lt#Mnrt A CATALOG OF THE COLEÓPTERA OF AMERICA NORTH OF MEXICO

FAMILY:

cz: -

rr- w ■• . CX3 CD. PN^ CD: CO

^^i^-. UNITED STATES AGRICULTURE PREPARED BY iliàM DEPARTMENT OF HANDBOOK AGRICULTURAL "^^^ AGRICULTURE NUMBER 529-95 RESEARCH SERVICE FAMILIES OF COLEóPTERA IN AMERICA NORTH OF MEXICO

Fascicle ' Family Year issued Fascicle ' Family Year issued Fascicle ' Family Year issued

1 Cupedidae 1979 45 Chelonariidae 98 Endomychidae _. 2 Micromalthidae 46 Callirhipidae 100 Lathridiidae 3 Carabidae 47 Heteroceridae 1978 102 Biphyllidae 4 Rhysodidae 48 Limnichidae 103 Byturidae 5 Amphizoidae 49 Dryopidae 104 Mycetophagidae 6 Haliplidae 50 Elmidae 105 Ciidae 1982 8 Noteridae 51 Buprestidae 9 Dytiscidae 52 Cebrionidae 107 Prostomidae 10 Gyrinidae 53 Elateridae 109 Colydiidae 13 Sphaeriidae 54 Throscidae 110 Monommatidae 14 Hydroscaphidae 55 Cerophytidae 111 Cephaloidae 15 56 Perothopidae 112 Zopheridae 16 Hydrophilidae 57 Eucnemidae 115 Tenebrionidae _- 17 Georyssidae 58 Telegeusidae 116 AUeculidae 18 Sphaeritidae 61 Phengodidae 117 Lagriidae 20 Histeridae 62 Lampyridae 118 Salpingidae 21 Ptiliidae 63 Cantharidae 119 Mycteridae 22 Limulodidae 64 Lycidae 120 Pyrochroidae —. 23 l>asyceridae 65 121 Othniidae 24 Micropeplidae 66 Nosodendridae 122 Inopeplidae 25 .— Leptinidae 67 Dermestidae 123 Oedemeridae — - 26 Leiodidae 69 Ptinidae 124 Melandryidae __ 27 Scydmaenidae 70 Anobiidae 125 Mordellidae 28 Silphidae 71 Bostrichidae 126 Rhipiphoridae __ 29 Scaphidüdae 72 Lyctidae 127___ Meloidae 30 Staphylinidae 74 Trogositidae 128 Anthicidae 31 Pselaphidae 76 Cleridae 129 Pedilidae 32 Lucanidae 78 Melyridae 130 Euglenidae 33 Passalidae 79 131 Cerambycidae _ _ 34 Scarabaeidae 81 Sphindidae 132 Bruchidae 35 Eucinetidae 82 Nitidulidae 133 Chrysomelidae _- 36 Helodidae 83 Rhizophagidae 134 Nemonychidae __ 37 86 Cucujidae 135 Anthribidae 38 Dascillidae 90 Cryptophagidae 138 Allocorynidae __ 39 Rhipiceridae 92 Languriidae 140 Brentidae 4p-___Byrrhidae 93 Erotylidae 141 Platypodidae _„ —- 1979 41 Psephenidae 94 Phalacridae 142 Scolytidae 42 Brachypsectridae 95 Cerylonidae 1982 143 Curculionidae _. 43 Artematopidae 96 Corylophidae 144 Stylopidae 44 97 Coccinellidae 145 Fossil Coleóptera

' Missing numbers are those assigned in the computer program to families not found in the United States and Canada.

For sale by the Superintendent oí Documents, U.S. Qovemment Printing Office, Washington, D.C. 20402 ^A CATALOG OF THE COLEÓPTERA OF AMERICA NORTH OF MEXICO, FAMILY: CERYLONIDAE i 1 ^

BY s j. F./LAWRENCE [r DEPARTMENT OF ENTOMOLOGY, CSIRO I CANBERRA, AUSTRALIA

UNITED STATES AGRICULTURE PREPARED BY DEPARTMENT OF HANDBOOK AGRICULTURAL AGRICULTURE NUMBER 529-95 RESEARCH SERVICE

October 1982 FOREWORD

Many species of are important pests of agricultural crops, stored food products, forests, wood products and structures, and fabrics. Many other species, in contrast, are beneficial in the biological sup- pression of pest and weeds, as well as in the decomposition of plant detritus, carcasses, and dung. Part of our national responsibility to American agriculture is to provide correct identification of species of American beetles so that appropriate controls can be apphed. Most information about animal species, whether agricultural, biological, or experimental, is filed under the species' scientific names. These names are therefore the keys to retrieval of such information. Because some species have been known by several names, a complete listing of these names for each species is neces- sary. For the user of scientific names, an up-to-date taxonomic catalog providing currently accepted names and pertinent bibliographic and distributional data is an indispensable tool. Although taxonomic hterature is constantly changing to reflect current work, the traditional published taxonomic catalog remains static with updating left to the individual user until it is revised. Production of catalogs in the past has been laborious with long printing delays resulting in data that are obsolete before being published. However, the computer now provides the capabihty of storing, updating, and retrieving taxonomic data; rapid publica- tion through computer-driven typesetting machinery; and a greater degree of currentness and flexibility. All 124 fascicles in this catalog of the beetles of America north of Mexico are produced by an original group of computer programs, designed and written during a pilot project by personnel of the Systematic Entomology Laboratory, Agricultural Research Service, and the Communications and Data Services Divi- sion, Science and Education Management Staff. The published information is stored on computer tape, is updated periodically to reflect taxonomic progress in the family, and is available in a data base for computer searching.

