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ut t wik Trans, tcpid, Soc, lapan 58 (1): 47-6S, January 2007

A review of the Latreille (, , ) from Korea

Bong-Woo LEEi'* and Yaiig-Seop BAE2'**

]i Entomological Laboratory, Graduate School of Li fe and Environmental Sciences, Osaka Prefecture University, Sakai, Osaka, 599-8531 Japan Zi Department of Biology, College of Naturul Scicnces, University of Incheon, Incheon, 402-749 Korea

Abstract 21 belonging to 14 genera of the tribe Pyralini, subfaniily Pyra]inae are recog- nized from Korea. Among thern, three species, Llypsopygia iwamotoi Kirp. & Yamanaka, fi. kawabei Yamanaka, and nanalis (Wilcrnan), are reported for the first time fOrm Korea.

Photographs of adults, illustrations of .uenitalia of both sexes, host plants, and flight period are pro- vided.

Key words Lepidotera, Pyralidae. Pyralinae, Pyralini, new record, Kerea.

Introduction

The Pyralinae belonging to the Pyralidae (Lepidoptera) comprises more than 900 species, which are mainly distributed in the tropical and Indo-Australian regions (Munroe & Solis, 1999). Many of them are notorious cosmopolitan pests of agricultural stored prod- ucts, and fbrest, such as R},ralis .farinalis (Linnaeus) (meal ) and dimidiata (Haworth) (tea tabby), The subfamily is distinguished from other by the fo]- lowing morphological characters: ocel]i present or absent; chaetosema present; maxillary pa]pus present; Iabial palpus porrect or ascending; proboscis well developed; fbrewing with R5 stalked with R, and R,, hindwings Sc+R, approximatcly or anastomesing with Rs. The Pyralinae is divided into two tribes, Pyralini and Endotrichini. Two tribes are defined by the venation of the hindwing: in the Pyralini Rs is not anastomosed with Sc+R,, whereas it is anastomosed with Sc+R, in Endotrichini, In Korea, two species, Orrybina regatis (Leech) and 7legulijbra bicoloratis (Leech), were initially reported by Leech (1 889), and further four species were added in 1891. Subsequently, several species were added to the Korean fauna by the Japanese specialists, Okamoto (1924), Shibuya (1927, 1928), and Maruda (1929). A Korean specialist, Park (1976, 1983, I993) added two more species. Recently Bae (2001) briefly reviewed the subfamily, covering 25 species.

In this anicle, the tribe Pyralini in Korea is reviewed, recognizing 21 species, among them,

Ilypsopygia iwamotoi Kirpichnikova & Yamanaka, H. kaivabei Yamanaka, and Bostra nanatis (Willeman), which are recorded for the first time from Korea,

Abbreviations used are as fOllows: CIS-Center for Systematics, Kangwon National University; FRI-Forest Research Institute, Seoul; NIAST-National Institute of Agricultural Sciences Tbchnology, Suweon; UIB-Department of Biology, University of Incheon, Incheon. TS-'Iiype species, TL-Ibrpe locality. HB-Hamgyeong-bugdo; HN-Hamgyeong- namdo; PN-Pyeongan-namdo; GW-Gangwon-do; GG-Gyeonggido; CB-Chungcheong-

bugdo; CN-Chungcheong-namdo; GB-Gyeongsang-bugdo; GN-Gyeongsang-namdo; JB-

*Present address: Division of Bio-Specimens & Genetic Resources, Korea National Arboretum, Pocheon, 487-821 Korea **Corresponding authon E-mail: [email protected]

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48 Bong-Woo LEE and Yang-Seop BAE

Jeonla-bugdo; JN-Jeonla-namdo; JJ-Jeju-do.

Subfamily Pyralinae Latreille

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Figs 1-21. Adults. 1.Aglossa diniidiata vsop)'gia regina (Butler, 1879). 3. H>pusop.vgia i}s,amotoi Kirpichnikova & Yamanaka, 1995, 4. H)rpsop>,gia kawabei Yamanaka, 1965, 5. P.vralis regalis [Denjs et SchitfermULIer], 1775, 6, P.yralis forinalis (Linnaeus, 1758). 7. albijimbrialis (Hampson, 1906). g. Stemmatophora valida (Butler, 1879). 9, 7Zimraca torridaiis (Lederer, 1863). IO. 712gulijlara bicoloralis (Leech, 1889). 11. Bostra nanaiis (Wilernan, 1911), 12, Orthop],・gia glat{cinalis (Linnaeus, 1758). 13, Orthom,gia nannodes (Butler, 1879). 14. 0rthopygia ptacens (Butlcr, 1879). 15-16. Hkrcutia peiasgalis (Walker, 1859), 17. Trebania flavij}'ontatis (Leech, 1889). 18. 5},brida mproxintans (Leech, 1888), 19. Datanoidesfasciatus Butler, 1878. 20. indicatorWalker. 1864. 21. 0r.v・hina regalis (Leech, 1889).

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Review of Pvralini 1'rom Korea 49

Tribe Pyralini Latreille

GenusAglossa Latreille, 1796

Aglossa Latreille, 1796, Pre'cis Carctctbreh' ge'ntiriqttes lnsectes: 145. TS: Phalaena pinguiiralih I.innaeus, 1758. Euctita HUbner, 1825, Verz bekannterSchmett.: 347, TS: Phalaena pinguinali.s Linnucus, 175g. Philotis Ragonot, 1891,Annls Soc. ent. Fr. 60: 24 (key), 81. TS: Philotis raclamalis Raguiiot, 1891. Agriqpe Ragonot, 1894,Annis Soc. ent. Fr, 63: 163. TS: Aglossa hrabanti Ragonat, l884.

Aglossa dimidiata (Haworth, 1810) (Figs 1, 22, 42)

Crambus dimidiata Haworth, 1810, Le7,id. Br.: 372, TL: India. Aglossa achatina But]er, 1 879, Iiiust. opical specimens Lci)id, neterocera Colln Br, Mus, 3: 72, TL: Japan. Pyraiis circulctris Motschulsky, 1860, Etudes ent. 9: 73. TL: Japan. Aglvssc; mic'atialis Walker, 1859, List Specimens lepid. CoUn Br. Mtts. 17: 227. TL: Chjna (Shanghai). Aglossa dimidiata: Matsumura, 1905i 201; Nakayama & Okamoto, 1940: 215: Inoue, 1954: 194; Mutuura, 1957: 106; Zool. Soc. Ker,, 1968: SO; Seok, 1970/ 144; Kor. Soc. Pi. Pret., 1972: I37; Inoue, 1982, 1: 380, 2: 246, pl. 45: 51; Park, 1983: 402, 887. pL 27: 445a, b; Yamanaku & Yoshiyasu, 1992: g9; Kim & Shin. 1996: 30; Bae, 2001: 172, fig. 162.

Aduit, male genitttlia, and female genitalia in Figs 1, 22, 42.

Materials. 13 males and 2 females examined in this study.

Distribution, Korea (GW, GG, GB, GN, JJ), Japan, China, Myanmar, and India,

Host, Storedgrain(Park, 1983).

Flight period. May to July.

Remarks. This spccies is one of the well-known pests for stored grain. It is well distin- guished frorn other species by the following external characters: fbrewing narrow and long, ante- and postmedial ljnes serrate, seven dark fuscous spots distinct along termen of forewing.

Genus Ilypsopygia HUbner, 1 g25

H}psop.vgia HUbner, 1825, Vkrz. hekannterSchmett.: 348. TS: Phalaena costalib' Fabricius, 1775.

