The malacologicalsocietymalacological society of Japan

.Rre VENUS CJap. Jour. Malac.) Vol. 49, No, 4 (1990):271-279

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Descriptions of New Mollusks from R6union Island II.

Cerithidae, Lamellariidae, Buecinidae, , and Veneridae

Jean DRIvAs and Maurice JAY

(50 rue Karl Marx, Sainte Therbse, 97419 Possession, Reunion, and 46 rue

Eugbne Dayot, 97434 Saint-Gilles les Bains, Reunion, Franee)

Description of New Taxa

Family CERITHIDAE FIeming, 1822

Cerithizam veitlardi n. sp.

(Fig. 1)

Description: Shell with a high spire. Protoconch of 31iS smooth whorls. Teleo- conch of 16 whorls, The earliest 3 whorls bear 2 main spiral eords, and a weaker

one just above suture; on the following whorls, 3 rnain spiral cords, secondary

spiral cords appear below the lowest main one; 14 spiral eords on last wherls,

Axial seulpture consists of strong axial ribs, numbering 10 to 11 per whorl, 'that strongly protruded at the level of the upper and lower main spiral cords, are tuberculated at crossing peints. One varix at each whorl, opposite to outer

lip on the last whorl. Very fine spiral cords and very fine irregular axial riblets

covering the main sculpture visible under mierescope. Interior canal recurved

and truncated. Aperture oval, outer lip denticulated; eolumella with a single

felcl at its upper part. Operculum eorneous and thin. Colour orange-brown,

spiral eords and tubereles creamy; protoconeh, columella, inside of aperture, and

a few tubereles white.

Di7n-ensions: Holetype 25.4mm in length, 9mm in width. Adults from 24 to

27 mm length.

Depository: Holotype in the Museum National d'I{istoire Naturelle, Paris,

Paratypes 1 to 25 in authors' eollection.

"Souris-ehaude'', T・ype locality: Off between La Saline and St. Leu, west coast

of R6union island, found alive in white coral sand, under big stones, at the depth

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272 VENUS: Vol. 49, No.4(1990)

of 50-75 m.

Etymolog・y: Named in honour of Mr. Mauriee Veillard, a well-known collector

on Reunion.

Remarks: This species eould be compared with Rhinoclavis icochi (Philippi,

1846), from whieh it differs by the lack of a columellar fold, less reeurved

anterier canal, more ventrieose shape, and details of sculpture.

Family LAMELLARIIDAE d'Orbigny, 1841

(;briocelta herberti n. sp. (Figs. 2-5)

Description: Shell internal, thin and fragile, ear-shaped, white and translucent,

covered with a very thin brownish periostracum. Very fiattened spire of 2Y.,

smooth whorls. Aperture wide and ellipsoidaL The dorsal face of is

occupied by the mantle, which is wide and oval. The foot, visib]e on the ventral

face, is small eompared with the mantle. Two elongate flat tentacles on the

head. Colour varies from lemon-yellow to b]ack, but all specimens bear a white

transverse band, white short bands next to the edges, and white fine dots on

the whole surface; tentacles of the same eolour as the mantle, with white

summits, Male duct in body wall. Radula formula O.1.R.1.0. Central tooth

delta shaped, lateral teeth with a single cusp.

Dimensions: Holotype: Diameter of retraeted animal, preserved in aleohol,

10mm Shell length 8mm, height 6mm,

Depository: Holotype in the Museum National d'Histoire Naturelle, Paris.

Paratype 1 in Natal Museum. Paratypes 2-5 in the authors' collection.

Type locality: Cape La Houssaye, Saint-Paul, west coast of R6union Island,

at the depth of 12m, found creeping under small stones. All the 7 known

specimens from the same locality.

Etymoeog・y: Named for Dr. Dai Herbert, Natal Museum, who has taken the

SEM photos for this paper.

Remarics: This new species differs from Coriocella bergh・ii (Deshayes, 1863), described from Reunion, and synonym of C. nigra (Blainville, 1824), by the

animal being much smaller, more flattened body, and the lack of dersal tubercles,

Family BUCCINIDAE Rafinesque, 1815

Siphonofusus bl(ntneri n.''sp.

