Museum, University of Nebraska State Insecta Mundi

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An illustrated descriptive catalogue of the genera Anidrytus Gerstaecker and Epopterus Chevrolat, with descriptions of new species (Coleoptera: Endomychidae)

H. F. Strohecker Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services, Gainesville, FL,

This paper is posted at DigitalCommons@University of Nebraska - Lincoln. http://digitalcommons.unl.edu/insectamundi/266 158 Volume 11, No. 2, June, 1997, INSECTA MUNDI

An illustrated descriptive catalogue of the genera Anidrytus Gerstaecker and Epopterus Chevrolat, with descriptions of new species (Coleoptera: Endomychidae)

H. F. Stroheckerl Research Associate, Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services 1911 SW 34th St. Gainesville, FL 32614-7 100 U.S.A.

Abstract: The endomychit1 beetle genera Artidrytus Gerstaecker and Epopt.erus Chevrolat have been confused for many years. This paper discusses the similarities ancl differencesof these genera, describes new species, makes some nomenclatural changes, and illustrates the genitalia for many species. New species described: Epo~>teruspicticollis,E. bioculatus, E. cortfirsns, E. grucilis, E. lor.etertsis,E. atrort~nl~rs,E. arcruacus,E. utriuerttris, E. crypticus, E. qrrechuanus, E. calleriar~~rs,E. fluuortotat~~s, E. srrbrttaclrlatus, E. pcrrucrs, Attidrytus rtirrtbiferus,A. parki, A. corrrpactus,A. gibbosus, A. circurrlcirrctus,A. becl~yr~eorurrt,A. bat,esi,A. rt~r~xicctr1us,A. trir~itntis, A. r~tctjor,A. Ir~l.rt~erosus,A. grurrdis, A. cardiosonta. New synonymies:E. uacrc.ns Gerstaecker (= E. sccrlaris Gorham);E. decerr~purtctutusGerstaecker (= E. bifasciutus Pic);E. uuriegutus Erichson (= E. decoratus Kirsch); A. cotrtr.ucti~sGerstaeclter (= A. dolosus Gorham); A. heluolus Gerstaecker (= Epitebus igrlobilis Gorham);A. parallelus Gerstaecker (= Ephebus depressirs Gorham). Changes in nomenclatural status or combination: Epopterus kirshi Strohecker is reduced to a subspecies of E. pclrtitirs Gerstaecker; E. arlgrrstatus Strohecksr is reduced to a subspecies of E. fascicttus (Fabricius);E. filligittosus GuBrin-MenBville is transferred to Artidrytus.

