WTU Herbarium Specimen Label Data
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Outline of Angiosperm Phylogeny
Outline of angiosperm phylogeny: orders, families, and representative genera with emphasis on Oregon native plants Priscilla Spears December 2013 The following listing gives an introduction to the phylogenetic classification of the flowering plants that has emerged in recent decades, and which is based on nucleic acid sequences as well as morphological and developmental data. This listing emphasizes temperate families of the Northern Hemisphere and is meant as an overview with examples of Oregon native plants. It includes many exotic genera that are grown in Oregon as ornamentals plus other plants of interest worldwide. The genera that are Oregon natives are printed in a blue font. Genera that are exotics are shown in black, however genera in blue may also contain non-native species. Names separated by a slash are alternatives or else the nomenclature is in flux. When several genera have the same common name, the names are separated by commas. The order of the family names is from the linear listing of families in the APG III report. For further information, see the references on the last page. Basal Angiosperms (ANITA grade) Amborellales Amborellaceae, sole family, the earliest branch of flowering plants, a shrub native to New Caledonia – Amborella Nymphaeales Hydatellaceae – aquatics from Australasia, previously classified as a grass Cabombaceae (water shield – Brasenia, fanwort – Cabomba) Nymphaeaceae (water lilies – Nymphaea; pond lilies – Nuphar) Austrobaileyales Schisandraceae (wild sarsaparilla, star vine – Schisandra; Japanese -
UNIVERSITY of CALIFORNIA SANTA CRUZ Refining Species
UNIVERSITY OF CALIFORNIA SANTA CRUZ Refining species distribution models for coevolving plant and insects using species interactions and phylogeography A thesis submitted in partial satisfaction of the requirements for the degree of MASTER OF ARTS in ECOLOGY AND EVOLUTIONARY BIOLOGY by Christopher Schwind December 2016 The Thesis of Christopher Schwind is approved: Professor John N. Thompson Professor Kristy Kroeker Professor Erika Zavaleta ________________________________ Tyrus Miller Vice Provost and Dean of Graduate Studies Table of Contents Abstract........................................................................................................................iv Acknowledgements......................................................................................................v Introduction..................................................................................................................1 Methods........................................................................................................................5 Results..........................................................................................................................12 Discussion....................................................................................................................21 Supplemental Information...........................................................................................28 References....................................................................................................................30 -
CDLT Mountain Home & USFS-Boundry Butte Plant List
CDLT Mountain Home Preserve- Boundary Butte Plant list CDLT Mountain Home & USFS-Boundry Butte Plant list Type Scientific Name Common Name Fern Pteridium aquilinum bracken fern Forb Achillea millefolium common yarrow Forb Agoseris heterophylla annual agoseris Forb Anemone oregana Oregon anemone Forb Antennaria racemosa raceme pussytoes Forb Boechera pauciflorus rockcress (Formerly Arabis) Forb Arnica cordifolia heart-leaf arnica Forb Balsamorhiza sagittata arrowleaf balsamroot Forb Brickellia oblongifolia Mojave brickellbush Forb Cacaliopsis nardosmia silvercrown (Formerly Luina) Forb Calochortus lyallii Lyall's mariposa lily Forb Camassia quamash common camas Forb Castilleja miniata scarlet Indian paintbrush Forb Claytonia lanceolata springbeauty Forb Collinsia parviflora small-flowered blue-eyed mary Forb Commandra umbellata bastard toadflax Forb Delphinium viridescens Wenatchee larkspur Forb Erythronium grandiflorum glacier lily Forb Erysimum species wallflower Forb Fragaria virginiana Virginia strawberry Forb Fritillaria affinis checker lily, chocolate lily Forb Fritillaria pudica yellow bells Forb Galium sp. bedstraw Forb Heuchera cylindrica roundleaf alumroot Forb Hydrophyllum capitatum ballhead waterleaf Forb Lathyrus pauciflorus few-flowered pea Forb Lithophragma parviflorum small-flowered woodland-star Forb Lithophragma glabrum bulbous woodland-star Forb Lithophragma tenellum slender woodland-star Forb Lomatium nudicaule barestem biscuitroot Forb Lomatium triternatum nineleaf biscuitroot Forb Lonicera ciliosa orange honeysuckle -