T. B. Kinney, Jr. Administrator Agricultural Research Service PREFACE The Coleóptera, or beetles, are represented in the world by about 220,000 described species, of which about 24,000 occur in the United States and Canada. A comprehensive taxonomic catalog of beetles for this area has not been available except the series of world-based "Coleopterorum Catalogus" volumes (1909-present, Junk, Berlin). The Leng ''Catalogue of the Coleóptera of America North of Mexico" (J. D. Sherman, Jr., Mt. Vemon, NY), which was published in 1920 with supplements to the end of 1947, is a checklist. However, it has served professional and amateur alike for nearly 60 years as the principal source of scientific names of beetles. Since 1947, many new taxa have been described and many changes in status and nomenclature have appeared in numerous scattered publications, but little effort has been made to summarize these changes. This catalog will supplant the Leng catalog and supply additional essential information. It is produced by an original suite of storage, retrieval, and printing programs written especially for automated taxonomic catalogs. The catalog for each family is published as a separate fascicle with its introductory text, bibliography, and index. Each family is numbered as listed, but the order of issuance of fascicles is not necessarily in numerical sequence. The publishing of separate fascicles makes data available shortly after they are assembled. Computer tapes for each fascicle are maintained for updating and necessary reprinting. The information on each family is the responsibility of the respective author or authors. The editors modify it only to correct obvious errors and to make it conform to the requirements of the computer programs. No original proposal for a new name, taxon, status, or classification is given, such data having been previously published, but new host and distributional data are often listed. The rules of "The International Code of Zoological Nomenclature" are followed. The geographic scope of this catalog includes the continental United States, Canada, Alaska, Greenland, and the associated continental islands. Names of taxa found only in other regions are excluded. If the range of a species extends outside these geographic limits, this fact is indicated, inside the back cover is a map of the 12 faunal regions based on historical and faunal criteria to simplify distribution recordings. Two-letter Postal Service style abbreviations are used for States and Provinces, and faunal regions are indicated in each distribution record by a diagonal line between groups of abbreviations. It is not the purpose of this catalog to present a complete scheme of higher classification within the order. The familial makeup is somewhat intermediate between that of R, H. Arnett in "The Beetles of the United States" (1960-62, Catholic University Press, Washington, DC) and that of R. A. Crowson in "The Natural Classification of the Families of Coleóptera" (1967, Biddies Ltd., Guildford, England). Modifications of these two systems are largely those advocated by J. F. Lawrence based in part on suggestions by taxonomic specialists for certain families. Generic groups and higher categories within the family are arranged phylogenetically as indicated by the author of the particular fascicle, and species group names with their respective synonyms are arranged alphabetically. Names referable to incertae sedis and nomen dubium are listed separately at the end of the nearest applicable taxon with notations as to their status. Each available name is followed by its author, date proposed, and page number referring to the complete bibliographic citation containing the original description. Following each generic name are vn the type-species and method of its designation, necessary explanatory notes, and pertinent references on immature stages, , redescription, ecology, and keys. After the specific name entry are the original genus (if different from the present placement), type-locality, geographical distribution by State, Province, and broad extralimital units, explanatory notes, pertinent references to immature stages, taxonomy, redescription, and ecology, depository of type-specimen and its sex, and hosts. In addition to the list under the map of faunal regions (inside back cover), the following abbrevia- tions are used in this catalog:

ABBREVIATIONS, GENERAL

Amer. Bor.—America Borealis Mus.—Museum Amer. Sept.—America Septentrionalis N. Amer.—North America Autom.—Automatic Orig. des.—Original designation C. Amer.—Central America Preocc.—Preoccupied Co.—County S. Amer.—South America Cosmop.—Cosmopolitan Sp.—Species Design.—Designated Subseq. monot.—Subsequent monotypy F.—Female Subsp.—Subspecies Holarc.—Holarctic Taut.—Tautonymy Isl.—Island Univ.—^University M.—Male USA—United States of America Mex.—Mexico Var.—Variety Monot.—Monotypy W. Ind.—West Indies

MUSEUMS IN THE CONTINENTAL UNITED STATES AND CANADA ^

AMNH—American Museum of Natural History, FSCA—Florida State Collection, Gainesville New York HAHC—H. & A. Howden Collection, Ottawa, ANSP—^Academy of Natural Sciences, Phila- Canada delphia, PA ICCM—Carnegie Museum, Pittsburgh, PA BYUC—Brigham Young University, Provo, UT INHS—Illinois Natural History Survey, Urbana CASC—California Academy of Sciences, San JGEC—J. G. Edwards Collection, San Jose, CA Francisco KMFC—K. M. Fender Collection, McMinnville, CISC—University of California, Berkeley OR CNCI—Canadian National Collections, Ottawa KSUC—Kansas State University, Manhattan CUIC—Cornell University, Ithaca, NY LACM—Los Angeles County Museum, CA CWOB—C. W. O'Brien Collection, Tallahassee, LSUC—Louisiana State University, Baton Rouge FL MCZC—Museum of Comparative Zoology, Har- DHKC—D. H. Kistner CoUection, Chico State vard University, Cambridge, MA College, CA MSUC—Michigan State University, East Lansing ELSC—E. L. Sleeper Collection, Long Beach, NCSM—North Carolina State University, Raleigh CA NYSM—^New York State Museum, Albany FMNH—^Field Museum of Natural History, Chi- OSEC—Oklahoma State University, Stillwater cago, IL OSUC—Ohio State University, Columbus OSUO—Oregon State University, Corvallis

' Abbreviations for U.S. and Canadian museums abridged from Arnett, R. H., Jr., and Samuelson, G. A., 1969, "Di- rectory of Coleóptera Collections of North America (Canada Through Panama)," Cushing-Malloy, Ann Arbor, MI, 123 pp. vm PMNH—Peabody Museum, Yale University, New SMSH—Stovall Collection, University of Okla- Haven, CT homa, Norman PSUC—Pennsylvania State Museum, University TAMU—Texas A. & M. University, College Park Station PURC—Purdue University, West Lafayette, IN UCDC—University of California, Davis RUIC—Rutgers University, New Brunswick, NJ UMMZ—University of Michigan, Ann Arbor SEMC—Snow Museum, University of Kansas, UMRM—University of Missouri, Columbia Lawrence USNM—U.S. National Museum of Natural His- SJSC—San Jose State College, CA tory, Washington, DC SLWC—S. L. Wood Collection, Provo, UT WSUC—Washington State University, Pullman