Hlypsopygia regina (Butler, 1879) (Figs 2, 23, 43)

P.}'ratis regina Butler, 1879. Ann, Mag, nat. Hist. (5) 4: 452, TL: Japan. Hmph'op.),gia regirta: Ragnot, 1g9I, 7Lran.s', ent. Soc'. Fr, 60: 41; Hainpson, 1896, fuuna Br. India () 4: 149; Hampson, lg96, 7:F'ans. ent. Soc. Lond. 1896: 507; Leech, 1901: 422; Matsumura. 1905: 201; Mutsumura, 1925: 187; Shibuya, 1928, 21: 157, pl, 3, fig. 3; Marumo, 1939, IVosi Kait},o Sir.vo 142: 10, p]. 1, fig. 2, pl. 3, fig. 10, p]. 4, fig. 9, pl. 6, fig. 4: Inoue, 19S5: 149; Mutuura, 1957: 107, pl. 18, fig. 570; Inoue, 1959: 243, pl. 168, fig. 20; Yamanaka, 1965: 68, pl. 10. figs 1, la, pl. 11, pl. 13, fig. 22; Wang, 1980i 94, pL 2, fig. 72; Inoue, ]982, 1: 381, 2: 247, pl. 45, fig, 55; Park, i983: 412, 891, pL 28: 460a, b (part,); Yamanaka & Yoshiyasu, 1992: 89; Park, 1993: 154; Bae, 200l: 176, fig. 166.

Adult, male genita]ia, and female genitalia in Figs 2, 23, 43.

Materials, 24 males and 20 females examined in this study,

Distribution. Korea (GWI GG, GB, GN, JN), Japan, Myanmar, and India.

Flight period. June to August.

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50 Bong −Woo LEE and Yang −Seop BAE

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《 総 論 戴 一黙藩 警概 嶝∵\ 勳 Figs 22− 27, Male genitalia. 22. Ag ’o ∬a ditnidiata (HawQrtb,18 ]1) .23 , H

θ pygl α ア ∫ regina (Butler,1879), 24 , H . vpsop )’gia irvamotoi Kirpichnikova &Y 1995 25 ρ‘ regatis et Schif a 副 (a, . , 恥 ∫, pygia kawabt ・i Yamanaka,1965.26. Py厂‘ilis [Denis rmUller ],1775. 27 .Pyrcttisfttr’natis( Linnaeus , 疋 7

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of Korea 51 Reyiew Pvralini" from

Remarks. This js a common species in Korea. It is similar to H. iwamotoi ill its superficial appeararice, but separable from the latter by the following character: presence between ante- and postmedial lines of hindwing of a suffusion of red scales.

Elyps(m},gia iwamotoi Kirpichnikova & Yamanaka, 1995 CKorean name: Norangtti-aegi- bidanmyeongnabang) (Figs 3, 24, 44)

H),psop.vgia i}t,amotoi Kirpichnikova & Yamanaka, 1995, Tinea 14: 20()-203, figs 1, 2, 4, TI.: Russja.

H)Jpsopygia sp.i Bae, 2001: 177, fig. 167.

Adult. Wingspan 11-18 mm (Fig. 3), Hcad dark gray on frons; vertex with tuft rnixed with dark fuscous. Labial palpus dark gray; basal segment reddish brown; apical segment ochre- ous terminally; rnedian and apical segment ascending. Antenna 4/5 of length of forewing, pale purplish fuscous mixed with gray. Thorax pale reddish purpIe; patagium dark gray ir- rorated with dark fuscous; upper side of tegulae pale gray, lower side pale reddish purple. Abdomen dark gray, mixed with reddish purple. Forewing: ground color purpIish red, light- ly scattered with blackish scales; antemedial ljne narrow, pale yellow, rather indistinct, slightly sinuate; pestmedial line also narrow, yellowish, arising from a yellow costal patch, and a]most straight from costa to dorsum; costa with several minute, yellowish specks be- tween ante- and postmedial Iine. Cilia bright yellow. Hindwing: ground coler of upper side conco]orous with forewing; ante- and postmedial lines narrow, yellowish. Cilia as in fOrewing.

Male genitalia (Fig. 24). Uncus rather short, stumpy, with short hairs dorsally; gnathos weakly sclerotized, slender central projection well developed, pointed at top, slightly hooked distally; valva rather long, simple, a little pilose; saccus rather narrow, long, ta- pered; juxta nearly circular, slightly swollen, a weakly sc]erotized plate. Aedeagus weakly sclerotized, rather long, about 1.3 times as ]ong as valva; cornutus sclerotized, rather long and straight, thorn-shaped; ductus ejaculatorius arising from near midd]e.

Female genitalia (Fig. 44). Apophysis posterioris and anterioris slender, long; ostium bur- sae long, slender, with sclerotized incomplete ring; ductus bursae long, weakly sclerotized;

ductus seminalis originating near top of ductus bursae; corpus bursae ovate; signum sclero-

tized, ellipsoidal, with sclerotized thorns.

Material examined, [GW] Jinburyeong: 1 9, 7. VII, 1998 (Bae et al,)-coll. UIB. [GG] Mt Kwangduk: 1 r, 20, VII. 1996 (Bae et al.); Mt Soyo: 2 4, 5, VIII. 1996 (Bae et at.). [GB] Mt Sokri: 1?, 17. VII. 1998 (Paek, Ahn, Kim); Mt Sobaek: 1r3\, 17. VII. 1998 (Bae, Lee & Song)-coll. UIB, [GN] Mt Jiri: 3 X, 14. VII. 1976 (K,T. Park), 1 l, 14. VH, 1976 (J,S, Hyun), 3 9, 14. VII. 1976 (Y YL Ha),3 9, 16. VII. 1976 (K.T. Park), I 4, 17. VII. 1976 (Y,Yl Ha)-coll. NIAST. [JNI Mt Baekun: 3 if' 12 4, 19. VII, 1998 (Bae et al.)-coll. UIB.

Distribution. Korea (GW, GG, GB, GN, JN) and USSR (Far East).

Flight period. July to August.

Remarks, This species is reported for the first time from Korea, where it has been confused with H, regina, Park C1983a) illustrated two moths under the name of H. regina from Korea, but one of them (fig. 460a) seems to be this species. External characters of this species resemble those of H. regina, but it is distinguished by the fo11owing characters: upper side of hindwing concolorous with forewing, with a remarkable medial yellow band; uncus wjder than that ofH. regina.

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52 Bong-Woo LEE and Yane-Seop BAE

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Figs 28-33, Male genitalia. 28. Stemmatophora alb(timbriatis (Harnpson, 1906). 29. Stemmaio- phora valida (Butler,1879). 30. Tamrat'a forridalis (Lederer, lg63). 31. Tegulijbra hicot- oratis (Leech, 1889), 32. Bostra nanatis (VVileman, 1911). 33. 0rthop.ygia glaucinatis (Linnaeus, 1758). Scales: 2mm.

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Review of Pvra]ini fi'om Korea 53

Ilypsopygia kawabei Yamanaka, 1965 (Korean name: Hwoisaek-aegibidanmyeongnabang) (Figs 4, 25, 45)

HJpsopygia kawahei Yamanaka. 1965, Tirtea 7: 68, pl. 1O: 2. TL: Japan.

Adult, Wingspan 16-20 mm (Fig. 4). Head yellowish brown, irrorated with dark fuscous on frons; vertex yellowish brown, Labial palpus yellowish brown, outer surface of median 'fuscous; segment irrorated with dark median and apical segments ascending. Antenna 3/4 length of forewing, yellowish ash-gray, annulated with dark fuscous; scape yellowish brown, Thorax yellowish brown, mixed with fuscous; patagium and tegulae yellowish brown, irrorated with fuscous, Abdomen yellowish brown, mixed with fuscous, Forewing yellowish ash-gray mixed with fuscous scales, costa narrowly edged and speckled with fus- cous and yellow; antemedial Iine cntirely absent; postmedia] line fuscous, parallel with ter- men. Hindwing pale bright gray; ante- and postmedial lines faint.

Male genitalia (Fig, 25). Uncus stumpy, with few hairs dorsally; gnathos weakly sclero- tized, slender; central projection rather short, pointed at top, slightly hooked distally; valva rather long, pilose; saccus long, tapered; juxta stumpy, slightly swo]len, a weakly sclero- tized plate. Aedeagus weakly sclerotized, rather long, about 1.4 times as long as valva; cor- nutus sclerotized, thorn-shaped; ductus ejaculatorius arising firom basal 113.

Female genitalia (Fig, 45). Apophysis posterioris rather long, about 1.4 times as long as apophysis anterioris. Ostium bursae funnel-shaped, with sclerotized ring; ductus bursae s]ender long, with strongly sclerotized strap, connected with basal area of signum; ductug

seminalis originating near middle of ductus bursae; corpus bursae ovate; signum three prominent thorns, the terminal one 1argest.

