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Schmidt, Powell, & Halvorson In Cooperation with the University of Arizona, School of Natural Resources Vascular Plant and Vertebrate Inventory of Tuzigoot National Monument Vascular Plant and Vertebrate Inventory of Tuzigoot National Monument Inventory of Tuzigoot Plant and Vertebrate Vascular U.S. Geological Survey Southwest Biological Science Center 2255 N. Gemini Drive Flagstaff, AZ 86001 Open-File Report 2005-1347 Southwest Biological Science Center Open-File Report 2005-1347 October 2005 U.S. Department of the Interior U.S. Geological Survey National Park Service In cooperation with the University of Arizona, School of Natural Resources Vascular Plant and Vertebrate Inventory of Tuzigoot National Monument By Cecilia A. Schmidt, Brian F. Powell, and William L. Halvorson Open-File Report 2005-1347 October 2005 USGS Southwest Biological Science Center Sonoran Desert Research Station U.S. Department of the Interior University of Arizona U.S. Geological Survey School of Natural Resources 125 Biological Sciences East National Park Service Tucson, Arizona 85721 U.S. Department of the Interior Gale A. Norton, Secretary U.S. Geological Survey P. Patrick Leahy, Acting Director U.S. Geological Survey, Reston, Virginia: 2005 Note: This document contains information of a preliminary nature and was prepared primarily for internal use in the U.S. Geological Survey. This information is NOT intended for use in open literature prior to publication by the investigators named unless permission is obtained in writing from the investigators named and from the Station Leader. Suggested Citation Schmidt, C.A., B.F. Powell, B.F., W.L. Halvorson. 2005. Vascular Plant and Vertebrate Inventory of Tuzigoot National Monument. USGS Open-File Report 2005-1347. U.S. Geological Survey, Southwest Biological Science Center, Sonoran Desert Research Station, University of Arizona, Tucson, AZ. This report supersedes Powell et al. (2003, 2004a, and 2005a) and Schmidt et al. (2005). Printed on recycled paper Cover photo: Kristen Beaupre ii Authors Cecila A. Schmidt and Brian F. Powell School of Natural Resources 125 Biological Sciences East Building The University of Arizona Tucson, AZ 85721 William L. Halvorson USGS SBSC Sonoran Desert Research Station 125 Biological Sciences East Building The University of Arizona Tucson, AZ 85721 U.S. Geological SBSC Survey Sonoran Desert Research Station Personnel Charles van Riper III, Station Leader Steve P. Gloss, Ecologist William L. Halvorson, Research Ecologist Cecil R. Schwalbe, Ecologist Michael R. Kunzmann, Ecologist (Emeritus) Kathryn Thomas, Ecologist Pamela Nagler, Physical Scientist Phil Rosen, Ecologist Program and Expertise Areas of USGS and UA Personnel Administration & Outreach Fire Management Kate Leonard Dennis Suhre Jennifer Meador Cori Dolan Wendy Parrish James Feldmann Emily Sherbrooke Bill Halvorson Charles van Riper III Invasive Species Research Avian Ecology Patricia Guertin Claire Crow Jim Malusa Glenn Johnson Phil Rosen Chris O’Brien Cecil Schwalbe Brian Powell Brent Sigafus Charles van Riper III Dennis Suhre Kathryn Thomas Data Management Brent Sigafus Inventory & Monitoring Patricia Guertin Ecology of Amphibians & Reptiles Bill Halvorson Kevin Baker Pamela Nagler Cristina Jones Brian Powell Dave Prival Cecilia Schmidt Phil Rosen Cecil Schwalbe Vegetation Mapping & Ecology Brent Sigafus Patricia Guertin Eric Stitt Bill Halvorson Jim Malusa Kathryn Thomas USGS Southwest Biological Science Center http://sbsc.wr.usgs.gov USGS Southwest Biological Science Center, Sonoran Desert Research Station http://sbsc.wr.usgs.gov/sdrs iii iv Table of Contents Report Dedication .......................................................................................................................................x Acknowledgements.................................................................................................................................... xi Executive Summary ................................................................................................................................ xiii Chapter 1: Introduction to the Biological Inventories.............................................................................1 Project Overview...................................................................................................................................1 Report Format and Data Organization...................................................................................................1 Verification and Assessment of Results ................................................................................................3 Technical Concepts ...............................................................................................................................4 Chapter 2: Monument Overview ...............................................................................................................5 Monument Area and History .................................................................................................................5 Natural Resource Overview...................................................................................................................5 Natural Resource Management Issues...................................................................................................9 Chapter 3: Plants.......................................................................................................................................11 Previous Research and Collections......................................................................................................11 Methods...............................................................................................................................................11 Results .................................................................................................................................................15 Inventory Completeness ......................................................................................................................15 Discussion ...........................................................................................................................................15 Chapter 4: Fishes.......................................................................................................................................19 Previous Research ...............................................................................................................................19 Methods...............................................................................................................................................19 Results .................................................................................................................................................20 Inventory Completeness ......................................................................................................................20 Discussion ...........................................................................................................................................22 Chapter 5: Amphibians and Reptiles ......................................................................................................23 Previous Research ...............................................................................................................................23 Methods: Overview .............................................................................................................................23 Methods: Active Searches ...................................................................................................................25 Methods: Trapping ..............................................................................................................................30 Specimen and Photographic Vouchers................................................................................................32 Results .................................................................................................................................................33 Survey-Method Comparisons..............................................................................................................37 Inventory Completeness ......................................................................................................................38 Discussion ...........................................................................................................................................40 Chapter 6: Birds ........................................................................................................................................43 Previous Research ...............................................................................................................................43 Methods...............................................................................................................................................43 Results .................................................................................................................................................46 Inventory Completeness ......................................................................................................................51 Discussion ...........................................................................................................................................52 Chapter 7: Mammals ................................................................................................................................55