MUSEUMS IN FOREIGN COUNTRIES

BMNH—British Museum (Natural History), NHRS—Naturhistoriske Riksmuseet, Stockholm London NMPC—Narodni Museum, Prague, Czechoslo- BPBM—Bernice P. Bishop Museum, Honolulu vakia GUHC—Glasgow University, Hunterian College, SCUT—Spinola College, University of Turin, Scotland Italy HMOX—Hope Museum, Oxford, England SMTD—Staatliches Museum für Tierkunde, HNHM—Hungarian Natural History Museum, Dresden, East Germany Budapest UNAM—Universidad Nacional Autónoma, Mexi- IPZE—Institut Pflanzenschutzforschung Zweig- co Citv stelle, Eberswalde, East Germany UZMC—University Zoological Museum, Copen- IRSB—Institut Royal Sciences Belgique, Brus- hagen, Denmark sels UZMH—University Zoological Museum, Hel- MFNB—Museum für Naturkunde (Humboldt), sinki, Finland Berlin ZMAS—Zoological Museum, Academy of Sci- MGFT—Museum G. Frey, Tutzing, Munich, ences, Leningrad West Germany ZMPA—Zoological Museum, Pohsh Academy of MHNL—Museum d'Histoire Naturelle, Lyon, Sciences, Warsaw France ZMUL—Zoological Museum, University of Lund, MNHP—Museum National d'Histoire Naturelle, Sweden Paris ZMUM—Zoological Museum, University of MNSL—Museum of Natural Sciences, Leipzig, Moscow East Germany ZSBS—Zoologische Sammlung Bayerischen MZBS—Museum Zoología, Barcelona, Spain Staates, Munich, West Germany IX ACKNOWLEDGMENTS We are indebted lo many individuals who contributed to the planning and development of this ca- talog. We are especially grateful to the following specialists who helped to make it as complete and accurate as possible: Richard H. Foote, Systematic Entomology laboratory (SEI), Agricultural Research Service (ARS), for his suggestions, guidance, and encouragement; C. W. Sabrosky, SEI, for valuable counsel on nomenclatural problems; J. F. L awrence. Division of Entomology, Com- monwealth Scientific and Industrial Research Organization, Canberra, Australia, for his recommenda- tions on higher categories; and more than 50 coleopterists in Canada, the United States, and Mexico for voluntarily contributing information about their specialty groups. We thank the following members of the Communications and Data Services Division, SEMS: San- dra Strauss and Marianne Kingston for designing and writing the computer programs, and Margaret Seldin for developing the editing system.

J. M. Kingsolver, editor in chief Systematic Entomology I aboratory. Agricultural Research Service Washington, D.C.

Editorial Board

J. M. Campbell, Biosystematics Research Institute, Agriculture Canada, Ottawa, ON T. L. Erwin, Department of Entomology, Smithsonian Institution, Washington, DC H. F. Howden, Department of Biology, Carleton university, Ottawa, ON P. J. Spangler, Department of Entomology, Smithsonian Institution, Washington, DC T. J. Spilman, Systematic Entomology Laboratory, ARS, Washington, DC R. E. White, Systematic Entomology Laboratory, ARS, Washington, DC

CONTENTS Page Family Cerylonidae 1 Subfamily Euxestinae 1 Subfamily Anommatinae 1 Subfamily Murmidiinae - 2 Subfamily Ostomopsinae 3 Subfamily -- 3 Bibliography 5 Index 10 Family CERYLONIDAE Billberg, 1820 By J. F. Lawrence Cerylonides Billberg, 1820: 47 The Cerylonidae includes about 40 genera and over 300 species from all the major Zoogeo- graphie regions. The North American fauna consists of 18 species in 10 genera. This was first recognized as an independent clavicorn family by Crowson (1955), who included the cerylonines, murmidiines, and euxestines, all of which were treated as tribes of the heteromeran family Co- lydiidae by both Hetschko (1930) and Arnett (1968). Sen Gupta and Crowson (1973) added the genera Anommatus Wesmael, Abromas Reitter, and Ostomopsis Scott while transferring Eidor- eus Sharp {'^Eupsilobius Casey) to the Endomychidae. The classification used here is from Lawrence and Stephan (1975), which represents a modification of that proposed by Sen Gupta and Crowson (1973). The latter authors treat the Anommatinae and Ostomopsinae as tribes of the subfamilies Euxestinae and Murmidiinae, re- spectively. The Cerylonidae is believed to contain relatively primitive members of the cerylonid series of Clavicornia, a group which also includes the Sphaerosomatidae, Endomychidae, Coc- cinellidae, Corylophidae, Discolomidae, Lathridiidae, and Merophysiidae. The phylogenetic rela- tionships among these and other clavicorn families are being studied by R. A. Crowson and T. Sen Gupta. Because of their small size and cryptic habits, cerylonids are still not very well known sys- tematically or biologically. Although Sen Gupta and Crowson have revised the world fauna at the superspecific level, several genera and numerous species remain to be described from all regions, and modification of present concepts may still be required, especially for the poorly known neotropical fauna. Several North American species have been discovered only recently, and further collecting in certain microhabitats may reveal other undescribed forms. The larvae of MychoceruSy Botrodus, and Ostomopsis are still unknown, as are those of various exotic genera. Cerylonids are most often collected in leaf litter and under bark, but some have been taken in fungus fruiting bodies, in ant and mammal nests, and in animal debris (ant refuse, bat guano). Examination of gut contents has shown fungal hyphae and spores to be the most common food source. This manuscript was received November 1980.

Subfamily EUXESTINAE Grouvelle, 1908 Pachyochthesinae Reitter, 1922: 5, 53; Tachyoryctidiini Jeannel and Paulian, 1945: 56; Cy- cloxenini Jeannel and Paulian, 1945: 57 Genus HYPODACNE LeConte Hypodacne LeConte, 1875: 170. Type-species: Hypodacne punctata LeConte (monot.). Pachyochthes Reitter, 1897: 248 (synonymized by Nikitsky and Belov, 1979: 849). Type- species: Pachyochthes edithae Reitter (monot.). Euxestus, of authors, in part, not WoUaston. IMMATURE STAGES: Sen Gupta and Crowson, 1973: 381, figs. 218-224; Nikitsky and Belov, 1979: 852. REDESCRIPTION: Sen Gupta and Crowson, 1973: 380, figs. 50-51, 54-60. punctata LeConte, 1875: 171. Eastern United States; MI ON/ KS lA MO IN OH KY/ NY PA NJ MD DC VA/ TX/ AR MS TN SC NC FL. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Lawrence and Stephan, 1975: 139, fig. 20. ECOLOGY: Stephan, 1968. HOST: Camponotus galleries in oak, elm, beech.

