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© CSIRO 2005 10.1071/IS04020 AC 1445-5226 Invertebrate Systematics, 2005, 19(2), 113–143
© CSIRO 2005 10.1071/IS04020_AC 1445-5226 Invertebrate Systematics, 2005, 19(2), 113–143. When and where did troidine butterflies (Lepidoptera : Papilionidae) evolve? Phylogenetic and biogeographic evidence suggests an origin in remnant Gondwana in the Late Cretaceous Michael F. BrabyA,B,D, John W. H. TruemanA and Rod EastwoodB,C ASchool of Botany and Zoology, The Australian National University, Canberra, ACT 0200, Australia. BMuseum of Comparative Zoology, Harvard University, 26 Oxford St, Cambridge, MA 02138, USA. CAustralian School of Environmental Studies, Griffith University, Nathan, Queensland 4111, Australia. DCorresponding author. Email: [email protected] The age, geographic origin and time of major radiation of the butterflies (Hesperioidea + Papilionoidea + Hedyloidea) are largely unknown. The general modern view is that butterflies arose during the Late Jurassic/Cretaceous in the southern hemisphere (southern Pangea/Gondwana before continental breakup), but this is not universally accepted, and is a best guess based largely on circumstantial evidence. The extreme paucity of fossils and lack of modern, robust, higher-level phylogenies of extant monophyletic groups have been major impediments towards determining reliable estimates of either their age or geographic origin. Here we present a phylogenetic and historical biogeographic analysis of a higher butterfly taxon, the swallowtail tribe Troidini. We analysed molecular data for three protein-encoding genes, mitochondrial ND5 and COI–COII, and nuclear EF–1α, both separately and in combination using maximum parsimony (with and without character weighting and transition/transversion weighting), maximum likelihood and Bayesian methods. Our sample included representatives of all 10 genera of Troidini and distant ingroup taxa (Baroniinae, Parnassiinae, Graphiini, Papilionini), with Pieridae as outgroup. -
Pmsp for Oregon and Washington Blueberries
PMSP FOR OREGON AND WASHINGTON BLUEBERRIES SOURCE Title PMSP for Oregon and Washington Blueberries Report URL https://ipmdata.ipmcenters.org/documents/pmsps/ORWABlueberry2004.pdf Type Pest Management Strategic Plan Source Date 03/24/2004 Workshop Date Settings Blueberry Region Western States Oregon, Washington Contacts EXECUTIVE SUMMARY Key Pests Insects Aphid (Ericaphis fimbriata) Black vine weevil (Otiorhynchus sulcatus) Cherry fruitworm (Grapholita packardi) Garden symphylan (Scutigerella immaculata) Green peach aphid (Myzus persicae) Leafrollers and olethreutine moths ( ) Lecanium scale (Lecanium spp.) Obliquebanded leafroller (oblr) (Choristoneura rosaceana) Obscure root weevil (Sciopithes obscurus) Orange tortrix (Argyrotaenia franciscana) Root weevils ( ) Rough strawberry root weevil (Otiorhynchus rugosostriatus) Strawberry root weevil (Otiorhynchus ovatus) Winter moth ( ) Pathogens Anthracnose ripe rot ( ) Avocado root rot (Phytophthora cinnamomi) Bacterial blight ( ) Bacterial brown spot of bean/canker of stone fruit, twig blight (Pseudomonas syringae pv. syringae) Bacterial canker (Clavibacter michiganensis pv. michiganensis) Godronia canker (Godronia cassandrae) Phytophthora root rot ( ) Scorch virus (Carlavirus BlSV) Shock virus (Ilarvirus B1ShV) Tomato ring spot virus (Nepovirus ToRSV) Weeds Barnyardgrass (Echinochloa crus-galli) Black nightshade (Solanum nigrum) Buckhorn plantain (Plantago lanceolata) Canada thistle (Cirsium arvense) Cane fruit (blackberry, raspberry) (Rubus spp.) Chickweed (Stellaria media) Clovers ( ) -
The 2014 Golden Gate National Parks Bioblitz - Data Management and the Event Species List Achieving a Quality Dataset from a Large Scale Event