Wildflowers of Twin Pillars North Trail, Bingham Springs, Trail Station
Wildflowers of Twin Pillars North Trail, Bingham Springs, Trail Station Meadows & upper Desolation Canyon Mill Creek Wilderness & Ochoco National Forest Data from personal observations & Data Provided by the Oregon Flora Project & Consortium of Pacific Northwest Herbaria Taxa with a question mark have been found within 10 miles of this site but not yet documented onsite. Last updated May 28, 2018 Common Name Scientific Name Family ____ Black Elderberry Sambucus racemosa v. melanocarpa Adoxaceae ____ Bulbil Onion Allium geyeri v. tenerum Amaryllidaceae ____ Tolmie's Onion Allium tolmiei v. tolmiei Amaryllidaceae ____ Gray’s Lovage Ligusticum grayi Apiaceae ____ Slender-fruited Lomatium Lomatium leptocarpum Apiaceae ____ Gray's Desert Parsley Lomatium papilioniferum ? Apiaceae ____ Common Sweet Cicely Osmorhiza berteroi Apiaceae ____ Western Sweet Cicely Osmorhiza occidentalis Apiaceae ____ Gairdner’s Yampah Perideridia gairdneri Apiaceae ____ Sierra Snakeroot Sanicula graveolens Apiaceae ____ Mountain Dogbane Apocynum androsaemifolium Apocynaceae ____ False Solomon Seal Maianthemum racemosum ssp. amplexicauleAsparagaceae ____ Star-flowered False Solomon SealMaianthemum stellatum Asparagaceae ____ Hyacinth Cluster-lily Triteleia hyacinthina ? Asparagaceae ____ Yarrow Achillea millefolium Asteraceae ____ Pale Agoseris Agoseris glauca v. glauca Asteraceae ____ Sagebrush Agoseris Agoseris parviflora Asteraceae ____ Pearly Everlasting Anaphalis margaritacea Asteraceae ____ Pussytoes Antenaria sp. Asteraceae ____ Woodrush Pussytoes Antennaria -
Slide Show Coevolution of Prodoxid Moths
One of the Classic Examples of Coevolution Prodoxid Moths and Their Host Plants Yucca Greya moths moths G. variabilis Some unknown G. subalbaancestor G. enchrysa G. obscura G. mitellae G. politella G. piperella Moderately Highly Antagonist Commensal/ Inefficient Efficient Efficient Obligate Antagonist Mutualist Mutualist Mutualist, mutualist (What we knew in 1979) Sometimes Exclusive Thompson, Pellmyr, Segraves, Althoff, Brown,… What We Now Know: Diversification of Traits and Ecological Outcomes Prodoxid Moths and Their Host Plants Yucca Greya moths moths G. variabilis G. subalba G. enchrysa G. obscura G. mitellae G. politella T. maculata G. piperella Antagonist Commensal/ Inefficient Moderately Highly Obligate Antagonist Mutualist Efficient Efficient mutualist Mutualist Mutualist, Sometimes Exclusive Thompson, Pellmyr, Harrison, Brown, Segraves, Althoff, Cunningham, Nuismer, Merg, Cuautle, Rich, Laine, Schwind, Friberg, Raguso,… Diversification of Derived Taxa in Drier Habitats Basal Prodoxidae Basal Greya Derived Greya Yucca moths Wahlberg et al. 2013 for Prodoxidae Pellmyr et al. (various) for Yucca moths Thompson et al. and Pellmyr et al (various) for Greya Pollination Mutualisms Evolved More than Once In Prodoxid Moths Pollinators 18 5 Monocot-feeders 5-6+ Eudicot-feeders Thompson 2014 in Grant and Grant, eds., In Search of the Causes of Evolution, Princeton Univ. Press These Mutualisms Involve Two Plant Families Agavaceae Agavaceae Saxifragaceae Thompson 2014 in Grant and Grant, eds., In Search of the Causes of Evolution, Princeton Univ. Press The Moths Ensure Developing Seeds for Their Offspring: Actively in Yucca Moths Apiaceae Agavaceae Passively in Greya moths: Greya politella on Woodland Stars (Lithophragma) Photos: John N Thompson Lithophragma (Woodland star) Traits Coevolved Unique traits in With Greya Moths Lithophragma parviflorum Variable stigma & style and height, shape, etc. -
Generalized Olfactory Detection of Floral Volatiles in the Highly Specialized Greya-Lithophragma Nursery Pollination System
