Upper Albian, Cenomanian and Upper Turonian Ammonite Faunas from the Fahdène Formation of Central Tunisia and Correlatives in Northern Algeria
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Acta Geologica Polonica, Vol. 70 (2020), No. 2, pp. 147–272 DOI: 10.24425/agp.2019.126459 Upper Albian, Cenomanian and Upper Turonian ammonite faunas from the Fahdène Formation of Central Tunisia and correlatives in northern Algeria WILLIAM JAMES KENNEDY Oxford University Museum of Natural History, Parks Road, Oxford OX1 3PW and Department of Earth Sciences, South Parks Road, Oxford OX1 3AN, United Kingdom. E-mail: [email protected] ABSTRACT: Kennedy, W.J. 2020. Upper Albian, Cenomanian and Upper Turonian ammonite faunas from the Fahdène Formation of Central Tunisia and correlatives in northern Algeria. Acta Geologica Polonica, 70 (2), 147–272. Warszawa. Over 130 species are documented from the Upper Albian, Cenomanian and Upper Turonian Fahdène Formation and correlatives in Central Tunisia and northern Algeria, based on material described by Henri Coquand (1852, 1854, 1862, 1880), Léon Pervinquière (1907, 1910), Georges Dubourdieu (1953), Jacques Sornay (1955), and new collections. The material consists predominantly of limonitic nuclei, together with adults of micromorphs. There is no continuous record, and a series of faunas are recognised that can be correlated with the zonation developed in Western Europe. These are the Upper Albian Ostlingoceras puzosianum fauna, Lower Cenomanian Neostlingoceras carcitanense and Mariella (Mariella) harchaensis faunas, the upper Lower to lower Middle Cenomanian Turrilites scheuchzerianus fauna, Middle Cenomanian Calycoceras (Newboldiceras) asiaticum fauna, Upper Cenomanian Eucalycoceras pentagonum fauna, and the Upper Turonian Subprionocyclus neptuni fauna. Two new micromorph genera are described, Coquandiceras of the Mantelliceratinae and Cryptoturrilites of the Turrilitinae. Most of the taxa present have a cosmopolitan distribution, with a minority of Boreal, North American and endemic taxa. Key words: Ammonites; Cretaceous; Albian; Cenomanian; Turonian; Algeria; Tunisia. INTRODUCTION wheel base Land Rover, and drive to northern Algeria and Central Tunisia (via Marseille and Algiers) to in- The origins of this contribution go back more vestigate the classic localities of Coquand (1852, 1854, than fifty years. It began in the spring of 1959 at the 1862, 1880), Péron (1866, 1883, 1896–1897), Thomas Colloque sur le Crétacé Supérieur Française, when and Péron (1889–1893) and Pervinquière (1903, 1907, the late Jake Hancock (1928–2004) and the late Jost 1910), together with those in the Monts du Mellègue Wiedmann (1931–1993) disputed on the affinities of (Dubourdieu 1953, 1956; Dubourdieu and Sigal the nuclei assigned to Submantelliceras Spath, 1923: 1949; Sornay 1955) that straddle the Algeria/Tunisia were they the nuclei of Mantelliceras Hyatt, 1903, or boundary. Our first visit to Algeria and Tunisia took Graysonites Young, 1958 (Basse de Menorval 1960, place between March 31st and April 22nd 1965. The p. 807)? In order to investigate the subject, Hancock trip was not without incident not least of which was (at that time my doctoral thesis supervisor) applied my rolling over the Land Rover in northern France. for and was awarded a grant from the then Natural We persisted, however, and arrived in Algiers. A visit Environment Research Council to purchase a long- to the British Consulate to seek advice on logistics 148 WILLIAM JAMES KENNEDY led to firm advice to return to the United Kingdom mens refigured in Kennedy and Wright (1981, 1984a, forthwith. We persisted, and carried out fieldwork in b), Wright and Kennedy (1978, 1979, 1996, 2015) and the area between Berrouaghia and Sour El-Ghozlane Kennedy et al. (2005). The Coquand Collection was (formerly known as Aumale: Péron 1866), made clas- subsequently returned to Budapest and is currently sic by Pervinquière (1910) on the basis of material housed in the Geological Museum of Hungary accord- collected by Phillipe Thomas, Alphonse Péron, and ing to Szives and Company (2011), but not all of the others, with slight results. Continuing to the Monts material I examined in 1985 has been traced. du Mellège, we left Algeria to discover that the bor- Mid-Cretaceous ammonites from Central Tunisia der zone, including some of Dubourdieu’s localities, were described and figured by Pervinquière (1907), was a sort of no man’s land several kilometres wide, and his material deposited in the Sorbonne Collections, bounded by barbed wire and minefields, set up during thereafter transferred to the basement store below the the Algerian War of Independence (1954–1962). Key Université Pierre et Marie Curie (Paris VI), where I outcrops were unvegetated, and strewn with tiny li- first studied it in the 1980’s. The specimens are now in monitic fossils. In