National Park Service U.S. Department of the Interior Natural Resource Stewardship and Science The 2014 Golden Gate National Parks BioBlitz - Data Management and the Event Species List Achieving a Quality Dataset from a Large Scale Event Natural Resource Report NPS/GOGA/NRR—2016/1147 ON THIS PAGE Photograph of BioBlitz participants conducting data entry into iNaturalist. Photograph courtesy of the National Park Service. ON THE COVER Photograph of BioBlitz participants collecting aquatic species data in the Presidio of San Francisco. Photograph courtesy of National Park Service. The 2014 Golden Gate National Parks BioBlitz - Data Management and the Event Species List Achieving a Quality Dataset from a Large Scale Event Natural Resource Report NPS/GOGA/NRR—2016/1147 Elizabeth Edson1, Michelle O’Herron1, Alison Forrestel2, Daniel George3 1Golden Gate Parks Conservancy Building 201 Fort Mason San Francisco, CA 94129 2National Park Service. Golden Gate National Recreation Area Fort Cronkhite, Bldg. 1061 Sausalito, CA 94965 3National Park Service. San Francisco Bay Area Network Inventory & Monitoring Program Manager Fort Cronkhite, Bldg. 1063 Sausalito, CA 94965 March 2016 U.S. Department of the Interior National Park Service Natural Resource Stewardship and Science Fort Collins, Colorado The National Park Service, Natural Resource Stewardship and Science office in Fort Collins, Colorado, publishes a range of reports that address natural resource topics. These reports are of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, conservation and environmental constituencies, and the public. The Natural Resource Report Series is used to disseminate comprehensive information and analysis about natural resources and related topics concerning lands managed by the National Park Service. -
Lepidoptera of North America 5
Lepidoptera of North America 5. Contributions to the Knowledge of Southern West Virginia Lepidoptera Contributions of the C.P. Gillette Museum of Arthropod Diversity Colorado State University Lepidoptera of North America 5. Contributions to the Knowledge of Southern West Virginia Lepidoptera by Valerio Albu, 1411 E. Sweetbriar Drive Fresno, CA 93720 and Eric Metzler, 1241 Kildale Square North Columbus, OH 43229 April 30, 2004 Contributions of the C.P. Gillette Museum of Arthropod Diversity Colorado State University Cover illustration: Blueberry Sphinx (Paonias astylus (Drury)], an eastern endemic. Photo by Valeriu Albu. ISBN 1084-8819 This publication and others in the series may be ordered from the C.P. Gillette Museum of Arthropod Diversity, Department of Bioagricultural Sciences and Pest Management Colorado State University, Fort Collins, CO 80523 Abstract A list of 1531 species ofLepidoptera is presented, collected over 15 years (1988 to 2002), in eleven southern West Virginia counties. A variety of collecting methods was used, including netting, light attracting, light trapping and pheromone trapping. The specimens were identified by the currently available pictorial sources and determination keys. Many were also sent to specialists for confirmation or identification. The majority of the data was from Kanawha County, reflecting the area of more intensive sampling effort by the senior author. This imbalance of data between Kanawha County and other counties should even out with further sampling of the area. Key Words: Appalachian Mountains, -
Lizard Predation on Tropical Butterflies
Journal of the Lepidopterists' Societ!J 36(2), 1982, 148-152 LIZARD PREDATION ON TROPICAL BUTTERFLIES PAUL R EHRLICH AND ANNE H, EHRLICH Deparhnent of Biological Sciences, Stanford University, Stanford, California 94305 ABSTRACT. Iguanid lizards at Iguac;u Falls, Brazil appear to make butterflies a major component of their diets. They both stalk sitting individuals and leap into the air to capture ones in flight. Butterfly species seem to be attacked differentially. These observations support the widespread assumption that lizards can be involved as selec tive agents in the evolution of butterfly color patterns and behavior. Butterflies have been prominent in the development of ideas about protective and warning coloration and mimicry (e.g., Cott, 1940; J. Brower, 1958; M. Rothschild, 1972), and the dynamics of natural pop ulations (Ford & Ford, 1930; Ehrlich et aI., 1975). In spite of the crucial role that predation on adults must play in evolution of defen sive coloration and may play in population dynamics, there is re markably little information on predation on adult butterflies in nature. This lack is all the more striking, considering the large numbers of people who collect butterflies and the abundant indirect evidence from bird beak and lizard jaw marks on butterfly wings (e.g., Carpen ter, 1937; Shapiro, 1974) that adult butterflies are quite frequently attacked. Published field observations of predation on butterflies deal almost exclusively with the attacks of birds and consist largely of accounts of individual attacks (Fryer, 1913). Observations of natural predation by lizards are very rare, although "birds and lizards have long been considered to be the major selective agents responsible for the ex treme diversity of unpalatable and mimetic forms of butterflies in nature" (Boyden, 1976). -