(Fig. 6)

De$cription: Shell fusiform in shape, spire slightly shorter than total length

of aperture and anterior canal. Protoconch of 2 smooth whorls, apieal ang]e

23e. Teleoconeh of 8 eonvex whorls bearing rounded axial plieae, more neatly

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Drivas and Jay: New Mollusks frorn R6union-II 273

marked on earlier wherls, disappearing on body whorl, numbering 7 on penulti-

mate whorl. Main spiral cords 11 on penultimate whorl, with seeondary weaker

cords, one in each interval, and 4 above the higher main eord. Numerous very

fine undulating axial riblets overeross the cords giving the shell a finely granulate

appearance. Siphonal canal large, slightly incurved. Columella eallus smooth.

Outer Iip of aperture thin, weakly denticulate, bearing small brown lines

eorresponding to the tinted spiral cerds of outer surfaee. More deeply inside

aperture present 14 lirae. Celor golden brown, with a subsutural spiral band, with the darker main spiral eords. Protoconch paler. Inside of aperture white,

Dimensions: Holotype height 54mm, breadth 18mm. Paratype 1 height

77,5mm, breadth 21.3mm. Paratype 2 height 62.2mm, breadth 21,3mm.

Paratype 3 height 61.5mm, breadth 20.8mm. Paratype 4 height 46.6mm,

bradth 16.2mm.

Depository: Holotype in the Museum d'Histeire Naturelle, Paris; all paratypes

in the authors' colleetions.

Type locatity: St. Paul's Bay, west eoast of R6union island, at the depth of

750 m. Found dead in shrimp-pots. All specimens frorn the same area.

Etymology: Nained for Mr. Hugh Bradner, Scripps Institution of Oceano-

graphy, University of California, San Diego, whom we thank for his aid for literatures.

Remarks: This spec'ies may be eompared with Siphenofusus chTysodomoides

(Sehepman, 1911) from whieh it differs by smaller size, regularly cenvex whorls

and lack of a stronger spiral cord at shoulder angle, and possession of a single

secondary weaker cord in each interval instead of 2 er 3.

Family MITRIDAE Swainson, 1831

Mitra rmphaeli n. sp.

(Figs. 7-8)

Description: Shell fusiform, elongate ovate in shape, solid and heavy. Pro-

toconeh unknown. Teleoconch of 9 whorls, the earlier 2 of them broken on

paratype. FIattened spiral cords 6 on penultimate whorl, about 22 on body whorl,

and 6 finer ones on base. Very fine, closely-set axial riblets, above 100 on last

whorl, and better visible in the intervals between the cords than over them.

Aperture oval and elongate, equal in height to spire. Outer lip thick, interior

smooth, Columella with 5 folds. Siphonal canal slightly recurved. Colour

creamy white, with brown blotches irregularly spaced on spiral cords; a spiral

cord at the middle of bedy whorl does bear no blotch.

Dimensions. Holotype height 30,7mm, breadth 9.6mm. Paratype height

55.6 mm, breadth 17.8 mm.

Depository: Holotype in the Museum National d'Histoire Naturelle, Paris.

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274 VENUS: Vol. 49, No.4(1990)

Paratype in the authors' collection.

``Souris-Chaude", Type locality: near Saint Leu, in the western part ef R6union

island, at the depth of 66 m. Beth specimens from this area.

Etymology: Named for Mr. Raphael Drivas, son of one of the authors.

RemaTks: This new species differs from Mz'tra maui Kay, 1979, by having

6 spiral eords instead of 7, the axial riblets overerossing the cords, and by

color pattern. The brown blotches on cords are not alined axially, except on the

juvenile, and are always less neatly marked under suture than on lower whorls.

It may be eempared with M. cardinalis (Gmelin, 1791), but the shape of the

latter is different, with a shorter spire and wider body whorl, bearing more

numerous axial ribs, and the brown blotches extending over several cords. These

ehaTacters are valuab]e to separate juveniles of these twe species, that are at a

first glanee, nearer than adults. Juvenile of this species may be eompared with

Ziba rehderi (Webb, 1959), but differs from it by shape and color pattern.

