformes, Eusuchia): Phylogenetic Implications
    The palate and choanae structure of the Susisuchus anatoceps (Crocodyliformes, Eusuchia): phylogenetic implications Karla J. Leite1,* and Daniel C. Fortier2,* 1 Departamento de Geologia, Universidade Federal do Ceará, Fortaleza, Ceará, Brazil 2 Departamento de Ciências Biológicas, Universidade Federal do Piauí, Floriano, Piauí, Brazil * These authors contributed equally to this work. ABSTRACT Crocodyliformes is a group with a broad fossil record, in which several morphological changes have been documented. Among known transformations the most iconic is perhaps the series of changes seen in the structural evolution of the choanae. The change in the position of the choanae was important during the evolutionary history of the Crocodyliformes. This structure is relevant in the phylogenetic position of many crocodyliforms. The new skull of Susisuchus anatoceps from the Crato Formation of the Santana Group (Lower Cretaceous) is described and the preservation in the ventral view allows character encoding not yet observed for the species. The new specimen shows a typical eusuchian palate for Susisuchus anatoceps, in which the choana is fully enclosed by the pterygoid. The Susisuchidae clade has been placed in different phylogenetic positions: as a sister group of Eusuchia, advanced Neosuchia and in Eusuchia. In Isisfordia there are reports that the choana of this taxon is or is not fully enclosed by the pterygoid. The encoding of the ventral characters of S. anatoceps places Susisuchidae in Eusuchia. However, this position must be further studied, since the matrices showed fragility in the reconstitution of the Neosuchia–Eusuchia transition. Subjects Paleontology Submitted 22 February 2018 Keywords Crocodyliformes, Eusuchia, Lower Cretaceous, Araripe Basin Accepted 12 July 2018 Published 10 August 2018 Corresponding author INTRODUCTION Karla J.
    [Show full text]