Alfred Russel Wallace and the Darwinian Species Concept
Gayana 73(2): Suplemento, 2009 ISSN 0717-652X ALFRED RUSSEL WALLACE AND THE Darwinian SPECIES CONCEPT: HIS paper ON THE swallowtail BUTTERFLIES (PAPILIONIDAE) OF 1865 ALFRED RUSSEL WALLACE Y EL concepto darwiniano DE ESPECIE: SU TRABAJO DE 1865 SOBRE MARIPOSAS papilio (PAPILIONIDAE) Jam ES MA LLET 1 Galton Laboratory, Department of Biology, University College London, 4 Stephenson Way, London UK, NW1 2HE E-mail: [email protected] Abstract Soon after his return from the Malay Archipelago, Alfred Russel Wallace published one of his most significant papers. The paper used butterflies of the family Papilionidae as a model system for testing evolutionary hypotheses, and included a revision of the Papilionidae of the region, as well as the description of some 20 new species. Wallace argued that the Papilionidae were the most advanced butterflies, against some of his colleagues such as Bates and Trimen who had claimed that the Nymphalidae were more advanced because of their possession of vestigial forelegs. In a very important section, Wallace laid out what is perhaps the clearest Darwinist definition of the differences between species, geographic subspecies, and local ‘varieties.’ He also discussed the relationship of these taxonomic categories to what is now termed ‘reproductive isolation.’ While accepting reproductive isolation as a cause of species, he rejected it as a definition. Instead, species were recognized as forms that overlap spatially and lack intermediates. However, this morphological distinctness argument breaks down for discrete polymorphisms, and Wallace clearly emphasised the conspecificity of non-mimetic males and female Batesian mimetic morphs in Papilio polytes, and also in P. -
Phylogenetic Relationships and Historical Biogeography of Tribes and Genera in the Subfamily Nymphalinae (Lepidoptera: Nymphalidae)
Blackwell Science, LtdOxford, UKBIJBiological Journal of the Linnean Society 0024-4066The Linnean Society of London, 2005? 2005 862 227251 Original Article PHYLOGENY OF NYMPHALINAE N. WAHLBERG ET AL Biological Journal of the Linnean Society, 2005, 86, 227–251. With 5 figures . Phylogenetic relationships and historical biogeography of tribes and genera in the subfamily Nymphalinae (Lepidoptera: Nymphalidae) NIKLAS WAHLBERG1*, ANDREW V. Z. BROWER2 and SÖREN NYLIN1 1Department of Zoology, Stockholm University, S-106 91 Stockholm, Sweden 2Department of Zoology, Oregon State University, Corvallis, Oregon 97331–2907, USA Received 10 January 2004; accepted for publication 12 November 2004 We infer for the first time the phylogenetic relationships of genera and tribes in the ecologically and evolutionarily well-studied subfamily Nymphalinae using DNA sequence data from three genes: 1450 bp of cytochrome oxidase subunit I (COI) (in the mitochondrial genome), 1077 bp of elongation factor 1-alpha (EF1-a) and 400–403 bp of wing- less (both in the nuclear genome). We explore the influence of each gene region on the support given to each node of the most parsimonious tree derived from a combined analysis of all three genes using Partitioned Bremer Support. We also explore the influence of assuming equal weights for all characters in the combined analysis by investigating the stability of clades to different transition/transversion weighting schemes. We find many strongly supported and stable clades in the Nymphalinae. We are also able to identify ‘rogue’ -
Namadgi National Park Plan of Management 2010