Arthropod-Plant Interactions https://doi.org/10.1007/s11829-021-09809-5 ORIGINAL PAPER Generalized olfactory detection of foral volatiles in the highly specialized Greya-Lithophragma nursery pollination system Florian P. Schiestl1 · Erika A. Wallin2 · John J. Beck3 · Magne Friberg4 · John N. Thompson5 Received: 20 October 2020 / Accepted: 2 February 2021 © The Author(s) 2021 Abstract Volatiles are of key importance for host-plant recognition in insects. In the pollination system of Lithophragma fowers and Greya moths, moths are highly specialized on Lithophragma, in which they oviposit and thereby pollinate the fowers. Floral volatiles in Lithophragma are highly variable between species and populations, and moths prefer to oviposit into Litho- phragma fowers from populations of the local host species. Here we used gas chromatography coupled with electroanten- nographic detection (GC-EAD) to test whether Greya moths detect specifc key volatiles or respond broadly to many volatiles of Lithophragma fowers. We also addressed whether olfactory detection in Greya moths varies across populations, consistent with a co-evolutionary scenario. We analyzed fower volatile samples from three diferent species and fve populations of Lithophragma occurring across a 1400 km range in the Western USA, and their sympatric female Greya politella moths. We showed that Greya politella detect a broad range of Lithophragma volatiles, with a total of 23 compounds being EAD active. We chemically identifed 15 of these, including the chiral 6, 10, 14-trimethylpentadecan-2-one (hexahydrofarnesyl acetone), which was not previously detected in Lithophragma. All investigated Lithophragma species produced the (6R, 10R)-enantiomer of this compound. We showed that Greya moths detected not only volatiles of their local Lithophragma plants, but also those from allopatric populations/species that they not encounter in local populations. -
Common Native Forbs of the Northern Great Basin Important for Greater Sage-Grouse Tara Luna • Mark R
United States Department of Agriculture Common Native Forbs of the Northern Great Basin Important for Greater Sage-Grouse Tara Luna • Mark R. Mousseaux • R. Kasten Dumroese Forest Rocky Mountain General Technical Report November Service Research Station RMRS-GTR-387 2018 Luna, T.; Mousseaux, M.R.; Dumroese, R.K. 2018. Common native forbs of the northern Great Basin important for Greater Sage-grouse. Gen. Tech. Rep. RMRS-GTR-387. Fort Collins, CO: U.S. Department of Agriculture, Forest Service, Rocky Mountain Research Station; Portland, OR: U.S. Department of the Interior, Bureau of Land Management, Oregon–Washington Region. 76p. Abstract: is eld guide is a tool for the identication of 119 common forbs found in the sagebrush rangelands and grasslands of the northern Great Basin. ese forbs are important because they are either browsed directly by Greater Sage-grouse or support invertebrates that are also consumed by the birds. Species are arranged alphabetically by genus and species within families. Each species has a botanical description and one or more color photographs to assist the user. Most descriptions mention the importance of the plant and how it is used by Greater Sage-grouse. A glossary and indices with common and scientic names are provided to facilitate use of the guide. is guide is not intended to be either an inclusive list of species found in the northern Great Basin or a list of species used by Greater Sage-grouse; some other important genera are presented in an appendix. Keywords: diet, forbs, Great Basin, Greater Sage-grouse, identication guide Cover photos: Upper le: Balsamorhiza sagittata, R. -
Annotated Checklist of the Vascular Plant Flora of Grand Canyon-Parashant National Monument Phase II Report