Subfamily ANOMMATINAE Ganglbauer, 1899 Anommatini Ganglbauer, 1899: 890 ¿ COLEÓPTERA CATALOG

Genus ANOMMATUS Wesmael Anommatus Wesmael, 1835: 338. Type-species: Anommatus terrícola Wesmael (monot.) duodecimstriatus (Mueller). TAXONOMY: Dajoz, 1965. KEYS: Dajoz, 1965. duodecimstriatus (Mueller), 1821: 190 (Lyctus). Germany: Odenbach; IL OH/ NY/ Old World. obsoletas Stephens, 1830: 98 (Tribolium). London. terrícola Wesmael, 1835: 339, pl. 4 (part). Belgium: Chenee, near Liege. pusíllus Schaufuss, 1862: 49. Saxony. basalís Reitter, 1883: 197. Germany. IMMATURE STAGES: Dajoz, 1968: 979, figs. 11-24. REDESCRIPTION: Crowson, 1944: 290; Sen Gupta and Crowson, 1973: 392; Lawrence and Stephan, 1975- fig. 23. ECOLOGY: Dury, 1928; Cooper, 1962; Dajoz, 1968; Peck, 1972; Sen Gupta and Crowson, 1973. HOST: Litter, leaf mold, damp soil, grass roots, grass cuttings, under stones, tree holes. Subfam ly MURMIDIINAE Jacquelin du Val, 1858 Murmidiides Jacquelin du Val, 1858: 227; Mychocerini Ganglbauer, 1899: 907 Genus MURMIDIUS Leach Murmidius Leach, 1822: 41. Type-species: Murmídius ferrugíneus Leach (monot.) ovalís (Beck). Ceutocerus Germar, 1824: 85. Type-species: Ceutocerus advena Germar (monot.) ovalís (Beck). Ceuthocerus (error) Marseul, 1885: 182. TAXONOMY: Hinton, 1942. KEYS: Hinton, 1942. ovalis (Beck), 1817: 7, pl. 1, fig. 1 (Híster). Bavaria; AK/ ON/ CA/ KS IL/ NY MD DC/ LA FL/ Old World. ferrugíneus Leach, 1822: 41. England? (imported from China). advena Germar, 1824: 85 (Ceutocerus). Europe?. IMMATURE STAGES: Boeving and Craighead, 1931; Halstead, 1968; Sen Gupta and Crowson, 1973: figs. 225, 232-234. REDESCRIPTION: Crowson, 1944: ñg. 73; Sen Gupta and Crowson, 1973: 395, figs. 80-90; Lawrence and Stephan, 1975. ECOLOGY: Halstead, 1968: 13; Hinton, 1945: 184. HOST: Fruits, seeds, corn, rice, wheat, flour, hay, straw, gall-nuts, oak galls, ginger, pepper, dead leaves, cut grass. Genus Erichson Mychocerus Erichson, 1845: 292. Type-species: Mychocerus ferrugíneus Erichson (monot.). TAXONOMY: Lawrence and Stephan, 1975: 142. REDESCRIPTION: Sen Gupta and Crowson, 1973: 399, figs, 99-105. KEYS: Lawrence and Stephan, 1975: 142 (North America). arizonensis Lawrence and Stephan, 1975: 144, figs. 3, 5-6. AZ: Pima Co., Arivaca; AZ/ Mex. TYPE DEPOSITORY: MCZC. HOST: Under dry mesquite bark, under oak bark. depressus (LeConte), 1866: 376 (Murmidius). DC; ON/ OH/ PA MD DC/ TX/ TN SC FL. TYPE DEPOSITORY: MCZC. TAXONOMY: LeConte, 1869. REDESCRIPTION: Lawrence and Stephan, 1975: 145, fig. 22. HOST: Under fungus-infested bark of oak, beech, hickory. Genus BOTRODUS Casey Botrodus Casey, 1890: 319. Type-species: Botrodus estriatus Casey (monot.). REDESCRIPTION: Sen Gupta and Crowson, 1973: 396, figs. 91-98. estriatus Casey, 1890: 320. TX: Columbus; TX/ FL. TYPE DEPOSITORY: USNM. CERYLONIDAE 3 Subfamily OSTOMOPSINAE Sen Gupta and Crowson, 1973 Ostomopsini Sen Gupta and Crowson, 1973: 400 Genus OSTOMOPSIS Scott Ostomopsis Scott, 1922: 250, pi. 19, figs. 6, 11. Type-species: Ostomopsis solitaria Scott (orig. des.). TAXONOMY: Lawrence and Stephan, 1975: 146. REDESCRIPTION: Sen Gupta and Crowson, 1973: 400, figs. 43-49. neotropicalis Lawrence and Stephan, 1975: 146, figs. 4, 7, 24. Canal Zone: Barro Colorado IsL; FL/ Mex., C. Amer. TYPE DEPOSITORY: MCZC. HOST: Rotten palm wood, leaf litter.

