(Diptera, Conopidae) in the Collection of the Museum Für Naturkunde Berlin
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Dtsch. Entomol. Z. 66 (1) 2019, 41–53 | DOI 10.3897/dez.66.33814 Type catalogue of the thick-headed flies (Diptera, Conopidae) in the collection of the Museum für Naturkunde Berlin Jens-Hermann Stuke1, Joachim Ziegler2 1 Roter Weg 22, 26789 Leer, Germany 2 Museum für Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity at the Humboldt University Berlin, Invalidenstraße 43, 10115 Berlin, Germany http://zoobank.org/40FA7125-B15A-4678-843C-2B3D67B66344 Corresponding author: Jens-Hermann Stuke ([email protected]) Abstract Received 11 February 2019 Accepted 18 April 2019 Type material of thick-headed flies (Diptera, Conopidae) in the collection of the Museum Published 15 May 2019 für Naturkunde Berlin (ZMHB) is documented. The entire collection holds primary type material (i.e. holotypes, lectotypes, syntypes) of 73 species. Five Conops species with Academic editor: previously unknown subgenus belong to the subgenus Asiconops: C. frontosus Kröber, Dominique Zimmermann 1916; C. indicus Kröber, 1916; C. maculiventris Kröber, 1916; C. nigrofasciatus Kröber, 1916; and C. punctifrons Kröber 1916. Two new synonyms are introduced: Conops vag- inalis Rondani, 1865 syn. nov. of Conops truncatus Loew, 1847 and Siniconops fuscatus Key Words Qiao & Chao, 1998 syn. nov. of Physocephala sepulchralis Brunetti, 1912. Asiconops Conops new synonym new subgenus position Physocephala Siniconops Introduction terial of some 73 species. Only the Natural History Mu- seum (London; formerly the British Museum, Natural The recently published catalogue of Conopidae lists the History) and the National Museum of Natural History depositories of primary type material based on published (Smithsonian Institute) (Washington, DC) hold more information (Stuke 2017). This present paper presents a type material. revised catalogue of the type material of Conopidae de- posited in the collection of the Museum für Naturkunde, Leibniz Institute for Evolution and Biodiversity Science, Methods Berlin, Germany (ZMHB). Due to impressive historical and recent research in The original labels are listed as citations. The labels Diptera at the museum, several dipterists have added are listed and numbered in the order found, commenc- type material to the collection, including Hermann Loew ing with the uppermost. Line breaks on labels are in- (1807–1879; 10 species), Theodor Becker (1840–1928; dicated by a slash (“ / “), but if there are “/” signs 10 species); Anton Ferdinand Franz Karsch (1853–1936; on the labels themselves, these are included without 2 species), Oswald Duda (1869–1941; 1 species), Otto spaces before and after. If text on labels could not be Kröber (1882–1969; 41 species and 2 lost species) and identified properly the line is marked by “[?]”. The Jens-Hermann Stuke (1967; 9 species). interpretation of cited locations is given in Table 1. The Berlin collection is one of the most important The following acronyms are used for collections men- for Conopidae worldwide, containing primary type ma- tioned in the text: Copyright Jens-Hermann Stuke, Joachim Ziegler. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 42 Jens-Hermann Stuke & Joachim Ziegler: Type catalogue of the thick-headed flies... MNHN Muséum National d’Histoire Naturelle (Paris, Australoconops unicinctus (Kröber, 1939) France) NHML Natural History Museum (London, UK) Conops unicinctus Kröber 1939b: 603–604; type local- NMW Naturhistorisches Museum (Vienna, Austria) ities: ‟S. W. Australien, Kalamunda”, ‟Port Philipp” – SDEI Senckenberg Deutsches Entomologisches In- available, valid. stitut (Müncheberg, Germany) ♀ syntype: (1) ‟3039”; (2) ‟Typus”; (3) ‟Conops / USNM Smithsonian Institution National Museum of unicinctus / n sp”; (4) ‟pica / pica et aureorufa / Macq. ? Natural History; formerly, United States Na- / Pt. Phil. Coulon”. tional Museum (Washington DC, USA) Kröber (1939) described the material he used for the ZMHB Museum für Naturkunde, Leibniz Institute for description of A. unicinctus as follows: ‟♂♀. ♀ als pica Evolution and Biodiversity Science (Berlin, von mir beschrieben. Type: Berlin. ♂ von S. W. Austral- Germany) ien, Kalamunda, 9.iii.–11.iii. Type: London” [♂♀. ♀ de- ZMUC University of Copenhagen, Zoological Muse- scribed as pica by me. Type: Berlin. ♂ from S. W. Austral- um (Copenhagen, Denmark) ia, Kalamunda, 9.iii.-11.iii. Type: London].” Schneider (2010) and Stuke (2017) misinterpreted this statement and suggested only ‟Australien, Kalamunda” as the type Results locality. In fact Kröber (1916) also redescribed ‟Conops picus Mcq.” [= Australoconops picus (Macquart, 1851)] Abrachyglossum capitatum (Loew, 1847) and gave ‟Port Phillipp” as the location. Therefore ‟Port Phillipp” should be treated as a second locus typicus and Conops capitatus Loew 1847: 2; type localities: ‟Vater- the specimen described above regarded as the female syn- land: Es scheint nicht über das Alpengebiet hinauszuge- type of A. unicinctus. hen und kömmt auch im schlesischen Gebirge vor.” – available, valid. ♀ syntype: (1) “Reinerz / 13/7 41”; (2) ‟Brachyglos- Brachyceraea brevicornis (Loew, 1847) sum ♀ / capitatum Lw / O. Kröber det. 1912”; (3) ‟Ab- Conops brevicornis Loew 1847: 23; type locality: ‟Brusa rachyglossum / capitatum ♀ / Stuke det. 2004”. im nördlichen Kleinasien” – available, valid. ♂ syntype: (1) ‟Riesengeb. / VII”; (2) ‟capitatus ♂ holotype: (1) ‟Brusa / 8/42 A. m”; (2) ‟Coll. / H. / Loew”; (3) ‟Abrachyglossum / capitatum ♂ / Stuke Loew”; (3) ‟Typus”; (4) ‟Brachyceraea ♂ / brevicornis det. 2004”. LW” / O. Kröber det 1912”. ♂ syntype: (1) ‟Altenbg. / 7/8”; (2) ‟Coll. / Gerst.”; (3) ‟Brachyglossum ♂ / capitatum Lw / O. Kröber det. 1912”; Conops acuticornis Loew, 1847 (4) ‟Abrachyglossum / capitatum ♂ / Stuke det. 2004”“. Loew (1847) described only the female of A. capitatum Conops acuticornis Loew 1847: 8-9; type localities: but obviously had several specimens at hand because he ‟Das Alpengebiet; auch in Württemberg und Schlesien” gave a range for the distribution of the new species. In – available, invalid: junior synonym of Conops ceriae- ZMHB there are only three specimens likely to have been formis Meigen, 1824. collected before 1847, and which, therefore, should