Annotated Checklist of the Vascular Plant Flora of Grand Canyon-Parashant National Monument Phase II Report By Dr. Terri Hildebrand Southern Utah University, Cedar City, UT and Dr. Walter Fertig Moenave Botanical Consulting, Kanab, UT Colorado Plateau Cooperative Ecosystems Studies Unit Agreement # H1200-09-0005 1 May 2012 Prepared for Grand Canyon-Parashant National Monument Southern Utah University National Park Service Mojave Network TABLE OF CONTENTS Page # Introduction . 4 Study Area . 6 History and Setting . 6 Geology and Associated Ecoregions . 6 Soils and Climate . 7 Vegetation . 10 Previous Botanical Studies . 11 Methods . 17 Results . 21 Discussion . 28 Conclusions . 32 Acknowledgments . 33 Literature Cited . 34 Figures Figure 1. Location of Grand Canyon-Parashant National Monument in northern Arizona . 5 Figure 2. Ecoregions and 2010-2011 collection sites in Grand Canyon-Parashant National Monument in northern Arizona . 8 Figure 3. Soil types and 2010-2011 collection sites in Grand Canyon-Parashant National Monument in northern Arizona . 9 Figure 4. Increase in the number of plant taxa confirmed as present in Grand Canyon- Parashant National Monument by decade, 1900-2011 . 13 Figure 5. Southern Utah University students enrolled in the 2010 Plant Anatomy and Diversity course that collected during the 30 August 2010 experiential learning event . 18 Figure 6. 2010-2011 collection sites and transportation routes in Grand Canyon-Parashant National Monument in northern Arizona . 22 2 TABLE OF CONTENTS Page # Tables Table 1. Chronology of plant-collecting efforts at Grand Canyon-Parashant National Monument . 14 Table 2. Data fields in the annotated checklist of the flora of Grand Canyon-Parashant National Monument (Appendices A, B, C, and D) . -
Statement of Work
Validation and Verification of Rare and Mislabeled Plant Specimens at Whiskeytown National Recreation Area: Final Report Windy Bunn, Biological Science Technician June 2006 Introduction Whiskeytown National Recreation Area’s vascular plant list is based largely on an inventory conducted by David Biek (a California Native Plant Society volunteer) from 1985 to 1987. Biek’s flora (1988) and associated voucher specimens contain the most comprehensive information on the park’s flora gathered to date. However, findings from recent vegetation related projects in the park (Smith et al 2003, Mathiasen 2005, Bunn 2005, Bunn et al 2005, Bradley et al 2006) have lead to a growing uncertainty in the accuracy of the park’s plant list. Of particular concern are the findings of Bunn (2005) that illustrate that 13 out of 26 CNPS listed rare plants on the park’s list are either misidentifications or range extensions of the species. Due to this growing uncertainty and its potential to compromise the park’s NPS Biological Inventory goals, Whiskeytown staff drafted a proposal to begin validating the accuracy of the park’s vascular plant list. During the 2004 certification of the park’s plant list in NPSpecies, the existence of a voucher specimen in the park’s herbarium or the Shasta College Herbarium was a primary factor in determining a species “Park Status” rating. Therefore, the first priority of the proposal was to obtain some measure of the reliability of these voucher specimens as evidence for a species presence in the park. Thanks to funding provided by the Klamath Network Inventory and Monitoring Program, voucher specimens from the Whiskeytown and Shasta College Herbaria were validated by experts at the Jepson Herbarium. -
White River National Forest Rare Plant Survey 2006
White River National Forest Rare Plant Survey 2006 Prepared for White River National Forest By Peggy Lyon, Janis Huggins and Georgia Doyle Colorado Natural Heritage Program 254 General Services Building Colorado State University Fort Collins CO 80523 Acknowledgments This project would not have been possible without the help of many people in the White River National Forest: Kevin Atchley for first requesting this survey and working to get a contract in place; Beth Brenneman and Keith Giezentanner who took over for Kevin when he left the area; Joe Doerr and Philip Nyland, for sharing their experience and caring about the rare plants on the forest. Also, we depend on the support of the staff of CNHP in Fort Collins, especially Dave Anderson, Jill Handwerk, Susan Spackman, and Amy Lavender. Thanks also to Tim Hogan at the CU Herbarium for assisting with identification of problem plants; Dee Malone for her expertise and transportation to Taylor Pass; and John Chapman, FS Volunteer, for his time and transportation to Warren Lakes. Colorado Natural Areas Program, under the energetic direction of Brian Kurzel, and members of the Colorado Native Plant Society assisted with a survey of the Hoosier Ridge State Natural Area. Finally, thanks to our spouses, George and Rick, for putting up with our absences over the summer, and our faithful companions Molly and Misia, who protected us from Ptarmigan and other scary things. 2 Executive Summary The White River National Forest contracted with Colorado State University in 2005 to have the Colorado Natural Heritage Program (CNHP) conduct a survey of sensitive plants on the forest. -