Subfamily CERYLONINAE Billberg, 1820 Cerylonides Billberg, 1820: 47; Pleosomides Fauvel, 1891: 162; Lapethini Sharp, 1894: 445; Aculagnathidae Oke, 1932: 22; Dolosidae Dajoz, 1963: 96 Genus Aube Philothermus Aube, 1843: 93. Type-species: Philothermus montandoni Aube (orig. des.). IMMATURE STAGES: Sen Gupta and Crowson, 1973: 423, figs. 226-231 TAXONOMY: Heinze, 1944; Lawrence and Stephan, 1975: 150. REDESCRIPTION: Sen Gupta and Crowson, 1973: 423, figs. 125-132. KEYS: Lawrence and Stephan, 1975: 150 (North America). glabriculus LeConte, 1863: 69. PA; WI MI ON PQ/ lA MO IL IN OH KY/ NY PA NJ DE MD DC WV VA/ ME NH VT MA RI CT/TX OK/ AR LA MS AL TN GA SC NC FL. TYPE DEPOSITORY: MCZC. REDESCRIPTION: Lawrence and Stephan, 1975: 151, fig. 25. HOST: Under bark oí Acer, Carya, Fagus, Pinus, Quercus, Tsuga, Ulmus; under boards, in rotten logs, sawdust piles, malt traps, leaf litter, tree holes, forest debris. occidentalis Lawrence and Stephan, 1975: 152, figs. 8, 9, 26. CA: Tuolumne Co., Mather; CA. TYPE DEPOSITORY: MCZC. HOST: Coniferous forest litter. puberulus Schwarz, 1878: 361. FL; GA FL/ Mex., C. Amer. TYPE DEPOSITORY: USNM. REDESCRIPTION: Lawrence and Stephan, 1975: 153, fig. 27. HOST: Under bark of Pinus, swamp litter. Genus CERYLCAUTOMUS Sen Gupta and Crowson Cerylcautomus Sen Gupta and Crowson, 1973: 439. Type-species: Cerylcautomus floridensis Sen Gupta and Crowson (orig. des.). floridensis Sen Gupta and Crowson, 1973: 441, figs. 205-213. FL: Leon Co., Chaires; FL. TYPE DEPOSITORY: FMNH. HOST: Old sawdust pile. Genus LAPETHUS Casey Lapethus Casey, 1890: 317. Type-species: Lapethus discretas Casey (monot.). Lapecautomus Sen Gupta and Crowson, 1973: 409, figs. 171-178 (synonymized by Lawrence and Stephan, 1975: 156). Type-species: Lapecautomus dybasi Sen Gupta and Crowson (orig. des.). Lytopeplus Sharp, 1895: 494 (synonymized by Hinton, 1936). Type-species: Lytopeplus compactus Sharp (monot.). Brachylon Gorham, 1898: 256. Type-species: Brachylon breve Gorham (monot.). TAXONOMY: Hinton, 1936: 185; Lawrence and Stephan, 1975: 154. REDESCRIPTION: Sen Gupta and Crowson, 1973: 404, figs. 159-178. KEYS: Lawrence and Stephan: 155 (North America). 4 COLEÓPTERA CATALOG discretas Casey, 1890: 318. CA: Humboldt Co., near Humboldt Bay; WA/ CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Hatch, 1962: 248, pi. 52, ñg. 2; Lawrence and Stephan, 1975: 156, figs. 12, 28. HOST: Red rotten conifer logs. striatus (Sen Gupta and Crowson), 1973: 412, fig. 178 (Lapecautomus), TN: Sevier Co., Great Smoky Mts. N. P.; TN. The type was found below Ramsay Cascades, 3500'. TYPE DEPOSITORY: FMNH. HOST: Leaf litter under Rhododendron; mixed hardwood and Tsuga forest. Genus Latreille Cerylon Latreille, 1802: 228. Type-species: Lyctus histeroides Fabricius (subseq. monot.: Latreille, 1810: 431). Cerylus Agassiz, 1846: 75 (unjustified emendation). Type-species: Lyctus histeroides Fabricius (autom.). Aphardion Gozis, 1886: 11. Type-species: Lyctus histeroides Fabricius (orig. des.). IMMATURE STAGES: Sen Gupta and Crowson, 1973: 421, figs. 235-240. TAXONOMY: Heinze, 1944; Lawrence and Stephan, 1975: 157. REDESCRIPTION: Crowson, 1938: pi. 3, fig. 1; Sen Gupta and Crowson, 1973: 420, figs. 115-124. KEYS: Lawrence and Stephan, 1975: 157 (North America). californicum Casey, 1890: 316. CA: Lake Tahoe; OR/ CA. TYPE DEPOSITORY: USNM. REDESCRIPTION: Lawrence and Stephan, 1975: fig. 31. HOST: Coniferous forest litter. castaneum Say, 1827: 259. Eastern United States; BC/ AB/ WI MI ON PQ/ KS lA MO IL IN OH KY/ NY PA NJ MD DC WV VA/ ME NH VT MA CT/ TX/ TN SC NC. TYPE DEPOSITORY: MCZC (neotype). clypeale Casey, 1897: 636 (synonymized by Lawrence and Stephan, 1975: 158). KS. TYPE DEPOSITORY: USNM. sticticum Casey, 1897: 636 (synonymized by Lawrence and Stephan, 1975: 158). lA: Iowa City. TYPE DEPOSITORY: USNM. TAXONOMY: Lawrence and Stephan, 1975: 158 (neotype designation). REDESCRIPTION: Lawrence and Stephan, 1975: figs. 15, 17. HOST: Under bark of Acer, Fagus, Picea, hardwoods; in fruiting bodies of Pleurotus sp., Fomes fomentarius, Phellinus igniarius, Bjerkandera adusta. conditum Lawrence and Stephan, 1975: 159, figs. 13, 16. AZ: Pima Co., Santa Catalina Mts., 8000'; AZ NM. TYPE DEPOSITORY: MCZC. HOST: Under bark of fungus-infested conifer logs. distans Lawrence and Stephan, 1975: 161, figs. 14, 29. AZ: Cochise Co., Chiricahua Mts., 8500'; AZ. TYPE DEPOSITORY: MCZC. HOST: Under bark of fungus-infested conifer logs. unicolor (Ziegler), 1845: 270 (Latridius). Carolina; BC WA OR ID/ MT ND/ MN WI MI ON PQ/ CA/ NE KS lA MO IL IN OH KY/ NY PA NJ MD DC WV VA/ ME NH VT MA CT/ TX/ AR LA MS TN GA SC NC FL. TYPE DEPOSITORY: MCZC. simplex LeConte, 1857: 39 (synonymized by Lawrence and Stephan, 1975: 162). CA: San Jose. TYPE DEPOSITORY: MCZC. angustulum LeConte, 1863: 69 (synonymized by Lawrence and Stephan, 1975: 162). GA. TYPE DEPOSITORY: MCZC. sylvaticum Casey, 1897: 635 (synonymized by Lawrence and Stephan, 1975: 162). ID: Coeur d'Alene. TYPE DEPOSITORY: USNM. REDESCRIPTION: Hatch, 1962: 247, pi. 51, fig. 7; Lawrence and Stephan, 1975: 162, figs. 18-19, 30. HOST: Under bark of Carya, Fagus, Liriodendron, Picea, Pinus, Platanus, Populus, Liquidambar, Salix, Tsuga-, in rotten logs, leaf mold, sap, ears of corn, malt traps; fruiting bodies oí various fungi, including Phellinus gilvus, Bjerkandera adusta. BIBLIOGRAPHY Agassiz, L. 1846 Nomenclatoris Zoologici. Index universalis, continens nomina systematica clas- sium, ordinum, familiarum et generum animalium omnium, tam viventium quamfossilium, secundum ordinem alphabeticum unicum disposita, adjectis ho- monymiis plantarum, nee non variis adnotationibus et emendationibus. Jent and Gassmann, Soloduri, viii and 393 pp. Arnett, R. H. 1968 The beetles of the United States. (A manual for identification), American Entomo- logical Institute, Ann Arbor, Michigan, xii and 1112 pp., illus. Aube, C. 1843 Notes sur une nouvelle espèce de Coleóptera tetramere, qui devra servir de base a une coupe générique nouvelle. Annales de la Société Entomologique de France, ser. 2, vol. 1, pp. 93-95, illus. Beck, L. von 1817 Beitraege zur baierischen Insektenfaune, oder Beschreibung und Abbildung neuentdeckter Kaefer mit angehaengtem Namensverzeichnisse der Eleuteraten des Landgerichtsbezirks Zusmeshausen. Wolff, Augsburg, 45 pp., illus. Billberg, J. G. 1820 Enumeratio insectorum in museo. Gadelianis, Stockholm, 138 pp. Boeving, A. G. and F. C. Craighead 1931 An illustrated synopsis of the principal larval forms of the order Coleóptera. Ento- mológica Americana, n. ser., voL 11, pp. 1-351, illus. Casey, T. 1890 Coleopterological notices. II. Annals of the New York Academy of Sciences, vol 5 pp. 307-504, illus. Casey, T. 1897 Coleopterological notices. VII. Annals of the New York Academy of Sciences, vol 9, pp. 285-684. Cooper, K. W. 1962 A remarkable anophthalmic immigrant to North America: Anommatus duodecim- striatus Mueller (Coleóptera: Colydiidae). Entomological News, vol. 73, pp. 187- 190. Crowson, R. A. 1938 The metendosternite in Coleóptera: a comparative study. The Transactions of the Royal Entomological Society of London, vol. 87, pp. 397-415, illus. Crowson, R. A. 1944 Further studies on the met-endosternite in Coleóptera. The Transactions of the Royal Entomological Society of London, vol. 94, pp. 273-310. Crowson, R. A. 1955 The natural classification of the families of Coleóptera. Nathaniel Lloyd & Co., London, 187 pp., illus. Dajoz, R. 1963 Dolosas leleupi n. g., n. sp. et Dolosus basilewskyi n. sp., types d'une famille nou- velle de (Coléoptères). Revue de Zoologie et de Botanique Africaines, vol. 67, pp. 91-96. Dajoz, R. 1965 Revision des Colydiidae anophthalmes de la faune palearctique (Insectes, Coléop- tères). Revue d'Ecologie et de Biologie du Sol, vol. 2, pp. 239-269. 6 COLEÓPTERA CATALOG