Material examined, LGWI Mt Dukga: l9, 24. VII. 1997 (YIS. Bae & N.H. Ahn)l Mt Balgyo: 1 8, 7. VII, 1998 (Paek, Lee, Kim & Song); Mt Jembong: 1 gi 3 9, 24, VII, 5. VIII. 1997 (YS. Bae & N,H, Ahn)-coll. UIB. Gwangleung: 1 9, 10, VI, 1998 (B,K. Byun &, Y,S, Park), 1\, 29. VI, 1998 (B.K. Byun), 13. VII. 1998 (B.K. Byun)-coll, FRI. Chuncheon: 18', 1. V 1989 (K.T. Park)-coll. NIAST, LGG] Mt Gwangdeok:1-',19. VITI. 1997 (M.K. Paek & U,"l Jang); Mt Hwaak: 2r3\, 19. VII, 25. VII. 1998 (Paek, -9, Lee, Ahn & Kim); Mt Ybngmun: 1 11. VIII. 1998 (Lee, Ahn & Kim)-coll. UIB. Mt Cheonggae:5\, 16. VII. 1998 (D.S. Gu)-coll. NIAST. [GB] Mt Sokri:2?, 17. VIL 1998 (Paek, Ahn & Kim)-coll. UIB. [GN] Mt Jiri: 2 4, 5-6. VI, 29, VII. 1997 (M,J, Han & D.S, Gu); Is. Geeje: 1 \, 4-5. VI. 1997 (D,S. Gu)-coll. NIAST, [JB] Mt Mireuk: 1 ?, 23, VIJI. 1997 CB,W, Lee &N,H, Ahn)-coll. UIB. Mt Palgong, Gunwi: 1 8 1 ?, 27, VIL 1998 (D.S. Gu)-coll. NIAST. IJN] MtBaekun:1\, 19. VII. 1998 (Bae etaL)-coll. UIB. Mt Baekun: 1 9, 21, VIII. 199 1 (S.B. Ahn)-coll. NIAST.

Distribution. Korea (GW, GG, GB, GN, JB, JN) and Japan.

Flight period. July to August

Remarks. This species is reported for the first time from Korea. It has been known as an endcmic species in Japan, but it was fbund throughout the country during this study, lt is easily distinguished from the other species by the following characters: adult small in size (wingspan 11-18 mm); ground color ash-gray; antemedial line faint; signum three promi- nent thorns.

Genus Ib;ralis Linnaeus, 1758

Pyralis Linnaeus, 1758. S),st. Nat. (Edn 10) 1: 533. TS: Phalaena.farinatis Linnaeus, L7S8. Aletes Rafinesque, 1815, Analyse de ta Nature: 129.

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S4 Bong-Woo L]・/]E and Yang-Seop B.4E

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Figs 34-39. Male genitalia. 34, Orthopygia nannodes (Butler, l879). 35, Orthop},gia ptacens (Butlcr, 1879), 36. Herct{tia petasgatis (Walker, L859). 37. Trehania .flav(frontatis CLeeeh, 1889), 38. S:bride approximans (Leech, 1888), 39. Datanoides,1'i sciaius Butler, 1878. Scales: 2 rmn.

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Review of Pyralini from Korea 5S

tt 40 41

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Figs 40-41. Male genitalia. 40. Armpara indicator Walker, 1864. 41. 0Tl},bina regalis (Leech, 1889). Sca]es:2mm,

spr rella Rafinesque, 18t5, Anatyse de la IVIiture: 129. Ceropsina Rafinesque, ]815, Anal}'se de la IVature: 129. Asopia Treitschke, 1828, Schfnett. Euf: 6 (2): 316. TS: Phalaenafarinatis Linnaeus, 1758. Sacatia Walker, 1863, List Specimens tqpid. inse(rts CoUn Br. Mus. 27: 123. TS: Sacatia laudatella "[alker, 1863, Tlierapne Ragnot. 1890,Annls Soc. ent, Fr, (6) 10 (Bull.): xciii, TS: Stemmatqphora obsoletalis Mann, 1864. Eutrichodes Warren, 1891, Ann. Mag, nat. Hist. (6) 7: 498. TS: P.yralis ravotalis Walker, 1859,

Ilyratis regalis LDenis et SchiffermUller], 1775 (Figs 5, 26, 46)

R},ralis regatis [Denis et SchifierrnUller], 1775, Anktindung s.vst, Werkes Schmett, Wienet:gegend: 124; Matsumura, 1905: 202; Inoue, 1955: 150; Mutuura, 1957: 108; Inoue, 1959: 242, p]. 168: 23; Yamanaka, 1965: 70; Zeel. Soc. Kor., l968: 50; Kor. Soc. Pl, Prot,, 1972: 140; [noue, 1982, 1: 381, 2: 242, pL 46: 4; Park, 1983[ 404, 887, pl. 28: 447; Park & Ju, 1990i 392: Yalnanaka & Yoshiyasu, 1992: 89; Kim, ]999: 2] ; Bae, 2001: 183, fig. 172. TL: Austria. P.vralispulchellalis TvTilliere, 1873, Cat. Lep.Aipes-Adarit.: 221. TL: Africa (Congo), Ayralis princeps Butler, I889, nlust, mpical Specimens Lqpid, Heterocera Colln Br. Mus. 7: 91. TL: India (Dhartnsala).

Adult, male genitalia, and female genitalia in Figs 5, 26, 46.

Materials, 83 males and 87 females examined in this study.

Distribution. Korea (GW; GG, CB, CN, GB, GN, JB, JN, JJ), Japan, China, Taiwan, Russia (Siberia), Myanmar, India, and Central & Seuthern Europe.

Hest. A comb of wasps in Japan (Inoue, 1982).

F]ightperiod. June to September.

Remarks. This species is common in Korea and can be distinguished frorn other species by

the forewing marking and Iong corpus bursae.

IlyraIis farinatis (Linnaeus, 1758) (Figs 6, 27, 47)

Phaiaenafarinatis Linnaeus, 1758, S.vst, Nat, (Ed. 1O): 533, TL: Sweden (Tunisia). P.vraiisforinalis: Butler, 1879, Itlust, 4vical specimens Lepid. Heterocera Colln Br. Mus. 3: 70; Matsumura,

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56 Bong-Woo LF.F. and Yang-Seop BAE

1905: 202; Okamoto, 1924: 148; Inoue. 1954: 150; Mutuura, 1957: 108; lnouc, 1959: 242, pl. 168: 21; Yamanaku, 1965: 69; ZooL Soc. Kor., l968: 50: Seok, 1970: 147: Kor. Soc. P]. Prot., 1972: ]40; Inoue, 1982, 1: 381, 2: 247, Pl, 46: 1; Park, 1983: 403, 887, pl. 28: 446a. b: Yamanaka & Yoshiyasu, 1992: 89; Park, 1993: 154; Kim, 1999: 21; Bae,lOOI: 183, fig. 172. Ry,ratis,fraterna Butler, 1879, lltust. typical Specimens Lepid. Heterocera Coltrt Br. Mus. 3: 70. TL: Japan. P.vralis meridionaiis Schfnidt, 1934, Aren. Mag. nat. Hist, (1O) 14i 543. TL: Gt, At]as, ll}'ralis orientalis Amsel, 1961, Ark. Zbot, (N. S.) 13: 401, TL: Persia,

Adult, male genitalia, and female genitalia in Figs 6, 27, 47.

Materials. 12 males and 4 females examined in this study,

Distribution. Korea (GG, GB, GN. JJ), Japan, China. [faiwan, Ethiopian region, Aighanistan, Nearctic & Neotropica] region, Australia, and Ncw Zealand.

Host. Stored grain (Park, t983; Inoue, 1982).

Flight period, May to August.

Remarks. This species is cosmopolitan, and is a serious pest of stored grain and grain prod- ucts. Its common name is meal moth. It probably has one or two generations yearly and overwinters in the larval stage, This species is often collectcd near or in houses.

