Zootaxa 4951 (1): 041–079 ISSN 1175-5326 (print edition) https://www.mapress.com/j/zt/ Article ZOOTAXA Copyright © 2021 Magnolia Press ISSN 1175-5334 (online edition) https://doi.org/10.11646/zootaxa.4951.1.2 http://zoobank.org/urn:lsid:zoobank.org:pub:557825A0-714A-426A-917F-1C9AB7372C30 Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation S. BRUCE ARCHIBALD1,2,3 & VLADIMIR N. MAKARKIN4* 1Department of Biological Sciences, Simon Fraser University, 8888 University Drive, Burnaby, British Columbia, V5A 1S6, Canada. https://orcid.org/0000-0002-4397-2497 2Museum of Comparative Zoology, 26 Oxford Street, Cambridge, Massachusetts, 02138, United States of America 3Royal British Columbia Museum, 675 Belleville Street, Victoria, British Columbia, V8W 9W2, Canada 4Federal Scientific Centre of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences, Vladivo- stok, Russia *Corresponding author. �
[email protected]; https://orcid.org/0000-0002-1304-0461 Abstract Four new species of Raphidiidae are described from the early Eocene of western North America: Megaraphidia antiquissima sp. nov. from McAbee, M. ootsa sp. nov. from Driftwood Canyon, M. hopkinsi sp. nov. from the Allenby Formation (all from British Columbia, Canada), M. klondika sp. nov. from Republic (Washington, United States of America). Archiinocellia Handlirsch, 1910, Archiinocellia oligoneura Handlirsch, 1910 from Horsefly River (British Columbia, Canada), and A. protomaculata (Engel, 2011), comb. nov., from the Green River Formation (Colorado, United States of America) are redescribed. Archiinocellia is assigned to Raphidiidae, sit. nov. The apparent absence of sclerotized gonocoxites 9 in the Archiinocellia protomaculata male is probably plesiomorphic at the family level.