Extreme Diversification of Floral Volatiles Within and Among Species of Lithophragma (Saxifragaceae)
Extreme diversification of floral volatiles within and among species of Lithophragma (Saxifragaceae) Magne Friberga,1, Christopher Schwindb, Paulo R. Guimarães Jr.c, Robert A. Ragusod, and John N. Thompsonb aDepartment of Biology, Lund University, SE-223 62 Lund, Sweden; bDepartment of Ecology and Evolutionary Biology, University of California, Santa Cruz, CA 95060; cDepartamento de Ecologia, Instituto de Biociências, Universidade de São Paulo, 05508-090 São Paulo, Brazil; and dDepartment of Neurobiology and Behavior, Cornell University, Ithaca, NY 14853 Edited by May R. Berenbaum, University of Illinois at Urbana–Champaign, Urbana, IL, and approved January 15, 2019 (received for review June 16, 2018) A major challenge in evolutionary biology is to understand how insects (10–12). Single volatiles can mediate interactions be- complex traits of multiple functions have diversified and codiversified tween plants and pollinators (13, 14), but complex floral scent across interacting lineages and geographic ranges. We evaluate bouquets are common in many taxa (15). These bouquets may intra- and interspecific variation in floral scent, which is a heighten the attraction of preferred pollinators, but they may complex trait of documented importance for mutualistic and function simultaneously as cues for resource detection by seed antagonistic interactions between plants, pollinators, and herbi- predators or herbivores (16–19) and mediate interactions with vores. We performed a large-scale, phylogenetically structured microbes (20, 21). Hence, floral scent should be sensitive to se- study of an entire plant genus (Lithophragma, Saxifragaceae), of lection imposed by the local assemblage of mutualist and an- which several species are coevolving with specialized pollinating tagonist insects, and it may vary among populations within floral parasites of the moth genus Greya (Prodoxidae). -
Monophyly and Relationships of the Enigmatic Family Peridiscaceae
TAXON 56 (1) • February 2007: 65–73 Soltis & al. • Monophyly and relationships of Peridiscaceae Monophyly and relationships of the enigmatic family Peridiscaceae Douglas E. Soltis1, Joshua W. Clayton1, Charles C. Davis2, Matthew A. Gitzendanner1, Martin Cheek3, Vincent Savolainen3, André M. Amorim4 & Pamela S. Soltis5 1 Department of Botany, University of Florida, Gainesville, Florida 32611, U.S.A. [email protected] (author for correspondence) 2 Harvard University Herbaria, Department of Organismic and Evolutionary Biology, Cambridge, Massachusetts 02138, U.S.A. 3 Royal Botanic Gardens, Kew, Richmond TW9 3DS, U.K. 4 Departamento de Ciências Biológicas, Universidade Estadual de Santa Cruz, Illhéus, 46.650-000, Bahia, Brazil 5 Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, U.S.A. Peridiscaceae, comprising Peridiscus, Soyauxia, and Whittonia, are an enigmatic angiosperm family of uncertain composition and placement. Although some have placed Soyauxia in other families (e.g., Flacourtiaceae, Medusandraceae), rather than in Peridiscaceae, sequence data for five genes (material of Whittonia could not be obtained) provide strong support for a clade of Soyauxia and Peridiscus. This evidence, combined with the strong morphological similarity of Peridiscus and Whittonia, support a monophyletic Peridiscaceae of three genera. Molecular analyses of a three-gene (rbcL, atpB, 18S rDNA) dataset for 569 taxa indicate that Peridiscus + Soyauxia together with Daphniphyllaceae form a clade that is sister to the rest of Saxifragales. Maximum likelihood and Bayesian analyses of Saxifragales using a five-gene (rbcL, atpB, matK, 18S rDNA, 26S rDNA) dataset place Peridiscaceae (posterior probability of 1.00) Peridiscaceae as sister to the remainder of Saxifragales, albeit without high posterior probability (pp = 0.78).