Dajoz, R. 1968 Revision des Colydiidae anophthalmes de la faune palearctique IV. Etudes sur les genres Anommatus et Langelandia. Annales de la Société Entomologique de France, n. ser., vol. 4, pp. 975-988. Dury, C. 1928 Blaps mucronata Latr. in Cincinnati, O, Bulletin of the Brooklyn Entomological Society, vol. 23, p. 180. Erichson, W. F. 1845 Naturgeschichte der Insecten Deutschlands. Nicolai, Berlin, Abtheilung 1. Coleóp- tera. Band 3. Lieferung 2, pp. 161-320. Fauvel, A. 1891 Les Coléoptères de la Nouvelle-Calédonie et dépendances, avec descriptions, notes et synonymies nouvelles (Suite). Revue d'Entomologie, vol. 10, pp. 148-182. Ganglbauer, L. 1899 Die Kaefer von Mitteleuropa. Die Kaefer der oestereichisch-ungarischen Monar- chie, Deutschlands, der Schweiz, sowie der franzoesischen und italienischen Al- pengebietes. Carl Gerold's Sohn, Wien, Dritter Band, erste Haelfte. Familien- reihe Staphylinoidae. 2 Theil, 408 pp., illus. Dritter Band, zweite Haelfte. Fami- lienreihe Clavicornia, pp. iii and 409-1046, illus. Germar, E. F. 1824 Insectorum species novae aut minus cognitae, descriptionibus illustratae. L Hendel, Halae, xxiv and 624 pp., illus. Gorham, H. S. 1898 Erotylidae, Endomychidae and Coccinellidae (part). In Biología Centrali-Ameri- cana, Insecta. Coleóptera. F. Godman and O. Salvin, eds. R. H. Porter, London, vol. 7, pp. 241-256. Gozis, M. des 1886 Recherche de Tespece typique de quelques anciens genres. Rectifications synonymi- ques et notes diverses. Herbin, Montlucon, France, 36 pp. Grouvelle, A. 1908 Coléoptères de la region Indienne. Rhysodidae, Trogositidae, Nitidulidae, Colydii- dae, Cucujidae. Annales de la Société Entomologique de France, vol. 77, pp. 315- 495, illus. Halstead, D. G. H. 1968 Observations on the biology of Murmidius ovalis (Beck) (Coleóptera: Cerylonidae). Journal of Stored Products Research, vol. 4, pp. 13-21. Hatch, M. H. 1962 The beetles of the Pacific Northwest. Part III: Pselaphidae and Diversicornia L University of Washington Press, Seattle, Washington, ix and 503 pp., illus. Heinze, E. 1944 Beitraege zur Kenntniss der Tribus Cerylini und Metacerylini (nov.). (Coleóptera: Colydiidae). Arbeiten ueber morphologische und taxonomische Entomologie aus Berlin-Dahlem, vol. 11, pp. 19-32. Hetschko, A. 1930 Colydiidae. In Coleopterorum catalogus auspiciis et auxilio W. Junk editus a S. Schenkling. Junk, Berlin, pt. 107, (vol. 15), 124 pp. Hinton, H. E. 1936 Notes on some American Colydiidae. (Coleóptera). Entomological News, vol. 47, pp. 185-189. CERYLONIDAE 7