Genus Stemmatophora Guenee, 1854

Stefitntat(nyphora Guenee, 1854, in Boisduval & Guenee, Hist, nat. I}Tsectes (Spec. g6n. L6pid,) 8: 129. TS: Asopia combusTalis Fischcr von ROslerstamm, 1842. `:t884-7'i), Moore, 1884 (not Lepid. Ceylon3: 27. TS: Mdradana rivttlata Moore, 1884. Korematepis Hanipson, 1891, Illust. typical Specimens Lepid. Heterocefu Coltn Br. Mus. 8: 34, 129. TS: Koremalepis' scopttia Hampsen, 1 891 ,

Stemmatqphora athijimbrialis (Hampson, 1906)

Her(:ttlia alb(fimbrialis Hampson, 1906, Ann. Ma.o. , nat. Hist. (7) 17: 264. TL: China. Stemmatophora albijin'tbriaii.s: Inoue, 1985, Akitct (N. S.) 72: 4; Bae, 2001: 189, fig. 176. StemJnatophora th'i.{shimensis Inoue, 1982, 1: 382, 2: 247, pl. 46: 9; Purk, 1993: 154. TL: Japan,

Adult, male genitalia, and female genitalia as Figs 7, 28, 48.

Materials. 23 males and 21 females cxamined in this study.

Distribution, Korea (GW, GG, GB, GN, JN), Japan. China, and Tlaiwan.

Flight period. April to August

Remarks. This species was recorded by Park (1993) frem the collection of North Korean materials preserved in the Hungarian Natural History Museum. It is similar to S. validti, but can be di'stinguished by the fbllowing: forewing ground color yellowish brown; postmedia) line pale yellow; va]va wider than that of S. valida; corpus bursae long,

Stemmatophora valida (Butler, 1879) (Figs 8, 29, 49)

Pyralis vati.da Butler, 1879. Ann, Mdg. nat. Hist. (5) 4: 4S1. TL: Japan, Stemmatq]horct v'alida: Matsumura, 1905: 202; Inoue, 1955: I50; Inoue, 19S9: 243, pl. 168: 25; Park, 1976r 17; Inoue. 1982, 1: 382, 2: 247, p], 46: 8; Park, 1983: 410, 890, pl. 28: 456; Park, 1993i 154; Bae, 2001: 190,fig.177.

Adult, male genitalia, and female genitalia in Figs 8, 29, 49,

Materials, 14 males and 9 females examinecl in this study.

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57 Review of Pyralini from Korea

Distribution. Korea (GG, GN, JN, JJ), Japan, China, and N.W. Himalaya.

Ftight period. June to Aus,ust.

Remarks. This species is similur to Ztgt{.l(fera hicotoratis in pattern and wing coloration, but it is distinguished from the latter by the fo11owing characters: larger wingspan (17-23 mm); forewing postmedial line arising from a small pale yellow speck on costa; ground color pale grayish brosu'n; basal line black; male genitalia with short saccus; cornutus ab- sent; in fcmale narrow ostium bursae with sclerotized ring.

Genus 7timraca Moore, 1888

7'antraca Moore, 1888, Lepid. Ceylon 3: 554. TS: Asopia torridalis I.ederer, 1863.

7hmraca torritlalis (Lederer, 1863) (Figs 9, 30, 50)

Asqpia torridalih' Lederer, 1863, Wien. ent. Monatsc'hr. 7: 342, TL: Sri Lanka. 7timraca torridalis: Matsumura, 1905: 202; lnoue, 1954: 151; Mutuura. 1957: 108; Lee & Park, 1958: 6; Inoue, 1959: 243, pl. 168: 26; Zool. Soc. Kor., 1968: 50; Inoue, 1982, 1: 3g2, 2: 247, pl. 46: 10; Park, 1983: 411, 891, pl, 28: 4S8; Yamanaka & Yoshiyasu, 1992: 89; Bae, 2001t191, fig. 178. TL: Sri Lanka. VZmina incerta Wa[ker, 1865, List 5pecimens tepid. Ihsec'ts Colln Bn Mus, 33: 829, TL: Indiu. Coenodomtts brunneus Suzuki. 19l5, after Yamanuka & \oshiyusu (1992).

Adult, male genitalia, and female genitalia in Figs 9, 30, 50.

Materials. 15 males and 18 females examined in this study.

Distribution, Kerea (GG. GB, GN, JB, JN, JJ), Japan, China, India, Sri Lanka, Myanmar, and Java.

Flight period. July to August.

Remarks. This species seems to be common in Korea, and is easily distinguished from other species by the fOllowing characters: fbrewing with serrated postmedial line; juxta cylindrical, with small thorn-shaped top; ma]e genitalia with zigzag-shaped cornutus; fe- male genitalia with rather small corpus bursae without signum.

Genus 71?gulijlera SaalmUller, 1880

Tlegt`lijlera SaalmUller, 1880, Ber, senckenb. natucfi Ges. 1879-i880: 305. TS: Tl?rgul(fera rubicttndatis SaalmUlter, 1880.

f'tv'imacia Marumo, 1939, IVbzi Kitiryo Sis),・o 142: 20. TS: hicoloralis Leech, 1889: 65.

71?gulijlena bicoloralis (Leech, 1.889) (Figs 10, 31, 51)

Endotricha bi(,otoNtlih' Leech, 1889, Entomomogist 22: 65. pl. 4: 17. TL: Korea. 1'egulijiera bicoloratis: lnoue, 1982, 1: 382, 2: 247, pL 46: 11; Park, 1983: 41l, S90, pl. 28: 457; Park & Ju, 1990: 392: Yamanaka & Yoshiyasu. 1992: 89; Park. 1993: 154; Bae. 2001: 193, fig. 179. Stemmatophora bicotoraiis: Matsumura, 1905: 202.

FLofimacia bicoloratis: Marumo, 1939, IVt)zi Kainy,o Siryo 142: 20; Inoue, 1954: 151; Mutuuru, 1957/ 108; Inoue, 1959: 243, pl. 168: 27; Zool. Soc. Kor. 1968: 50. Pvratis dtttcicutatis Swinhoe, 1889, Proc, zoot, Soc, Lond. 18S9: 418. TL: India.

Adult, male genitalia, and female genitalia in Figs 10, 31, 51.

Materials, 18 males and 18 females examincd in this study.

rlaiwan, Distribution. Korea (GW, GG, GN, JB, JN), Japan, China, and lndia.

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58 Bong-Woo LEE and Yang-Seop BAE

. zle

mN-y ec -f,,,,7 :t.

x " 'l・ ii' isi;・ikit 1 i ){,ajN /. kti,

,k-l., .tilgili)x 't/ i'f/i;i.・'--15.'IX,, i..i>,,1, ・N ,/i.-1.tl/f//fii' l/"ti .:. t ,1 E.ki',ti:g..,,..・x.X .. .. ,t llkk ..111.l.1i,,,,, E,ii llifR ,f.,fl T-"-S " t. t t ttt ・,- ・. ・ ,/ ii/ ,i. . 1&x,/t IX di'q・. t''' '{ / 'i'' ' 'l //t ' .tt t ttt A lj11・5X t tt / t, t/t t tt /K ・t 'V':"Vny'''`-/11'...u..."-. st tt /) IK . 1, ttt t/ t' "x' .t t tt j" /} t '1 i 'tt'{1/il/'' /1 T tl" // ./`' 42 'v..・ ,t,43 'K.L..!t44 "

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,g.7g.

'"l' ttl'lit",l X) ・tts kl ,//' ]li f.- 'S---lj---:;i TT,-R", .

X" '・,

/ g・.g ?・'tt ; ' t "' ''"t ' ' re ''X lu ' ,・"' /ii't i" ,・/' 'xLg..ltt"' t.t L ../') ij IIs,/1'""'lil , l i ' y IS X ff '1 ,n 1t. .1 ' .t t Lt.;,,,,./2> s' ' !. 'i.' S,i-iS)>,, t ,r U 7 (C, '45 --' v.・ R 46 47

Figs 4247. Female genitalia.42. Agtossa dimidiata (Haworth, t81t), 43, H)ipsop.vgia regina (Butler, 1879). 44. H>psop.vgia iivamotoi Kirpichnikova & Yamanaka, 199S. 45.

Ll}'psop.vgia ka}vabei Yamanaka I965. 46, et SchiffennUllerJ , [Denis ,1775. 47. Py・ratisf}xrinalis (Linnaeus, 1758). Scales: O.5 mm.

