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Arquivos De Zoologia ISSN 0066-7870 Arquivos de Zoologia COMPARATIVE MORPHOLOGICAL STUDY OF REPRESENTATIVES OF THE THREE FAMILIES OF STROMBOIDEA AND THE XENOPHOROIDEA (MOLLUSCA, CAENOGASTROPODA), WITH AN ASSESSMENT OF THEIR PHYLOGENY Luiz Ricardo L. Simone VOLUME 37, FASCÍCULO 2 MUSEU DE ZOOLOGIA DA UNIVERSIDADE DE SÃO PAULO SÃO PAULO, 2005 UNIVERSIDADE DE SÃO PAULO MUSEU DE ZOOLOGIA Prof. Dr. Adolpho José Melfi Prof. Dr. Carlos Roberto Ferreira Brandão Reitor Diretor Prof. Dr. Hélio Nogueira da Cruz Profª. Drª. Eliana Marques Cancello Vice-Reitor Vice-Diretora Prof. Dr. Adilson Avansi de Abreu Pró-Reitor de Cultura e Extensão Universitária COMISSÃO EDITORIAL Hussam El Dine Zaher (editor) EDITORES ASSOCIADOS Antonio Carlos Marques (Universidade de São Paulo) Mário de Pinna (Universidade de São Paulo) Sérgio Antonio Vanin (Universidade de São Paulo) SEÇÃO DE PUBLICAÇÕES Airton de Almeida Cruz (arte-finalista) Marly dos Reis (bibliotecária) NORMALIZAÇÃO BIBLIOGRÁFICA Teresa Beatriz Nunes Guimarães (bibliotecária) Maria Regina de Mendonça Ferreira (bibliotecária) MUSEU DE ZOOLOGIA DA UNIVERSIDADE DE SÃO PAULO Av. Nazaré, 481, Ipiranga, CEP 04263-000, São Paulo, SP, Brasil Publicado com o apoio financeiro da Fundação de Amparo à Pesquisa do Estado de São Paulo (FAPESP), Programa de Apoio às Publicações Científicas Periódicas da USP (Comissão de Credenciamento) e Museu de Zoologia da Universidade de São Paulo (MZUSP). Arquivos de Zoologia. v.15- 1967- Museu de Zoologia da Universidade de São Paulo. v. ilust. ISSN 0066-7870 Continuação de: Arquivos de Zoologia do Estado de São Paulo, v.1-14; 1960-66. 1. Zoologia. I. Museu de Zoologia, USP. As instruções aos autores são publicadas em inglês e português no fascículo introdutório de cada volume, assim como a lista dos consultores que assessoram a Comissão Editorial. Arquivos de Zoologia MUSEU DE ZOOLOGIA DA UNIVERSIDADE DE SÃO PAULO CONTEÚDO LUIZ RICARDO L. SIMONE COMPARATIVE MORPHOLOGICAL STUDY OF REPRESENTATIVES OF THE THREE FAMILIES OF STROMBOIDEA AND THE XENOPHOROIDEA (MOLLUSCA, CAENOGASTROPODA), WITH AN ASSESSMENT OF THEIR PHYLOGENY ................................ 141 ISSN 0066-7870 Arq. Zool., S. Paulo São Paulo v. 37 n. 2 p. 141-267 2005 Contents Abstract ............................................................................................................................................... 141 Introduction ........................................................................................................................................ 142 Material and Methods ......................................................................................................................... 142 Systematics.................................................................................................................................... 155 Family STROMBIDAE ........................................................................................................... 155 Genus Strombus Linné, 1758............................................................................................. 155 Strombus pugilis Linné, 1758 ...................................................................................... 155 Strombus alatus Gmelin, 1790 ..................................................................................... 164 Strombus gracilior Sowerby, 1825 ............................................................................... 169 Genus Eustrombus Wenz, 1940 ......................................................................................... 170 Eustrombus goliath (Schröter, 1805) new comb. ......................................................... 170 Eustrombus gigas (Linné, 1758) .................................................................................. 178 Genus Aliger Thiele, 1929 ................................................................................................. 181 Aliger costatus (Gmelin, 1791) new comb................................................................... 181 Aliger gallus (Linné, 1758) .......................................................................................... 185 Genus Tricornis Jousseaume, 1886 ................................................................................... 189 Tricornis raninus (Gmelin, 1791) ................................................................................. 189 Genus Conomurex P. Fisher, 1884..................................................................................... 192 Conomurex luhuanus Linné, 1758 ............................................................................... 192 Genus Canarium Schumacher, 1817 .................................................................................. 196 Canarium urceus Linné, 1758 ...................................................................................... 196 Genus Lambis Röding, 1798 ............................................................................................. 199 Lambis lambis (Linné, 1758) ....................................................................................... 199 Genus Terebellum Röding, 1798 ....................................................................................... 203 Terebellum terebellum (Linné, 1758) .......................................................................... 203 Genus Tibia Röding, 1798 ................................................................................................. 207 Tibia insulaechorab Röding, 1798 ............................................................................... 207 Family STRUTHIOLARIIDAE .............................................................................................. 209 Genus Tylospira Harris, 1897 ............................................................................................ 209 Tylospira scutulata (Gmelin, 1791) .............................................................................. 209 Genus Struthiolaria Lamarck, 1816 ................................................................................... 215 Struthiolaria papulosa (Martyn, 1784) ......................................................................... 215 Family APORRHAIDAE ........................................................................................................ 217 Genus Cuphosolenus Piette, 1876 ..................................................................................... 217 Cuphosolenus (?) serresianus (Michaud, 1828) new comb. ........................................ 217 Genus Aporrhais DaCosta, 1778 ....................................................................................... 222 Aporrhais occidentalis Beck, 1836 .............................................................................. 222 Aporrhais pespelicani (Linné, 1758)............................................................................ 228 Family XENOPHORIDAE ..................................................................................................... 231 Genus Onustus Swainson, 1840 ......................................................................................... 231 Onustus caribaeus (Petit de la Saussaye, 1856) ........................................................... 231 Onustus indicus (Gmelin, 1971) .................................................................................. 238 Genus Xenophora Waldeheim, 1807 ................................................................................. 240 Xenophora conchyliophora (Born, 1780) .................................................................... 240 Discussion of Characters .................................................................................................................... 247 Shell......................................................................................................................................... 247 Head-foot................................................................................................................................. 248 Operculum ............................................................................................................................... 251 Mantle organs .......................................................................................................................... 252 Kidney ..................................................................................................................................... 254 Digestive system ...................................................................................................................... 255 Buccal mass and odontophore ................................................................................................. 255 Radula ..................................................................................................................................... 256 Salivary glands ........................................................................................................................ 256 Esophagus................................................................................................................................ 257 Stomach ................................................................................................................................... 257 Intestine ................................................................................................................................... 258 Male genital system ................................................................................................................. 258 Female genital
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