(;tincitla () salisburyi n. sp.

(Figs. 9-10)

Description: Shell fusiform in shape; height of aperture equal to spire. Pro-

toeoneh of 4 slightly convex smooth brown whorls, Teleoconch of 7 fiattened

whorls with a weakly impressed suture, Spiral sculpture made of slightly

rounded spiral eords, numbering 4 on whorls; the second cord under suture

slightly stronger than others, and the lowest one slightly weaker, Intervals

between cords about 3 times wider than cords, bearing a weak spiral cord in

the middle, and numerous axial riblets that never reaeh the summit of eords;

the interval between the secend and the third eords slightly wider than others,

and secondary spiral eord slightly stronger, espeeially on the last whorl. Aperture

narrow and elongate, pointed at the base. Columella with four oblique folds.

Outer Iip thin and faintly crenulated. Siphonal eanal nearly straight. Colour

cream, with brown blotches on the main cords, reaching the neighbouring parts

of interva]s, and sometimes confiuent into elQngate brown blotches,

Dimensions: Holotype height 15.3mm, breadth 4,8mm; Paratype specimens

1 to 22: from 12.2 × 4mm to 20.5 × 6.2mm.

Depository: Holotype in the Museum National d'Histoire Naturelle, Paris.

Paratypes 1 to 20 in the authors' collections. Paratype 21 in the Natal Museum.

Paratype 22 in Riehard Salisbury's collection.

"La Type locatity: Possession" in the western part of R6union Island, at

the depth of 54m, where this species was found alive in muddy black sand.

More dead speeimens frem Saint Paul's Bay, western part of the island, at

the depth of 24 m.

Etymology: Named for Mr. Riehard Salisbury, for appreeiation of his help.

Remarks: This speeies is easily separated from Subcancilea flammigera

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Drivas and Jay: New Mollusks from R6union-II 275

(Reeve, 1844) and Subcancitla annulata (Reeve, 1844), that have definitely

strogner cords, numbered only three per whor]; from Ziba intersc'ulpta (Sowerby,

1870) which has three main eords per whorl and a different colour pattern; from Neocancitla circula (Kiener, 1839) whieh has a more elevated spire; from juvenile NeocanciLla MaTis (Linn6, 1771), of which the axial furrows overcut the spiral eords; from (Domiporta) interlirata (Reeve, 1844) forma pia (Dohrn, 1861) which bears six coTds on penultimate whorl; and from

Caneilla (DomipoTta) rujitirata (Adams & Reeve, 1850) which has a shorter spire and only three spiral cords per whorl.

Family VENERIDAE Rafinesque, 1815

(;outclia (orenocirce) kilburni n. sp.

(Figs. 11-12)

Deseription: Shell quadrangular round in shape; umbo net elevated, and

situated about at va of the ]ength from anterior end. Posterior dorsal margin

straight and nearly parallel to ventral margin; anterior dorsal margin concave;

posterior margin vertieal. Ligament external pale brown. Hinge with three

divergent cardinal teeth, and strong anterior tooth. Inner margin crenulated.

Sculptured on the surface by elosely-set, flattened somewhat gemmulate, and

ceneentric lirae, and weak radial lirae, visible on anterior and posterior parts.

This sculpture is more evident and more extended on juvenile specimens.

Colour pattern of angulose, irregular gray and milky-white blotches, limited

to the posterior area, on a ereamy white back ground; umbonal area paler.

Interior is yellowish on fresh specimens. 'Dimensions: Holotype height 4.8mm, length 5.5mm, breadth 3.lmrn; Para-

type 1 height 4 mm, length 4.7 mm, breadth 2.5 mm; Paratype 2 height 5.8 mm,

length 7mm, breadth 4mm.

Depository: Holotype (a eomplete shell) in the Museum National d'Histoire Naturelle, Paris, paratype (a complete shell) in Natal Museum, paratypes 2 to

20 in the autohrs' collections.

Type locatity: Cape La Houssaye St. Paul, west coast of Reunion Island, at

the depth of 12m. Other specimens were collected off this area, to the depth

of 50 m; commonly found dead, and only rarely alive.