PLAN OF MANAGEMENT 2010 Namadgi National Park Namadgi National NAMADGI NATIONAL PARK PLAN OF MANAGEMENT 2010 NAMADGI NATIONAL PARK PLAN OF MANAGEMENT 2010 NAMADGI NATIONAL PARK PLAN OF MANAGEMENT 2010 © Australian Capital Territory, Canberra 2010 ISBN 978-0-642-60526-9 Conservation Series: ISSN 1036-0441: 22 This work is copyright. Apart from any use as permitted under the Copyright Act 1968, no part may be reproduced without the written permission of Land Management and Planning Division, Department of Territory and Municipal Services, GPO Box 158, Canberra ACT 2601. Disclaimer: Any representation, statement, opinion, advice, information or data expressed or implied in this publication is made in good faith but on the basis that the ACT Government, its agents and employees are not liable (whether by reason or negligence, lack of care or otherwise) to any person for any damage or loss whatsoever which has occurred or may occur in relation to that person taking or not taking (as the case may be) action in respect of any representation, statement, advice, information or date referred to above. Published by Land Management and Planning Division (10/0386) Department of Territory and Municipal Services Enquiries: Phone Canberra Connect on 13 22 81 Website: www.tams.act.gov.au Design: Big Island Graphics, Canberra Printed on recycled paper CONTENTS NAMADGI NATIONAL PARK PLAN OF MANAGEMENT 2010 Contents Acknowledgments ............................................................................................................................... -
Observations on Lycaenid Butterflies from Panbari Reserve Forest and Adjoining Areas, Kaziranga, Assam, Northeastern
Journal of Threatened Taxa | www.threatenedtaxa.org | 26 December 2015 | 7(15): 8259–8271 Observations on lycaenid butterflies from Panbari Reserve Forest and adjoining areas, Kaziranga, Assam, northeastern India ISSN 0974-7907 (Online) ISSN 0974-7893 (Print) Communication Short Monsoon Jyoti Gogoi OPEN ACCESS Ph.D Student, Department of Ecology & Environmental Science, Assam University, Silchar, Assam 788011, India [email protected] Abstract: A checklist of 116 taxa of Lycaenidae (Blues) along with made to document the Lycaenidae notes on important species in low elevation forest of Panbari Reserve, of Kaziranga-Karbi Hills Kaziranga - West Karbi Hills, upper Assam is reported in this paper based on surveys conducted during 2007–2012 and some recent sightings till date. Important sightings include Blue Gem Poritia Methods erycinoides elsiei, Square-band Brownie Miletis nymphys porus, Plain Plushblue Flos apidanus ahamus, Blue Royal Ancema carmentalis, Study area Elwes Silverline Spindasis elwesi, Artipe skinneri, etc. The Panbari Reserve Forest (26036’N & 93030’E) is protected under the Kaziranga National Park (KNP) Keywords: Butterfly diversity, Kaziranga, Lycaenidae, northeastern India, Panbari Reserve. as its fourth addition (Images 1a,b & 2). The average elevation of the forest is around 90m. The altitude however ranges from 70–300 m. The reserve is very close to National Highway 37 (NH37) on the Guwahati- The Lycaenidae (Blues) butterfly diversity in low Jorhat route. The reserve falls between Golaghat and elevation forests of Panbari Reserve, Kaziranga - West Karb Anglong (KA) districts of Assam. To the north of Karbi Hills, upper Assam is reported in this paper. Karbi the reserve lies Dollamora proposed reserve in Karbi Hills constitue a chain of hill ranges lying in middle Assam Anglong District and on the southern boundary is a in the southern bank of the river Brahmaputra. -
HAMADRYAS BABOON (Papio Hamadryas) CARE MANUAL
HAMADRYAS BABOON (Papio hamadryas) CARE MANUAL CREATED BY THE AZA Hamadryas Baboon Species Survival Plan® Program IN ASSOCIATION WITH THE AZA Old World Monkey Taxon Advisory Group Hamadryas Baboon (Papio hamadryas) Care Manual Hamadryas Baboon (Papio hamadryas) Care Manual Published by the Association of Zoos and Aquariums in collaboration with the AZA Animal Welfare Committee Formal Citation: AZA Baboon Species Survival Plan®. (2020). Hamadryas Baboon Care Manual. Silver Spring, MD: Association of Zoos and Aquariums. Original Completion Date: July 2020 Authors and Significant Contributors: Jodi Neely Wiley, AZA Hamadryas Baboon SSP Coordinator and Studbook Keeper, North Carolina Zoo Margaret Rousser, Oakland Zoo Terry Webb, Toledo Zoo Ryan Devoe, Disney Animal Kingdom Katie Delk, North Carolina Zoo Michael Maslanka, Smithsonian National Zoological Park and Conservation Biology Institute Reviewers: Joe Knobbe, San Francisco Zoological Gardens, former Old World Monkey TAG Chair, SSP Vice Coordinator Hamadryas Baboon AZA Staff Editors: Felicia Spector, Animal Care Manual Editor Consultant Candice Dorsey, PhD, Senior Vice President, Conservation, Management, & Welfare Sciences Rebecca Greenberg, Animal Programs Director Emily Wagner, Conservation Science & Education