be ♂ syntypus: (1) ‟Märkl.”; (2) ‟Coll. / H. Loew”; (3) accepted as syntypes. ‟Typus”; (4) ‟Conopilla ♂ / ceriaeformis mg / O. Kröber det 1912”; (5) ‟Zool. Mus. / Berlin”; (6) ‟Conops ♂ / Australoconops perbellus (Kröber, 1939) ceriaeformis / Stuke det. 2003”. ♂ syntypus: (1) ‟Siles. / Sellind.”; (2) ‟Coll. / H. Conops perbellum Kröber 1939b: 601–602; type locali- Loew”; (3) ‟Paratypus”; (4) ‟Conopilla ♂ / ceriaeformis ty: ‟W. Australien, Marloo Station, Wurarga” – available, mg / O. Kröber det 1912”; (5) ‟Zool. Mus. / Berlin”; (6) valid. ‟Conops ♂ / ceriaeformis / Stuke det. 2003”. ♂ holotype: (1) ‟Type”; (2) ‟Typus”; (3) ‟W. Austral- ♂ syntypus: (1) ‟cc. C. signat / v. Roser * / Stuttg.”; ien / Marloo Station / Wurarga 8.1935 / Gebr. Goerling S. (2) ‟Coll. / H. Loew”; (3) ‟Paratypus”; (4) ‟Conopilla ♂ / G.”; (4) ‟Conops / perbellum / ♂ Kröb.”. ceriaeformis mg / O. Kröber det 1912”; (5) ‟Zool. Mus. / Berlin”; (6) ‟Conops ♂ / ceriaeformis / Stuke det. 2003”. Australoconops pseudocellifer (Kröber, 1939) Conops pseudocellifer Kröber 1939b: 601; type locality: Conops atrimanus Kröber, 1939 ‟Neuholland” – available, valid. Conops apicalis var. atrimanus Kröber 1939a: 379; type lo- ♀ holotype: (1) ‟Type”; (2) ‟5518”; (3) ‟Typus”; (4) cality: ‟Uam, Kamerun” – available, invalid: available, in- ‟Conops / pseudocellifer / 1938 ♀ Krb”; (5) ‟Conops ♀ valid: junior synonym of Conops ferruginosus Kröber, 1915. / aurosa Herm / O. Kröber det. 1914”; (6) ‟aureorufa / ♀ holotype: (1) ‟S. O. Kamerun / Uam-Gebiet / G. Macq. / Nov.Holl. Schultz”. Tessmann S. G. / bei Bossum [written perpendicularly] / 29.6.14 [written perpendicularly]”; (2) ‟Conops apicalis / var atrimanus Krb. / det. Kröber 1938”. dez.pensoft.net Dtsch. Entomol. Z. 66 (1) 2019, 41–53 43 Table 1. Historical locations on labels mentioned in the text and their modern interpretation. Adeleide Adelaide [Australia, South Australia] Alai Geb. Alay Mountains [Kyrgyzstan & Tajikistan] Altenbg. Altenburg [Germany, Thuringia] Amboin. Amboina [Indonesia, Maluku Islands] Argentinien / Prov. Buenos Aires Buenos Aires Province [Argentinia] Asia minor / Taurus cilic. Central Taurus Mountains [Turkey] Assam Assam [India] Astrabad Gorgan [Iran, Golestan Province] Asuncion, / Paraguay Asunción [Paraguay] Asuncion, / Paraguy / Villa Morra Villa Morra, neighborhood of Asunción [Paraguay] Bampur Bampur, town and river [Iran, Sistan and Baluchestan Province] Basman Bazman, town and mountain [Iran, Sistan and Baluchestan Province] Berlin Berlin [Germany] Betschmanaland-Prot. / Kalahari / Severelela - Kooa between ‟Severelela” and ‟Kooa” in the Kalahari Desert [Botswana] Bozen Bolzano [Italy, South Tyrol] Brusa Bursa [Turkey, Bursa Province] Carolina North- and South Carolina [USA] Costa Rica / S. Jose San José [Costa Rica] Dalmat. Dalmatia [Croatia] Dech-i-Papid (=Дэх-и-Пабид) Deh-e Pabid, village NW Eskelabad 28°37’N 60°47’E [Iran, Sistan and Baluchestan Province] Dus-ab (=Дузъ-абъ) Zahedan town [Iran, Sistan and Baluchestan Province] Georgia Georgia [USA] Graecia [Greece] Hungria [Hungaria] Kala-i-bid (=Кала-и Бидъ) Qal’eh-ye Bid, village, 28°39’N 60°22’E [Iran, Sistan and Baluchestan