Etymotogy: Named for Dr. Richard Kilburn, Natal Museum,

Remarks: This species may be compared with Gouldia cooki (Dall, Bartsch &

Rehder, 1938), from which it differs by its shape, namely longer than high

while G. cooki being higher than long, and having rnuch stronger axial seulpture.

Pitur roseoIJrotlisso(x}nchus n. sp.

DescTiption: Shell rather thin, subtrigonal and round in shape, with elevated

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276 VENUS : Vo1 ,49, No .4 (1990)

straigh 七 umbones ,situated justanterior 七〇 the middle ; anterior dorsal 皿 argin ,

posterior dorsal margin concave ; rest of margin round except for angulate

an 七erior part . Lunule slightly concave . Ligamen 七 external and pale brown ,

one simple Three divergent cardinal teeth on each valve ,the Ieft anterior being ,

righ 七 bifid one Iateral too 七h. Inner margin thick, the anterior one , anterior

and Surface sculptured with very weak irregular concentric lirae, deep irregular

coneentric growth furrows . No radial sculpture . The glossy outer surface is

white 七 covered to a greater or lesserdegree ,by a soft chalky deposit, tha is

easily removed by nails .Colour pattern with irregular , pale fawn blotehes, oll − amilky white background . Prodissoconch with a vivid pink flammule anteriorly ,

with the second paler one posteriorly.

: and maximum size single right valve Dimensions Holotype ( ), , height

. mm . mm . right single valve 15 7mm , 1ength 18 , breadth 5 7 Paratype 1 ( ),

mm mm right single height 14 .5 , length 17 , breadth 5.1mm . Paratype 2 (

valve mm mm mm ), height 15 .7 , length 13 .7 , breadth 4.3 . Paratype 3 (left

single valve mm 七 mm ), height 12 .6 , length 14 .6mm , bread h 4,3 . ’ Depositorty : Holotype and paratype l in the Museum National d His 七〇ire

Nature]le, Paris , Paratypes 2 and 3 in Natal Museum . Paratypes 4 to 20 in

, authors col ]ection . ・ 丁 610e α : at the of m off aye − 鯉 1ity Found dead depth 12 , Cape La Housg. ,Saint

west as a nor alive Paul , coast of R6union Island。Never found complete .g hell, .

All specimens from the same locality.

Etymology : Named after the celour of its prodissoconch .

αna Remarles: The nearset species is Lioconchα torenzi (Dillwyn , 1817 ) from

which it differs by its colour pattern , and pink blotch on prodissoconch .

要 約

レ ユ= ソ 二 オ 島 か ら 腹足 類 5 新 種 , 枚 只 類 2 新種 を 記 載す る 。

ニノ ノ ン ル ー Cerithium veittardi n .sp .(オ ツ ガ イ 科) 殻 長 25.4mm 。 島 西 岸 の ラ サ リ ネ と サ の . 聞,水深 50〜75m サ ン ゴ 砂 中。 . ベ コ タ マ の サ ン ール Ceriocella herberti n ,sp .( ッ ウ ガ イ 科) 殻 長 8mm ,殻 高 6mm 。 島 西 岸 ポ ,

ル ウ ッ セ 岬 ,水 深 12m 。 ー Siphonofusus bradneri n . sp .(エ ゾ バ イ 科) 殻長 54 mm 。サ ン ポ ル 湾,水深 750 m 。死殻。

フ . サ ソ ル ス シ ー Mitra raPhaeli n .sp ,( デ ガ イ 科 ) 殻長 30,7mm 。 ウ 近 く の リ ョ ,水深 66 m 。

7 ゼ シ ン Cancilla (Domiporta ) salisburyi n .sp .( デ ガ イ 科) 殻 長 15.3mm 。 ラ ポ ョ ,水深 54 m 。 黒 色砂.泥。 ’ 一e ル ス レ サ ソ ール Gouldia (Crenocirce) hilburni n .sp .( ダ ガ イ 科 ) 殻 長 4.8mm 。 ポ , ラ ウ ッ

セ 。水 深 12m 。死 殻 が 多 い 。

マ ル ス レ . 皿1m ン ー ル セ Pitar roseoProdissoconchus n .sp .( ダ ガ イ 科) 殻 長 15.7 。 サ ポ 、 ラ ウ ッ 岬 ,水 深 12m 。死 殻 。 References

Adams , A . and Reeve , L . A .1848 −1850 . The Zoelogy o / the Voy α ge of H 、M .S .