Hinton, H. E. 1942 A synopsis of the old world species of Murmidius Leach (Coleóptera, Colydiidae). The Proceedings of the Royal Entomological Society of London. Series B. Taxon- omy, vol. 11, pp. 39-45. Hinten, H. E. 1945 A monograph of the beetles associated with stored products. British Museum (Nat- ural History), London, vol. 1, viii and 443 pp. Jacquelin du Val, P. 1858 Manuel entomologique. Genera des Coléoptères d'Europe, comprenant leur classifi- cation en familles naturelles, la description de tous les genres, des tableaux dich- tomiques destines a faciliter Tetude, le catalogue de toutes les espèces, de nom- breux dessins au trait de caracteres par Jacquelin du Val, et près de treize cents insectes représentant un ou plusieurs types. DeyroUe, Paris, vol. 2 (part), pp. 113-285. Jeannel, R. and R. Paulian 1945 Mission scientifique de TOmo. Faune des terriers des rats-taupes IV. Coléoptères. Mémoires de la Musée National d'Histoire Naturelle, n. ser., vol. 19, no. 1, pp. 51-147. Latreille, P. 1802 Histoire naturelle, genérale et particulière des Crustacés et des Insectes. Dufart, Paris, Tome 3, 468 pp. Latreille, P. 1810 Considerations generales sur Tordre naturel des animaux composant les classes des Crustacés, des Arachnides, et des Insectes. Schoell, Paris, 444 pp. Lawrence, J. F. and K. Stephan 1975 The North American Cerylonidae (Coleóptera: Clavicornia). Psyche, vol. 82, no. 2, pp. 131-166. Leach, W. E. 1822 Characters of a new genus of coleopterous of the family Byrrhidae. The Transactions of the Linnean Society of London, vol. 13, p. 41. LeConte, J. L. (as J. T.) 1857 Report upon insects collected on the survey. War Department, Washington, D. C, 72 pp., illus. The date for the species described in this work was taken from a preprint fírst issued in 1857. It was published again in 1860: In Reports of explorations and surveys to ascertain the most practica- ble and economical route for a railroad from the Mississippi River to the Pacific Ocean, vol. 12, bk. 2, pt. 3, no. 1, Zoological Report. LeConte, J. L. 1863 New species of North American Coleóptera. Prepared for the Smithsonian Institu- tion. Part I (part). Smithsonian Miscellaneous Collections, no. 167, pp. 1-86. LeConte, J. L. 1866 Additions to the coleopterous fauna of the United States. No. 1. Proceedings of the Academy of Natural Sciences of Philadelphia, pp. 361-394. LeConte, J. L. 1869 Synonymical notes on Coleóptera of the United States, with descriptions of new species, from the MSS. of the late Dr. C. Zimmermann. Transactions of the American Entomological Society, vol. 2, pp. 243-259. LeConte, J. L. 1875 Descriptions of new Coleóptera of the United States with notes on geographical distribution. Transactions of the American Entomological Society, vol. 5, pp. 169- 176. 8 COLEÓPTERA CATALOG

Marseul, S. A. de 1885 Catalogue des Coléoptères de Tancien monde: Anisotomides, Clambides, Corylo- phides, Ptiliides, Scaphidiides, Histerides, Phalacrides, Nitidulides, Trogositides, Colydiides. L'Abeille, Journal d'Entomologie, vol. 23, pp. 169-192. Mueller, P. W. J. 1821 Neue Insekten, beschreiben von Ph. W. J. Mueller. Magazin der Entomologie, vol. 4, pp. 184-230. Nikitsky, N. B. and V. V. Belov 1979 Novii i maloizvestnii Clavicornia (Coleóptera) iz Talisha. Zoologicheskii Zhurnal, vol. 58, no. 6, pp. 849-854. Oke, C. 1932 Aculagnathidae, a new family of Coleóptera. Proceedings of the Royal Society of Victoria, vol. 44, pp. 22-24. Peck, S. B. 1972 The eyeless European soil colydiid Anommatus duodecimstriatus, in North Amer- ica (Coleóptera: Colydiidae). The Coleopterists Bulletin, vol. 26, no. 1, pp. 19-20. Reitter, E. 1883 Ueber die deutschen Anommatus -Arten. Wiener Entomologische Zeitung, vol. 2, pp. 195-197. Reitter, E. 1922 Bestimmungs-Tabellen der europaeischen Coleopteren. VI. Heft. Enthaltend die Familien: Colydiidae, Rhysodidae, Ostomidae. E. Reitter, Troppau, 73 pp. Say, T. 1827 Descriptions of new species of coleopterous insects inhabiting the United States (part). Journal of the Academy of Natural Sciences of Philadelphia, vol. 5, no. 2, pp. 237-284, 293-304. Schaufuss, L. W. 1862 (Untitled note). Sitzungs-Berichte der naturwissenschaftlichen Gesellschaft Isis zu Dresden (1861), pp. 47-51. Schwarz, E. A. 1878 The Coleóptera of Florida. Proceedings of the American Philosophical Society, vol. 17, pp. 353-472. Descriptions of new species by E. A. Schwarz, pp. 354-372. Additional descriptions of new species by J. L. LeConte, pp. 373-434. A list of species by E. A. Schwarz, pp. 434-472. Scott, H. 1922 Reports of the Percy Sladen Trust Expedition to the Indian Ocean in 1905. Vol. 7, No. 4; Coleóptera: Scydmaenidae, Scaphidiidae, Phalacridae, Cucujidae (supple- ment), Lathridiidae, Mycetophagidae (including Propalticus), Bostrychidae, Lyc- tidae. The Transactions of the Linnean Society of London, ser. 2, Zoology, vol. 18, pp. 195-260, illus. Sen Gupta, T. and R. A. Crowson 1973 A review of the classification of Cerylonidae (Coleóptera, Clavicornia). Transac- tions of the Royal Entomological Society of London, vol. 124, pp. 365-446, illus. Sharp, D. 1894 Fam. Colydiidae (part). In Biología Centrali-Americana, Insecta. Coleóptera. F. Godman and O. Salvin, eds. R. H. Porter, London, vol. 2, pt. 1, pp. 443-488. Sharp, D. 1895 Fam. Colydiidae (part). In Biología Centrali-Americana, Insecta. Coleóptera. F. Godman and O. Salvin, eds. R. H. Porter, London, vol. 2, pt. 1, pp. 489-496. Stephan, K. (as Stephens) 1968 Notes on additional distribution and ecology of Euxestus punctatus LeC. (Coleóp- tera: Colydiidae). The Coleopterists' Bulletin, vol. 22, no. 1, p. 19. 9