Intern Raven Spencer, Conservation, Management, & Welfare Sciences Intern Hana Johnstone, Conservation, Management, & Welfare Sciences Intern Cover Photo Credits: Jodi Neely Wiley, North Carolina Zoo Disclaimer: This manual presents a compilation of knowledge provided by recognized animal experts based on the current science, practice, and technology of animal management. The manual assembles basic requirements, best practices, and animal care recommendations to maximize capacity for excellence in animal care and welfare. The manual should be considered a work in progress, since practices continue to evolve through advances in scientific knowledge. -
57. Accident Prevention Plan INTRODUCTION Like Everything Else in a Farm Enterprise, Health and Security Must Be Managed
Wild Blueberry in a Context of Production Guide... Sustainable Development 57. Accident Prevention Plan INTRODUCTION Like everything else in a farm enterprise, health and security must be managed. This means dedicating a certain amount of time to these concerns, and emphasizing teamwork. The employer must show that he cares about accident prevention, and employees must be involved in fi nding causes and solutions. BILL C-21 AND EMPLOYER OBLIGATIONS In force since March 31, 2004, the Act to amend the Criminal Code of Canada (Bill C-21) included substantial changes with regard to health and safety in the workplace. Employers are now subject to criminal proceedings after a serious accident if measures were not in place to protect the health and safety of workers. In fact, anyone who manages or supervises one or more workers must show that health and safety obligations are respected, taking all reasonable precautions to prevent bodily injury due to accident. Bill C-21 reinforces the Québec Act respecting Occupational health and safety (see References), which stipulates that every worker has a right to working conditions that have proper regard for his health, safety and physical well-being (section 9). The employer thus has an obligation to take whatever measures are necessary to protect the health and safety of workers (section 51). He must ensure that risks are eliminated at the source by organizing the workplace in a safe manner and by training workers on safe techniques and procedures. He is also obligated to provide individual protection equipment, free of charge, and to require workers to use it. -
A Molecular Phylogeny of the Neotropical Butterfly Genus Anartia
MOLECULAR PHYLOGENETICS AND EVOLUTION Molecular Phylogenetics and Evolution 26 (2003) 46–55 www.elsevier.com/locate/ympev A molecular phylogeny of the neotropical butterfly genus Anartia (Lepidoptera: Nymphalidae) Michael J. Blum,a,b,* Eldredge Bermingham,b and Kanchon Dasmahapatrab,c a Department of Biology, Duke University, Durham, NC 27705, USA b Smithsonian Tropical Research Institute, Naos Island Molecular Laboratories, Unit 0948, APO-AA 34002-0948, Panama, FL, USA c Department of Biology, Galton Laboratory, University College, 4 Stephenson Way, London NW1 2HE, UK Received 2 August 2001; received in revised form 17 June 2002 Abstract While Anartia butterflies have served as model organisms for research on the genetics of speciation, no phylogeny has been published to describe interspecific relationships. Here, we present a molecular phylogenetic analysis of Anartia species relationships, using both mitochondrial and nuclear genes. Analyses of both data sets confirm earlier predictions of sister species pairings based primarily on genital morphology. Yet both the mitochondrial and nuclear gene phylogenies demonstrate that Anartia jatrophae is not sister to all other Anartia species, but rather that it is sister to the Anartia fatima–Anartia amathea lineage. Traditional bi- ogeographic explanations for speciation across the genus relied on A. jatrophae being sister to its congeners. These explanations invoked allopatric divergence of sister species pairs and multiple sympatric speciation events to explain why A. jatrophae flies alongside all its congeners. The molecular phylogenies are more consistent with lineage divergence due to vicariance, and range expansion of A. jatrophae to explain its sympatry with congeners. Further interpretations of the tree topologies also suggest how morphological evolution and eco-geographic adaptation may have set species range boundaries.