174 PRINCIPES [VOL. 26

Principes.26(4). 1982. pp. 174-177 Pogonotium moorel, aNew Species from Sarawak

JOHN DRANSFIELD Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey 7W9 3AB, UK

In my article "A Day on the Klingkang close, oblique and horizontal, complete or Range" (Dransfield 1982) I mentioned the partial whorls of horizontal or reflexed excitement of discovering a possible new spines and abundant, pale brown, floccose species of Pogonotium from the summit indumentum; spines varying from minute of Gunung Gaharu in Sarawak. I have black spicules scarcely 1 mm long, to sin­ now been able to examine material of this uous papery, black or straw colored spines curious relative in detail and here to 22 X 1 mm with swollen bases; knee describe it as new. I name it for Hal scarcely developed; auricles very conspic­ Moore, who was much in my thoughts on uous, one on each side of the petiole, erect, the day I discovered it and who, I hope, to 8.5 cm long, ca. 1.5 cm wide at the would have been equally excited by its base, tapering to ca. 5 mm wide for much strange morphology. of the length, channeled adaxially, armed as the sheath. Leaf to ca. 1.1 m; petiole to 50 cm, 7 mm wide at the base, Pogonotium moorei J. Dransfield, sp. tapering to ca. 5 mm, flattened adaxially nov. (Fig. 1). near the base, rounded in distal region, Palmijuncus exiguus a P. ursino et P. covered with inflated hairs and scabrid due divaricato caule brevi, folio foliola c. 9, to papillae, and bearing sparse, mostly lat­ lata et remota divaricataque ferenti, fructu eral or abaxial solitary or paired, short, obpyriformi seriebus verticalibus 15-16 reflexed, bulbous-based black-tipped spines squamarum aliquantum convexarum tecto to 5 mm; rachis as the petiole, tapering, bene distincta. Typus: BORNEO, SARA­ very sparsely armed, sometimes ending in WAK, G. Gaharu, ]. Dransfield et al. a minute cirrus to 15 mm; leaflets distant, jD6102 (holotypus K; isotypi BH, BO, L, very regularly arranged, ca. 9 pairs in all, PNH, SAR). alternate proximally, distally in opposite, Solitary, slender, pleonanthic, spiny, divaricate pairs, ca. 20-32 X 2.2-2.5 cm, dioecious palm with short erect stem to 2 except for the smaller apical pair ca. 15 X m only; stem without sheaths ca. 10 mm 2.5 cm; leaflet margin armed with sparse diam., with sheaths ca. 18 mm, internodes short bristles; adaxial surface smooth, ca. 5 cm long. Leaf sheaths splitting oppo­ abaxial surface matt,. minutely dotted, site the petiole, armed with abundant, sparsely indumentose along the fold;

I. Pogonotium moorei. A, leaf sheath with base of petiole and auricles X%; B, sheath with removed from sheath in A X%; C. petiole and base of first leaflet X%; D, leaf tip X%; E, tip of rachilla with pistillate X4; F, sterile staminate flower dissected X4; G, X4; H, vertical section of X2; J. seedling X%. Drawn from Dransfield JD6102 by Mary Millar Watt. 176 PRINCIPES [VOL. 26

Table 1. Comparison ojspecies oj Pogonotium

Pogonotium ursinum Pogonotium divaricatum Pogonotium moorei Leaflets 90-100 40-50 ca. 9 close, ciliate narrow, rather distant, broad, remote, bristly, limp sparsely bristly, rigid very sparsely bristly, rigid Mature fruit ± barrel shaped globose obpyriform Scales ± flattened, grossly swollen and slightly convex, pinkish-brown, grooved, magenta, in pinkish brown, in in 16-17 21-22 vertical rows 16-17 vertical vertical rows rows

transverse veinlets conspicuous. Inflores­ mm, covered in pale brown scales tipped cence, only pistillate known, erect, held with 3 sinuous fleshy stigmas ca. 3.5 X between the auricles, ca. 18 cm long; 0.5 mm. Mature fruit borne on persistent peduncle adnate to sheath of following leaf, perianth whorls, obpyriform, ca. 13 X 9 with free portion lOX 7 mm; prophyll mm, tipped with a short beak ca. 1 X 2 beaked, ca. 17 cm, split longitudinally mm, and stigmatic remains; pericarp cov­ along adaxial (always?) face, covered in ered in 15-16 vertical rows of pinkish­ dense indumentum and in distal ¥3 bearing brown scales; seed basally attached, ellip­ partial whorls of spines as on sheath; two soidal, ca. 11 X 7 mm; sarcotesta thin; primary bracts also present, included homogeneous, basal. within prophyll and much smaller, and Germination adjacent-ligular; eophyll with more sparsely armed; prophyll and pri­ 2 small divergent leaflets, ca. 15 X 3 mm, mary bracts subtending 1st order tipped with bristles. branches, the inflorescence branching to BORNEO: SARAWAK, 1st Division, 2 orders in all; rachillae sinuous, bearing Serian District, Sabal Tapang Forest short tubular, triangular tipped, striate, Reserve, G. Gaharu, summit plateau, brown bracts each subtending a diad of montane forest developed on sandstone sterile staminate and fertile pistillate flow­ blocks, alt. 700 m above sea level, ]. ers, all ± included in the prophyll. Sterile Dransfield et al. JD6102 (holotype K; staminate flower ca. 4.5 mm long, trigo­ isotypes BR, BO, L, PNR, SAR). nous; calyx striate, ca. 2 mm high, shal­ Pogonotium moorei differs from the lowly 3-lobed; corolla split almost to the two other species of the genus, P. ursinum base into 3 narrow triangular lobes to 4 X (Becc.) J. Dransf. and P. divaricatum J. 1.5 mm; sterile 6, filaments fleshy Dransf. in the low habit, the leaf with only ca. 1.5 X 0.4 mm; anthers oblong ca. about 9 broad, remote divaricate leaflets 0.9 X 4 mm, pistillode minute. Pistillate (P. ursinum has about 90-100 close leaf­ flower ca. 7.5 X 3 mm; calyx with a basal lets and P. divaricatum about 40-50), tube ca. 2.2 mm and 3 broad triangular and the obpyriform fruit with about 15­ lobes ca. 1.5 X 2 mm; corolla with a basal 16 vertical rows of slightly convex scales. tube ca. 2.0 mm long and 3 striate lobes The seedling leaf of the new species with ca. 5 X 2.5 mm; staminodes 6, epipetal­ its single pair of minute divergent leaflets ous, filaments flattened, ± triangular ca. is very different from the pinnate eophyll 1.5 X 1 mm, empty anthers elongate, ca. of P. ursinum; unfortunately, the eophyll 0.75 X 2 mm; ovary ovoid ca. 2.5 X 1.8 of P. divaricatum is not known. These 1982] DRANSFIELD: POGONOTIUM 177 diagnostic features of P. moorei may be LITERATURE CITED contrasted with those of P. ursinum and P. divaricatum in Table 1 and in my DRANSFIELD, J. 1980. Pogonotium (Palmae: Lep­ idocaryoideae) a new genus related to Dae­ account of Pogonotium (Dransfield 1980). monorops. Kew Bulletin 34: 761-768. ---. 1982. A Day on the Klingkang Range, Principes 26: 19-33.

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