Flight period. July to August.

Remarks. This species is superficially similar to Stemmatophotu but is distin- guishedfrom the latter by the fbllowing characters: adult sma]1 invatida, size (wingspan 15-

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Review of Pyralini frQm Korea 59

18mm); ground color of forewing dark red; male genitalia with swollen sacculus at base, saccus rather large, aedeagus strongly curved, cornutus present; in female signum absent.

Genus Bostrti Walker, 1863

Bostra Wa]ker, 1 863, List opecimens tepid. insects CoUn Br. Mtts. 27: 1 23, TS: Bosra illttseUa Walker, 1863.

Bostra nanalis (Wileman, 1911) (Korean name: Bora-bidanmyeongnabang) (Figs 11, 32, 52)Ryralis

nanalis Wileman, ].911, 7)'ans, ent. Soc. Lond. 1911: 369. TL: Japan.

Bostra nanalis: Inoue, 1982.. 1: 382, 2i 247, pl, 46: 12. Endotricha rvijicosta Wileman & South, 1917, Entomologist 50: 175, TL: Taiwan.

Adult. Wingspan 9-11 mm (Fig. 11). Head reddish purple, with yellowish brovvn laterally; vertex irrorated with pale yellowish brown, Labial palpus reddish purple, mixed with dark gray; apical segment dark gray, ochreous terminally; median and apical segment ascending, Antenna 112 length of forewins,, reddish purple, mixed with pale yellowish brown. Thorax dorsally pale reddish purple, ventrally ochreous; patagium and tegulae reddish purple; pos- terior crest ochreous, irrorated with pale reddish purple, Abdornen reddish purple, mixed with paie yellowish brown, Forewing narrow; ground color reddish purple; costa finely edged with black and marked with whitish, and with a black spot on cell; antemedial line pale blackish brown, faint; postmedial line blackish outwardly, angled below costa, Hindwing reddish purple; ante- and postmedial lines rather distinct, curved and sinuous,

Male genitalia (Fig, 32). Uncus long, tongue-shaped, with few hairs dorsally; gnathos weakly sclerotized, slender; centrul projection well developed, tip pointed, slightly hooked distally; valva simple, narrower apically, slightly pilose; saccus rather narrow, slightly swollen; juxta stumpy, weakly sclcrotized. Aedeagus weakly sclerotized, rather long, about same length as valva; cornutus sclerotized, rather long, thorn-shaped; ductus ejaculatorius arising from near middle,

Fema]e genitaiia (Fig. 52). Apophysis posterioris slender, very short, about O.4 times as long as apophysis anterioris. Ostium bursac bowl-shaped, with sclerotized i'ncompletc ring;

ductus bursae weakly sclerotized, long, slightly spiraled from 1/2; ductus seminalis origi- nating near top of ductus bursae; coipus bursae ovate; signum sclerotized, romboid, with three minute thorn-shaped projections.

et al,)-coll. Materialexamined. [JB]Mt Jangan:4 8 1 9 , 21.VII.1998 (Bae UIB. [GB] Kyeoaju: 1\, 5, VIII. 1998 (S.K, Lee)-coll, UIB. [JN] Naju: 18, 15. VII. 1998 (S.H. Kim)-coll. FRI.

Distribution. Korea (GB, JB, JN), Japan, and [[aiwan,

Flight period. July to August.

Remarks. The genus Bostra Walker and this species are recorded fbr the first time from Korea. This species is easily distinguished from the other species by the fo11owing charac- ters: adult small in size (wingspan 9-11 mm); ground color reddish purple; valva simple and tapered; ductus bursae slightly spiraled.

Genus Orthopygia Ragonot, 1891

Orthopygia Ragonot, 1891, Annls Soc. ent. Fr. (6) 10 (Bull.): xciii. TS: Phataena gtaucinalis Linnaeus. 1758.

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60 Bong-Woe Lm and Yang-Seop BAE sts

V ,k' /.tt(1.--..,L・, 1 it x 'v

xltttt '・-.. 9 -,-.・' A, //

1 / 'xETte'' // ,, tt!1 ttJ/ f' 'j 1,1 // / /" f 7t/1,1. .tl -N '. .tte・ i 1 . t/'t/ .. I '・sLt,.V'50 1./t.k .,K. 1 x& ・ ---/ 48 ya 49

ttttt

w.,i .tw.xti>t.x ft'z /,lrJ,. f t11 i' tf' . gi'IS,4 '11"i''t'i': //t"../tt/tttt''.tl'.',/.i,.. t ,'ix・1 Rx t1. ,/ i l,/f' ・t 4--k' ' 1. ' /"'//'' ,x -t",,UL Lt.U $l

tttiy -,i/' kee ti1! x'/x '/'-' ''・'F' ・ ;b t t.,.ttttt lt. }l, 1・1 '/'111'lr.v-=1'' g/k Ldeif

vk-'stRlfN "x,

.... 1, 9E/t ...... I "L ,vb xxk.vs7 .M." /kY53 (Z・・ t)y x.-.x". . 51 52

Figs48-53. Female genitalia. 48. Stemntatoj)hora albtfimhrialis (Hampson, 1906). 49, Stemmatophora vatida (Butlcr, IS79). 50. 7Zimracra torridatis (I.ederer, 1863), Sl, Tlegutijkra bicotoratis (Lccch, 1889). 52. Bostra nanalis CWileman, 191 t). 53. 0rthop.vgia glaucinalis (Linnaeus, 1758). Scalcs: O.5 mm.

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6[ Rcview of Pyralini from Korea

Orthopygia glaucinalis (Linnaeus, 1758) (Figs 12, 33, 53)

P.vralisgtaucinalis Linnaeus, 17S8, S>,st, Nat. Ed. Ier 533. TL: Eurupe, Orthopygia gtattcinaiis: Yamanaka, 1965: 71: Inoue, 1982, 1: 383,2: 247. pl. 46: 1O; Park, 1983: 407, 889, pl, 28: 452; Park & Ju. 1990: 392; Purk, 1993: 154; Baq 2001: 177. fig, 168, Hercttlia glaucinalis: MaLsumura, 1905: 202; Zool. Soc. Kor. 1968, 2: 50; Kor. Soc. Pl. Prot, 1972: 138; Kim & Shin. 1996: 30.

Hercttltia gtaueinalis: Okamute, 1924: 149; lnoue, 1955: 15l; Mutuura, 1957: 108; Inoue, 1959: 243, pl. 168: 29; Seok, 1970: 145. P.vralis nitidalis Fabricius, 1794, Ent, S.yst, 3 (2): 228. TL: Europc. Pyralis yokohamae Butler. 1879, An.n. Mag. nat, Lfist, (5)4: 452. TL: Japan.

Adult, male genitalia, and female genitalia in Figs 12, 33, 53.

Materials, 69 males and 104 females examined in this study.

Distribution, Korea (GW, GG, CB, GB, GN, JB, JN, JJ), Japan, China, and Europe.

Flight period. May to September.

Remarks. This species seems to be a common and widely distributcd in Korea. It is very

similar to O. nannodes superficially, but it is distinguished from the latter by the almogt straight postmediat line and very short valva,

Orthopygia nannodes (Butler, 1879) (Figs 13, 34, 54)

P),ragis nannodes Butler, 1879, Iltust. t.vv)ical 5Petriinens Lepid. Heterocera CoUn Br. Mbts. 3: 7l , TL: Jupan. Orthop.vgia nannvdes: YuinunHka, 1965: 71; Park, 1976: 17; Inoue, 1982. 1] 383, 2: 247, pl. 46: 16; Park, 1983: 408, 890. pl. 28: 453; Park & Kim 1976: 80; Yamanaka & Yoshiyasu, 1992: 90; Park, 1993: 154; Bac, 2001: 180, fig. 169, Hercutia nannodes: Matsurnura. 1905: 203; Inoue. 1955: l52; Inoue, 1959: 243, p]. 16g: 31.

Adult, male genitalia, and female genitalia in Figs 13, 34, 54.

Materials. 2 males and 4 females examined in this study.

Distribution. KoTea (HB, GG, GB, JB, JN) and Japan.

Flight period, June to September.