contrast, when revisited in 1984, the collections of the Laboratoire de Paléontologie of many of these outcrops were under cultivation, and the Muséum national d’Histoire Naturelle, immacu- fossil collecting no longer possible. lately documented and illustrated in the Muséum’s on- It had been Jake Hancock’s intention to describe line catalogues. The larger specimens, from limestone these faunas in his retirement. He handed them over sequences, and some of the tiny limonitic specimens to me shortly before his death in 2004, but only now, were described and figured by Kennedy and Wright more than 50 years after our original collecting, are (1981, 1984a, b), Wright and Kennedy (1984, 1994, they described. In what follows, details of localities 1995, 1996), Kennedy (2004), and Kennedy and Gale and outcrops are based on Hancock’s field notebooks; (2015). Material from the Monts du Mellègue was de- his annotated field maps are, sadly, lost; my contribu- scribed by Dubourdieu and Sigal (1949), Dubourdieu tion was to collect the fossils. And the solution to the (1953), and Sornay (1955). Originally deposited in the disagreement between Hancock and Wiedmann? Both Collections of the Collège de France, much of the were correct to a degree. Some of the limonitic nuclei described material is lost (letter from Jacques Sigal, are Mantelliceras; some are Graysonites, but the type circa 1985). The Lower Albian material, now housed species of Submantelliceras is a paedomorphic dwarf. in the collections of the Université Claude Bernard Lyon I, was revised by Kennedy and Klinger (2008a) and Latil (2011); material described by Sornay (1955) PREVIOUS WORK is housed in the same institution. Material from areas in Central Tunisia outside Limonitic ammonite nuclei of Cretaceous ammo- those covered by the present contribution is described nites from north-eastern Algeria were described by in Robaszynski et al. (1993, 1994, 2004, 2008). Henri Coquand in a series of publications (Coquand 1852, 1854, 1862, 1880). Many species were described, but few were figured. This was partially remedied by LITHO- AND BIOSTRATIGRAPHY Charles Heinz, who, in 1886, produced five photo- graphic plates of fossils described by Coquand. The The present collections mainly come from iso- subsequent history of Coquand’s collections is dis- lated outcrops of the Fahdène Formation of Burrolet cussed by Szives and Company (2011). They were pur- (1956) and correlatives in Central Tunisia and adja- chased, on Coquand’s death, by Count Andor Semsey cent parts of north-eastern Algeria (Text-figs 1, 2); in 1882, and presented to the Hungarian Geological the reader is referred to the reviews of previous work Institute. Some of these specimens were described and in Robaszynski et al. (1994), Touir et al. (2005), con- figured by Pervinquière (1910), who stated them as be- tributions in Arnaud-Vanneau and Zghal (2005), and ing in the ‘Musée de Budapesth’. I first studied this ma- Latil (2011). The key references for localities in the terial in the summer of 1965, when it was housed in the Monts du Mellègue (the source of many of the fossils Geological Institute, Nepstadion Korut, Budapest. By described below) are the works of Dubourdieu (1953, the time of a second visit in 1985, the material had been 1956; in Sornay 1955). transferred to a field station at Sümeg, where I was able A discontinuous sequence of faunas is assigned to to describe and photograph most of Coquand’s speci- the zonal and subzonal sequence developed in Western mens that were described and figured by Pervinquière Europe (Text-fig. 3; Kennedy and Gale in Wright in 1910. Some of this work was published and speci- and Kennedy 2017). The ill-defined Arrhaphoceras MID-CRETACEOUS AMMONITES, NE ALGERIA AND CENTRAL TUNISIA 149 Text-fig. 1. Locality map, northern Algeria and Central Tunisia. The classic faunas from Algeria described by Pervinquière (1910), collected by Phillipe Thomas, Alphonse Peron and others came from between Berrouaghia and Aumale in Algeria, the latter now known as Sour El- Ghozlane. The Monts de Mellègue, source of much of the material described herein, lies in western central Tunisia and adjacent parts of Algeria to the west of Tadjerouine (Text-fig. 2) Text-fig. 2. Locality map, eastern Central Tunisia and adjacent parts of north-eastern Algeria 150 WILLIAM JAMES KENNEDY SUBSTAGE ZONE, WESTERN EUROPE FAUNA, PRESENT STUDY Neocardioceras juddii UPPER CENOMANIAN Metoicoceras geslinianum Calycoceras (P.) guerangeri pentagonum Acanthoceras jukesbrownei asiaticum MIDDLE CENOMANIAN Acanthoceras rhotomagense Cunningtoniceras inerme scheuchzerianus Mantelliceras dixoni LOWER CENOMANIAN Mantelliceras mantelli carcitanenseand harchaensis Pleurohoplites briacensis UPPER () ALBIAN Pervinquiera S. perinflata puzosianum (part) Pervinquiera ()S. rostrata Text-fig. 3. Ammonite faunas recognized