一 NII-ElectronicN 工 工 Eleotronio Library Service The malacologicalsocietymalacological society of Japan

Drivas and Jay: New Mollusks from REunion-II 277

Samavang, under th・e Command of Capta・in Sir Edward Belcher. London, pt. 1-3,

1-87, pls. 1-24.

Blainville, H.M.D. 1824. Dictionnaire des Sciences naturelles; Vers et Zoophytes.

Dictionnaire des Sciences naturelles Pt. 2 Regne organis6 32: 1-392 (not seen).

Dall, W. H., Bartsch, P. and Rehder, H. P. 1948. A Manual of the Recent and F'ossil

Marine Pelecypod Mollusks of the Hawaiian Islands. Bernice P, BishQp Mus.

BulL, (153>: 1-233, pls. 1-53,

Deshayes, M.G. 1863. Catalogue des Mollusques de Vile de la Reunion, in Netes sur

Vile de ta Reunion, annexe E, Parie, Dentu, 1-144, 13 pls.

Dillwyn, L.W. 1817. A desert-ptive Catalogue of Recent SheUs. Vol. 2: 1-1092

(not seen).

Gmelin, J,F. 1791. Systema naturae per regna tria naturae. Editio decima tertia, aucta・, reformata Leipzig, 1(6): 3021-3910 (not seen).

Kay, E.A. 1979. Hawaiian Marine Shells. Bishop Museum, 64(4): 1-654, 135 text- figs.

Kiener, L.C. 1839-1841. Species gefteral et Ieonographie des Coquilles vivatztes: Mitra et Voluta. Rousseau, Paris, 3: 1-120, pls. 1-34.

Reeve, L.A. 1844-1845. Monograph of the Mitra. Cenchologia Ieonica. 10:

pls. 1-39.

Schepman, M.M. 1911. The Prosobranehia of the Siboga Expedition. Rtisz{ltats

Siboga Expeditie; Mon. 49(1), Pt. 4: 1-452, pls. 1-30.

Sowerby, G.B. 1870. Deseriptions of fourty-eight new species of shells. Proc. Zool. ' Soc. Lond. 249-259, pls.21-22.

Webb, J.H. 1958. A new Mitra from Japan. Venus, Jap. J. Malac.. 20(1): 30-31, text-figs.

[Reeeived: June 11, 1990]

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278 VENUS: Vol. 49, No. 4 (1990)

PIate 1. New Mollusks from R6union Island

L2-5.Cerithium veiltardi n, sp. holetype, 25.4 × 9 mm.

CorioceUa herberti n. sp.

2. Shell, paratype No. 2, 8× 6mm ; 3. Animal, paratype No. 2, diam. 26 mm ;

4. Radula, paratype, No, 1; 5. Jaw plate, paratype No, 1.

6. Siphonqft{sus brandneri n, sp., holotype, 54 × 18 rnm.

7-8. Mitra raPhaeli n. sp.

7, Holotype, 30.7 × 9.6rnm; 8. Paratype, 55.6 × 17,8mm,

9-10. Cancilla (DomiPerta) salisburpa' n. sp.

9. Holotype, 15. 3 × 4, 8 mm ; 10, Paratype, No. 4, 17.3 × 5, 3mm(photographed

under amrnonium chloride). 11-12.13-14.Gouldia (Crenocirce) kiiburni n. sp. Holotype, 4.8 × 5.5 × 3,1mm.

Pitar roseoProdissoconchus n, sp.

13. Holotype, 15.7 × 18× 5,7mm; 14, Paratype, 14.5× 17 × 5,1mm.

.

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Drivas and Jay: NewMollusks fromR6union-II 279

Plate 1.

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