Remarks. This species is similar te O. glaucinalis superficially, but it can be distinguished from the latter by the fbllowing characters: ante- and postmedial lines of fOrewing pale dull yellowish, with blackish fuscous; cilia brown; uncus tongue-shaped; valva extremely short; corpus bursae spherical; signum present.

Orthopygiaplacens (Butler 1879) (Figs 14, 35, 55)

Rhodariaplaceits Butlcr, 1879, lllusi. tppi(,at SPecimens Lepid. Hererocera Colln Br. Mcts. 3: 72, TL: Japan. Orrhop}'gia placens: Yamanaka, 1965: 71; Park, 1983: 407, 889, pl. 28: 451; Park & Ju, 1990: 392; Park, )993: 151; Kim, 1999: 21;Bae. 2001: 181, fig. 170. placens: Matsumura, 1905: 203; Shibuya, l927: 96; Inouc, 1955: 152; Mutuurtt, l957: 109; lnoue, 1959i 243, pl. 168: 32; Inoue, ]982, 1: 383, 2: 247, pl. 46: 18.

Adult, male genitalia, and female genitalia in Figs 14, 35, 55,

Materials, 16 males and 27 females examined in this study.

Distribution. Korea (HB, GW, GG, CN, GB, GN, JB, JN, JJ), Japan, and China.

Flight period. July to August.

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62 Bong-Woo LEE aiid Yang-Seop BAE {be・tw

,llii .・i.i, x・,,,,-A"tx E- igi' v- iig i li g`iSi,.x,,, ,,,gi /k, l'(--ipa:xt,-A

1su,", :lt(ii .,&gggee fi-ii-\・i-X t/t .'ttl ss / tt /t ' ''t " 'ttt x .'x - iYS ig ,, N ., x ee./.; ・,, i,i 1 / Yi'.lir tt/t "" i ' ,i'・ K, xte""k[t'::{-....,..-.,'"''itk'.//,, /" t'n> 1 C).Vi 11 1t/ Xg7-f-xtlYLttt, , x .L i l eE //t/. / f' "/' ,x ' ' '" 58 '] "yV' × ivtktw/7 57 59

Figs 54-59. Female genitalia. 54. 0rthtrpygia nannodes (But]er, 1879). 55. 0rth(uJygia placens (Butler, l879). 56. Herculiapelasgalis (Walker, 1859). 57. T}'ebaniaj7avij7'ontalis (Leech. 1879), 58. Sybrida approximans (Leech, 1889). 59. Datanoides.fasciatbts Butler, ]878. Scales: O,5 n-n.

Remarks, This species seems to be common in Korea. It is easily separable from the other species by the follewing characters: ditferent coloratien of fbrewing and hindwing; ante- and postmedial lines ycllow; cilia brightly pale yellow; corpus bursac gourd-shaped.

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Review of Pyralini from Korea 63

60 61

Figs 60-61. genitalia.60. Arippara indicator Walker, 1864, 61, Or>'bina regalis (Leech, 1889)Female. Scales: O.5 mm,

Genus Herculia Walker, 1 8S9

Hlerculia Walker, 1859, List specimens tepid. insects Colln Br. Mus. 18: 546. TS: Hercttlia bra['teratis Walker, 1859. Bojuda Walker, 1865, List Sl)ecitnens lepid. Insects Colln Br. Mus. 34: 1273. TSi Bojuda costigeratis Walker, l866. Bleone Ragonot, 1890, Annlh' Soc. ent. Fr. (6) 10 (Bull.): xcii. TS/ Asopiafi{lvocilialis Duponchel, 1832. Buz-ala Walker. 1863, List ,SPecimens tepid, Insects Coiln Br. Mus. 27: l29. TS: Bu:,alajuscicosta Walkcr, 1863.

Cisse Walker, 1863, List Speeiinens lepid. Insects Colln Br. Mus. 27: 125. TS: Cisse nigrivitta Walker, 1863. Dotichomia Ragonot, 1891,Annls Soc. ent, Fr, 6e: 19 (key), 32, TS: Asopia binode{laiis Zeller, 1872, Lixa Walker, 1865, List hpeciemens tepid. Insects CoUn Br, M"s. 34: 1254. TS: Lixa productalih' Walker, 1866.

Pseudasopia Grote, 1873, Bull, Bqfilalo Soc. nat. Sci. 1: 172. TS] Pseudosopi.a squamealis Grote, 1873. 7'richattchenia WaiTen, 1 892, Ann. A4dg. nat. Hist, (6) 9: 173. TS: E"ctita dharmsatae Butler, 1889. Herculea Amsel, 1956 [indcx], missp.

Herculia pelasgalis (Walker, 1859) (Figs l5, 16, 36, 56)

P.vraiispelasgatis Walker, 1859, List 5?)ecimens t(tpid. insects Coiln Br, Mus, 17: 269. TL: N, China.

Herculia pelasgalis: Matsumura, 1905: 203; Okamoto, 1924: 149: Shibuya, 1927: t7; Inoue, 1955: ISI; Inoue, 1959: 243, pL 168: 34; Mutuura, 1957: 109; Yamanaka, 1968: 130; Zool. Soe, Kor, 1968: 50; Seok. 1970: 146; Inoue. 1982, 1: 383, 2: 247, pl. 46: 25; Park. 1983: 406, 889, pl, 28: 450; Yamanaka & Yoshiyasu, 1992i 90; Bae, 2001/ 174, fig. 16S. H>psop.vgia J'aponica Warren, 1891, Ann. Mag. nat. Hist. (6) 7: 499, TL: Japan, Hlercuiia j'aponica: Hampson, lg96, 7)'ans. ent. Soc, Lond. 1896: 519; Leech. 1901, T)'ans. ent. Soc, Lond. 1901 : 427; Matsumura, 1905: 203; For. Rcs. Kor., 1969: 93. Hlrrculia marthatis Strand, 1919 (not Walker, lg59).

Hlerculia ignij7"alis tangus'ti Strand. 1919, TL: Taiwan.

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64 Bong-Woo LEE and Yang-Seop BAE

Adult, male genitalia, and temale genitalia in Figs 15, 16, 36, 56.

Materials. 40 males and 30 females examined in this study.

and Distribution.Korea(HB,GW, GG, CB, GB, JB,JN,JJ),.lapan, China.

Host plant. et{ercus acutissima C. (Fagaceae) in Korea (Park, 1983).

Flight period. July to August.

Remarks. This is a common species in Korea. The color of the wings x,aries from reddish

brown to dark blackish brown, but it can be distinguished from the other species by the shape of wing, and parallel ante- and postmedia] lines of the hindwing.

Herculia orthogramma Inoue, 1960

Hereulia orthogramma Inoue, t960, Kontydi 28i 170, pl, 1O, figs 1-3; Inoue, I982, 1: 384, 2: 247, pl. 46: 30; Park, 1993: 154; Bae, 2001: 174, fig. 164, TL: Japan.

Distribution. Korea (PN, GW) and Japan,

Remarks. This species was recorded by Park (1993) bascd on the collection of North Korean materials preserved in the Hungarian Natural History Museum, In this study, no specimen of this species was found from to South Korca. According to Park (1993), this species is the largest (wingspan 40 mm) in the genus. According te Inoue (1982), it is vari- able in wing length, The female is larger than 30 mm, but the male is usually small, It ap- pears in autumn in Japan.

Genus 7beebania Ragonot, 1891

n'ehania Ragonot, 1891, Annts Soc. ent. Fr. 60: S46. TS: Propachr)s.flavij>'ontcclis Leech, ]889,

7>'ebaniaflavij},onlalis (Leech, 1889) (Figs 17, 37, 57)

Propach.vsflavij}'ontalis Lccch, 18g9, Entoinologist22: 108. TL: JapanlChina.

Trehania .frat,ijhonlalis/ Matsumura, 1905: 203; Shibuya, 1927: 96; Inoue. 1955: 152; )tlutuura. 1957: 109; Inoue, 1959: 243, pl. 168: 3S; Inoue, 1982, 1: 384, 2: 248, pl, 46: 33; Purk, 1983: 409, 890; Yamanaka & Yoshiyasu. 1992: 90: Bae. 2001/ 195. fig. 180.