Stephens, J. F. 1830 Illustrations of British entomology; or, a synopsis of indigenous insects: containing their generic and specific distinctions; with an account of their metamorphoses, times of appearance, localities, food, and economy, as far as practicable. Mandi- bulata. Baldwin and Cradock, London, vol. Ill, 374 pp., illus. Wesmael, C. 1835 Nouveau genre d'insectes Coléoptères, de la famille des Xylophages, tribu des Bos- trichins. Bulletins de TAcademie Royale des Sciences et Belles-lettres de Brux- elles, vol. 2, pp. 338-340, illus. Ziegler, D, 1845 Descriptions of new North American Coleóptera. Proceedings of the Academy of Natural Sciences of Philadelphia, vol. 2, pp. 266-272. INDEX

Names are indexed as follows: depressus (LeConte), Mychocerus 2 CAPITALS: All names for taxa above the generic discretus Casey, Lapethus.... 4 level; distans Lawrence and Stephan, Cerylon 4 Boldface: Valid generic and subgeneric names; duodecimstriatus (Mueller), Anommatus 2 Ronian: Valid specific and subspecific names; estriatus Casey, Botrodus 2 Italic: All invalid names such as synonyms, nomina EUXESTINAE ...Z.'.".'.* 1 nuda, and extra-limital taxa even though valid. Euxestus 1 Parentheses around an author's name indicate that ferrugineus Leach, Murmidius 2 the specific name has been transferred from its original floridensis Sen Gupta and Crowson, Cerylcautomus... 3 genus. The generic name following the author's name in- glabriculus LeConte, Philothermus 3 dicates the present placement of the species. Synonyms Hypodacne LeConte i of species-group names are listed with the original spell- Lapecautomus Sen Gupta and Crowson 3 ing. Lapethus Casey 3 Lytopeplus Sharp 3 advena (Germar), Murmidius 2 MURMIDIINAE '.''ZZZ 2 angustulum LeConte, Cerylon 4 Murmidius Leach 2 ANOMMATINAE 1 Mychocerus Erichson 2 Anommatus Wesmael 2 neotropicalis Lawrence and Stephan, Ostomopsis 3 Aphardion Gozis 4 obsoletas (Stephens), Anommatus 2 arizonensis Lawrence and Stephan, Mychocerus 2 occidentalis Lawrence and Stephan, Philothermus 3 basalis Reitter, Anommatus 2 OSTOMOPSINAE 3 Botrodus Casey 2 Ostomopsis Scott 3 Brachylon Gorham 3 ovalis (Beck), Murmidius 2 californicum Casey, Cerylon 4 Pachyochthes Reitter 1 castaneum Say, Cerylon 4 Philothermus Aube 3 Cerylcautomus Sen Gupta and Crowson 3 puberulus Schwarz, Philothermus 3 Cerylon Latreille 4 punctata LeConte, Hypodacne 1 CERYLONINAE 3 pusillus Schaufuss, Anommatus 2 Cerylus Agassiz 4 simplex LeConte, Cerylon 4 Ceuthocerus 2 sticticum Casey, Cerylon 4 Ceutocerus Germar 2 striatus (Sen Gupta and Crowson), Lapethus 4 clypeale Casey, Cerylon 4 sylvaticum Casey, Cerylon 4 conditum Lawrence and Stephan, Cerylon 4 terricola Wesmael, Anommatus 2 unicolor (Ziegler), Cerylon 4 *^^^lTr.^P<^^

^á î?u%'-^^^

AB Alberta MB Manitoba ON Ontario AK Alaska MD Maryland OR Oregon AL Alabama ME Maine PA Pennsylvania AR Arkansas Ml Michigan PE Prince Edward Island AZ Arizona MN Minnesota PM St. Pierre-Miquelon BC British Columbia MO Missouri RQ Quebec CA California MS Mississippi Rl Rhode Island CO Colorado MT Montana SO South Carolina CT Connecticut NB New Brunswick SD South Dakota DC District of Columbia NC North Carolina SK Saskatchewan DE Delaware ND North Dakota TN Tennessee FL Florida NE Nebraska TX Texas GA Georgia NF Newfoundland UT Utah GL Greenland NH New Hampshire VA Virginia !A Iowa NJ New Jersey VT Vermont ID Idaho NM New Mexico WA Washington IL Illinois NS Nova Scotia Wl Wisconsin IN Indiana NT Northwest Territories WV West Virginia KS Kansas NV Nevada WY Wyoming KY Kentucky NY New York YT Yukon Territory LA Louisiana OH Ohio MA Massachusetts OK Oklahoma *US GOVERNMENT PRINTING OFFICE:1982-381-2