Adult, male genitalia, and female genitalia in Figs 17, 37, 57.

Materials. 3 males and 2 females exainined in this study,

Distribution. Korea (HN, GW, GB), Japan, and China.

Flight period. July,

Remarks. This is a rare species in South Korea, Only five specimens were collected in July

throughout the study. This species is easily separable frem the other species by the follow- ing: adult rather large (wingspan 33-35 mm); head reddish orange yellow; ground color of wings dark grayish brown; ante- and postmedial lines absent; uncus long tongue-shaped; valva rather wide; cornutus absent; appendix bursae present.

Genus Sybrida Walker, 1865

Sybrida Walker, 1865, List specientens tepid. Insects Colln Br. Mtts. 32: 465. TS: 5)?brida inordinata Walker, 1865.

Sacada Walker, 1862, J. Proc. Liitn. Soc. (Zool.) 6: 136. TS: decora Wa[ker, 1862,

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65 Rcvicw of Pyralini from Kurea

Sybrida upproximans (Leech, 1889) (Figs 18, 38, 58)

Datanoides approximans Leech, 1889, Proc. :・ ool Soc. Lond. 1889: 636. TL:Japan. S.vbrida approximans: Shibuya, 1927: 96; Inoue, 19S4: 153; Mutuura, 1957: 109; Inoue. 1959r 243, pl. 168: 36; Kor. Soc. Pl. Prot,, 1972: 140; lnoue, 1982, li 384, 2: 284, pl, 46: 34, Sacada approximans: Leech, 1901: 429; Park, 1976: 17; Park, 19g3t 405, 88S, pl. 28: 449; Bae, 2001: 187, fig. 174. Sacada CDataneides) approximans: Matsumura, 1905. : 203.

Adult, male genitalia, and female genitalia in Figs 18, 38, 58.

Materials. 34 males ancl 47 females examined in this study,

Distribution. Korea (HN, GWI GG, CB, GB, GN, JB, JN), Japan, China, Myanmar, and India.

Host plants. QueiTus acutissima C,, Querct-{s acuta T., Castanea crenata S. et Z. (Fagaceae) in Korea, (Park, 1983).

Flight period, May to August,

Remarks, This species seems to be common in Korea. Male and female adults differ in the length of wings and the ground color. It is similar to Datanoidesfasciata, but it can be dis- tinguished by the slightly curved postmedial line, and dark reddish brown or grayish brown ground color,

Genus Datanoides Butler, 1878

Datanoides Butler, 1 878, Entomologist's Jnon, Mag. 14: 206. TS: Datanoidesfasciata Butler, 1878.

Datanoidesfasciatus Butler, 1 878 (Figs 19, 39, 59)

Datanoidesjusciattfs Butler, 1878, Entortzologist's mon. Mag. 14/ 207; Inoue, 1982, 1: 384. 2/ 284, p]s. 46: 36, 37. TL:Japan, S.vbrida fasciata: Inoue, 1955: IS3; Mutuura, 1957: 110: ]noue, 1959: 244, pl. 169: 2; For. Rcs, Inst., 1969: 97; Kor. Soc. Pl. Prot., 1972: 140. Sacada JLisciata: Park, 1983: 405, 8g8, pl. 28: 448; Park & Ju, 1990; Park, 1993: 154: 392; Bae, 2001: 188, tig. 175 Sacada (Datartoides)juseiata: Matsumura, 1905: 203. Trebania fosciata: Zool. soc. Kor,, 196g: 50. Xestttta miractttosa Snellen, 1885, in Romanoff. Me'm. Lopid. 2: l95. TL: Russia

Adult, male genitalia, and female genitalia in Figs 19, 39, 59,

Materials. 21 males and 29 fema]es examined in this study.

Distribution. Korea (HB, HN, GW, GG, CN, GB, GN, JB, GN, JJ), Japan, China, Myanmar, and India.

Host plants, euereus acvttissima C., 2uercus serrata T,, euercus aliena B., and Lespedel.a bicolor Turcz. (Fagaceae) in Korea (Park, 1983),

Flight period, May to September.

Remarks. This species has been known as Sacada fasciata, but was removed to Dananoides by Inoue (l982). It seems to be a common and widely distributed species in Korea, It is similar to S>,hrida cu2proximans, but can be distinguished by the almost straight postmedial line and brownish olive ground color of the fbrewing.

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66 Bong-Woo LEE and YanLg.-Seop BAE

`Cl864") GenusAmppara Wa]ker, 1863 (not

"1864"), AriRpara Walkcr, 1863

Arippara indicator Walker, 1863 (Figs 20, 40, 60)

Arippara indicator Walker, 1863, J. Pro(,. Linn. Sec. (Zool.) 7: 74; Inoue, 1982, 1: 384, 2: 248, pl, 46: 40; Park, 1983: 412, 891, pl. 28: 459; Yamanaka & Yoshiyasu, 1992, Lqpid. 1lriwan, 1 (2): 90. TL: Malaysia(Sarawak). Bostra indicator: Inoue, 1955: 153; Mutuura, 1957: 110; Inoue, 1959: 244, pL 169: 3; Zool, Soc. Kor., 196g: 50; For. Res. Inst,, 1969: 98; Kor. Soc. Pl. Prot., 1972: 140, Bostra marginata Walker, 186S, List Speciemens lepid. Insects Cotln Br. Mtts. 33; 270. Bostra (Poaphila) marginata: Matsumura, 1905: 203, Palea retfescens Butler, 1879, Ann. Ma.o, nat. Hist. (5) 4: 354, TL: Japan. L`1882"), Paredra eogenalis Snellen, 1880 (not in Veth, Midden-Sumatra Lep, 4 (1): 60. TL: Indollcsia (Sumatra). Ryralis assamica Moore, 1888, in Hewitson & Moore, Des(/r. ne}v indian lepid. Inseets Colln late Mr Atkinson: 205. TL: India (Assam). Bostra bip),・ra Strand, 1920, Arch. Nat. 84 (A) (12): 190.

Adu]t, ma]e genitalia, and female genitalia in Figs 20, 40, 60.

Materials. 3 males and 4 females examined in this study,

Distribution. Korea (GW, GG, JN, JJ), Japan, Taiwan, Indonesia, Malaysia, and India.

Host plants, Cinnamomum camphorzz Sieb. (Lauraceae) and Rhus sp. (Anacardiaceae) in Korea (Park, 1983).

Flight period, July.

Remarks. This species is similar to the female of S, approximans superficially, but it can be distinguished by the rounded termen, almost straight antemedial line, and the wide gap be- tween ante- and postmedial lines at dorsum.

Genus 0i ybina Snellen, 1895

Or.vbina Snellen, 1895, 1:ijdschr. Enl. 38: 107. TS: Or),ba.flaviplaga Walker, 1863. Oiyba Walker, 1863, List specimens tepid. insects Colln Br. Mus. 32/ 518. TS: Oi7ba flaviplaga Walker, 1863.

Hira.vamaia Marumo, 1917, in Nagane, Colln Essa.vsfor Mr }?isushi NitTva: 36. TS: Orl,ba regalis Leech 1889,

Or:ybina regalis (Leech, 1889) (Figs 21, 41, 61)

Oi:yba regalis Leech, 1889, Entoni.ologist 22: 71 . TL: Korea. Hirayamaia regalis: Inoue, I955: 153; Mutuura, 1957: 1 1O; Inoue, 1 959: 244, pl. 169: 5; }'lyun & Woo, 1969: 168. O,)rbina regalis: Inoue, 1982, 1: 385, 2: 248, pl. 46: 42; Park, 19g3: 4{}9, 890, pl, 2g: 454; Kjm, et al. 1991: 175; Bae, 2001:]S2, fig, 171,

Adult, male and female in Figs 21, 41 genitalia, genitalia , 61,

Materials. 87 males and 43 females examined in this study.

Distribution, Korea (HN, GW, GG, CB, CN, GB, JN, JJ), Japan, and China,

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Review of Pyralini from Korca 67

Flightperiod, June to September,

Remarks, This species was described by Leech (1889) from the type locality, WOnsan, N. Korea. This is a common species in Korea. It is easily distinguished from other species by the large yellow spot on the forewing,

Acknowledgements

We express thanks to Prof, K. T. Park of CIS for his helpful comments and fbr giving us an opportunity to examine valuable materials. Wb wish also to express our cordial thanks to

Prof. T, Hirowatari and Prof. M. Ishii of Osaka Prefecture University, and Mr. H. Ybshimoto, [Ibkyo High School, Japan, for their valuable suggestions. For the generous loan of materials, our cordial thanks to Dr G. S, Lee (NIAST), Dr B. K. Byun, Korea National Arboretum, and to Drs M, K, Paek, N, H. Ahn, and Mr Yl K. Kim for their assis- tance in collecting the materials. This study was supported by a grant of the University of Incheon in the year ef 2006 (Title: Systematic study of the tribe Pyralini Latreilie (Lepidoptera, Pyralidae, Pyralinae) in Korea).

References

AmseL G. H., 1956. Microlepidopteru venezolana I. Boletin Ent. V2,nezolana 10: 1-336. Bae, Y. S.. 2001. : Pyraustinae and Pyralinae. Ecenomic Insects of Kerea 9. Insect kore- ana (Suppl.) 16: 251 pp. Hyun, J. S. and K. S. Woo, 1969, Insect fauna of Mt. Jiri (I), Bull. Seoul NLitional (hiiv. Forests 6: 171-173. Inoue, H., 1955. Pyralidae. in Inoue, H, (Ed,), Check List ofthe Lepidopteizi qfJapan 2: 120-196. Tokyo,

et al,, nat"rc;li , 1959.Pyralidae,in Inoue,H. Jconogrcrphiainsac:torummponicorumColore Edita 1: 232-257. pls 165-]73. Hokuryukan, Tokyo. et al, ,1982. Pyralidae.Jn Inoue,H. (Eds),Mbths ofJapan 1: 307-404, 2:223-253. Kodanshu, Tokyo.

, 1985.A new specics and synonymic notes on two taxa of the PyralinaefromJapan (Lepidoptera; Pyra]idae). Akitti (N. S.) 72: 1-4, Kim S, S., 1999. Moths collected from Is Ulleung-do, Korea. J. Iepid. Soc. Kbrea 11: 28-29, Kim S. S. and Y, H. Shin, 1996. Notes on eleven unrecorded and nineteen littlc known geomenid moths from Korea. J, lqt)id. Soc. Koi'ea9: 15-25, Kor. Soc, Pl, Prot., 1972. A List qfPlant Diseases, insect Pests, and i7beds in Korea: 137-140. Leech, J. H., 1889, New species of Deltoides and R}'ralis from Corca, North China and Japan. Entomologists 22: 62-71, pLs 2-4, 1901. Lepidoptera , HeteroceTat'romChina, Japan and Corea. 7'rans,enr. Soc. Lond, 1901: 417- 432. Maruda, S,, 1929. Survey of noctuinal t]yin.o insects. Ann. agric, eup. Stn Gov,-gen. Chosen 4 (6): 3I3-376, Matsumura, S., 1905. Cataloguff inh'ectorum JQponieum. 3, 307 pp, Keiseisha, Tokyo. 1925. An enumeratjon , of the butterfliesand moths from Saghalien,",ith descripLionsof new species and subspecies. J, Coll. Agric. Hokkaido lntp. Univ. 15: 83-196, pls. 8-11. Munroe, E. and M. A, Solis, 1998. The Pyraloidea. in Kristensen, N. P. (Ed.), Lepidoptera, moths and but- terflies, yol. 1: , systematies and biogeography. Handb. Z)ol. Berl, 4 (35): 233-256, Mutuura, A., 1957, Pyralidae, JJz Esaki, T. et al., k'ones Heterocerorum Japonicorum in Coioribus NintLtralibus 1: 108-145, pls 17-26. Hoikusha, Osuka. Nakayama, S. and H, Okamoto, 1940. List of fruits insect pests in Korea. Ann. agric. exp. Stn, Gov.-gen. Chosen 12 (3): l95-247, Okamoto, H., 1924. Insect faunaofQuclpart Island. Ann. agric. exp, Stn Gov, Chosen 1(2): 95-152. Park, K, T,, l976. 0n fifty-stx unrecorded species of Pyralidae (Lep{doptera) from Korea, Korean J. Ent, 6 C2)i 11-21.

et at. , 1983. Pyralidae.In Shin.Y. H. (Eds),Insecta(IX),IUustfzitedFtora & Fa"na ofKof'ea27: 29g-444, pls 1 8-30. Ministry of Education.

and , 1993. Pyralidae Thyridjdae (Lepidoptera)from North Korea. Korean ,L mpl. Ent. 32: 151-167.

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68 Bong−Woo LEE and Yang−Seop BAE ・ Park, K , T . and S. Y . Ju,1990. The insect fa巳na of Ml . Samak , J.∫‘ i. Tech. Kangwea.n natn .σ〃 iv.29: 392−393 . Seok, D ,M .,1970.1加 伽 覦 加 襯 α (ゾthe ls. Cttelpart.186 pp, Bojinje Co . ’ Shibuya, J,,1927. Some new and unrecorded species of Pyralidae from Cerea 〔Lepldoptera).加 、secta matsttm , − 2 : 87 1〔〕2, ’ ,1928.Thc systcmatic study on the Formosan Pyralidae.ノ:Facttl. Agric. ft「okkaido intJ).(Jniv.17 − (D : 107 138. − SoLis, M A .跏 d M . Shaffer,1999, Contribution towards the study of the Pyralinae(Pyralidae): historical re vicw , morphology , and nolnenclature .ノ、ム epid . Soc 、53 : 1−10. − − − ’ Strand, E ,1911 1922.∬n Seitz(Ed .),1908 1934. C 厂o ∬ Schme 〃ε 厂〜inge der Erd(デ〃 α (,rolepidol )tera ρ1 洗召 − World . Vol .10 : 1−4,292 −387,289−290 ,422 425 ,436 −442 .

− − − Wang , P . Y ,,1980. Lepidoptera: Pyrulidac. Econon7.iclnsectFat‘na of China 21:i xii ,1 229, pls l 32. ー Yamanaka , H .,1965. Revised classificaUoll of the 亅apanese PyraIinae, especially l〕ased on thc malc and fヒ − − nlale genitalia(1). Tinea 7: 67 76, Pls l O 13, Yamanaka H and Y Yoghiyasu 1992 Pyralidae 加 Heppner J B and H Inoue Eds Lepid‘) コtera , . . , . . ,, . , ( ), 1 qf Taiivan l (2):77−95. Taiwan,

ZI}ol Soc KoL 1968 1V翩 ina Animaliu 〃t Koreanorum 2 lnsecta:49−53 . . , . ( ), .

摘 要

・ 韓 国産 シ マ メ イ ガ 族 の 再 検討 〔李 峰雨 裴 良燮)

韓 国 産 の シ マ メ イ ガ 族 Pyralilliを 再 し た . で は こ れ ま で に 17 の の メ イ ガ が さ ( ) 検討 ’韓国 , 種 本族 記 録 れ て い る が ,今 回 新 た に 助塑 ∫qρ}・g’α iwafnotoi Kirpichnikova & Yamanaka , H 漉 α 贈 わ♂ Y とmlallaka ウ ス モ ン マ ル バ シ マ メ イ ガ , Bostra nanalis (Wilcman)ヒ メ ア カ シ マ メ ・イ ガ の 3種 が 追加 さ れ た .こ れ ら 3種 に ー つ い て は ,雌 雄 交尾 器 を含 め 訂記載 した.ま た ,ハ ン バ リ 隊 の 標 本 に基 づ い て 北 朝 鮮 か ら記録 され

た Herculia orthogramma lnoue バ シ マ メ ガ は は と っ い , で は オ オ イ , 韓 国か ら 依然未発見 な て る 本報 , 本種 を除 く20 種 に つ い て 成虫 の 全 形図な ら び に 雌雄交尾.器図 を提出 し ,国内分布 もま とめ た.

[文責 : 古本 浩IHiroshi Yoshimoto ]

(Accepted August 12,2006)

Published by the Lepidopterological Society〔,f Japan, − − 5 20, Motoyokoyama 2, Hachi〔’ji, Tokyo,192 0063 Japan

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