The genera A~bidrytu,sGerstaecker anclEpopter- Most previous descriptions have been drawn on us Chevrolat share many common features, among body form and color pattern. Many similarities in them: body form ovate to oblong; dorsum pubes- these features are found among the species. Yet, cent, but much abraded in many specimens; anten- strong differences in color pattern are shown by nomeres I11 & IV subequal; antennomeres IS & X, specimens of El)opterus which, on the basis of especially X, mesally acuminate; spinasternum aedeagal structure, are conspecific. Some of the horizontal, extencled beyond front coxae, broadly observed variation is due, no doubt, to dirferences spatulate with apex subtruncate. Maleshaveprotib- in maturity, but may also exist as color phases in ia curvecl, enlarged in apical third, often with a some species. I have used trinomials only where small tootth below; and abdominal sternite VI is names have already been published. visible. I consider the aedeagus the most reliable taxo- Gerstaecker (1858) clistinguished the two gen- nomic character. This organ is strongly sclerotized, era on the basis of mandibular and maxillary struc- often asymmetrical and bizarre in form, which ture: mantlibular apex in Ar~idrytrisis entire, in cliffers in appearance with change of position, ei- Epopterus it is bifid; maxillary galea is corneous at ther by rotation or inclination. The camera lucida base in Alr.idryt,rr.s,in Epopterus it is entirely mem- drawings are of destro-ventral aspect, the lesser branous. I doubt t,hat Gerstaecker subjectecl all the curvature of the base is consiclerecl ventral. species which he treated to much analysis but, Study of the aedeagi suggests relationships rather, fell into the habit of assigning plainly col- between species which transgress the nominal ge- ored and mostly larger forms to Ar~idrytusancl neric boundaries of these genera and suggests long those of strongly variegated pattern and mostly separated evolutionary lines not reflected in cur- smaller size to Epopterus. Later authors, notably rent nomenclature. Evaluation and action on these Gorham, followed him in this practice. ideas would be a long task, for which material and I have not so tested the many species now time left to me are inadequate. Therefore, I follow known but, after examining the holo- or lectotype of Gerstaecker's and Gorham's concepts in t,he assign- nearly all described forms, I think that a prelimi- men t of species. nary treatment may enable any future student of Because of interspeclfic similarities and in- the group to proceed without repeating my efforts, traspecific variations in external features, I found which have extended over many years. it impossible to construct a worthwhile key. I would be loathe to attach identity labels to most females Dr. Stroheclter finishetl this pnper before his death, November 14, 1988. The manuscript has been changed little from its original form, and lacks certain details generally required for modern systematicpapers and by Zrzsecta Mirrtdi. Even in its current state, it represents a step forward in the study of these genera and is here published for that reason. Paul E. Skelley, Florida State Collectio~lof Arthropocls, Florida Department of Agriculture & Co~lsumerServices, Division of Plant Intlustry, 1911 SW 34th St., Gainesville, FL 32614-7100, U.S.A. INSECTA MUNDI, Vol. 11, No. 2, June, 1997 unless in series with males, and I believe the most for the following nominal species: E. rr1,yops GuBrin- reliable path to identification of males is by compar- MBneville and E. trirnacu.la.tus GuBrin-MBneville, ing figures of the aedeagi with the specimen. Each which I have been unable to locate; E. gerbicu1a.tu.s figure is followed by a number in parentheses Pic, which is compared with E. picticollis n. sp.; E. which indicates t,he species number in the text. ryei Gorham, E. 1ir~eoguttu.tusGorham, E. pau- Material esamineclfor this study is foundin the tl~erirtusGorham, and E. ocellatus rriaculosus following institutions, the officers of which I thank Gorham. Habitus figures of the last four accompa- for their cooperation: American Museum of Natu- ny the original descriptions. ral History, New York (AMNI-I); California Acade- my of Sciences, San Francisco (CASC); Canadian 1. Epopterr~socellatus (Olivier) Museum of Nat,ure, Ottawa (CMNC); Deutsches Fig. 24, 33 Entomoligisches Institut, Eberswalde Finow Erotylus ocellatus Olivier 1792:437. Type reposit,ory (DEIC); Field Museum of Natural History, Chicago unknown. Eutrl.orphus ocellatus (Olivier) 1808:1067, pl. 1, fig. 6. (FMNH); Floricla State Collection of Arthropods, Epopterus ocellatus Gerstaecker 1858:275, pl. 3, fig. Gainesville (FSCA); Hungarian Natural History 43u, x; Strohecker 1953:pl. 3, fig. 30. Museum, Budapest (I-INI-IM); Funcacion e Institu- to Miguel Lillo, Tucumfin (IMLA); Instituto de Oblong, rather flat above, almost equally round- Zoologia Agricola, Universidacl Central de Venezu- ed in front and in rear. Legs, head and antenna1 ela, Maracay (IZAV); Ent,omologisches Institut, base reclclish yellow. Venter and pronot,um reddish Museum G. Frey, Tiitzing (MGFrI'); Museum brown, the latter witth 4 black spots in a transverse &Histsoire Naturelle, Geneva (MI-ING); Musdum row. Elytron blackish brown with suture reddish. A National cTI-Iistoire Nat,urelle, Paris (MNHN); Nat- yellow band on base of elytron encloses a dark ural I-Iist,ory Museum, London (NHML); Staatli- humeral spot and a large dark ocellus, almost or ches Museum fur Tierkuncle, Dresden (ShfTD); U. quite touching base. A yellow pre-apical band emits S. National Museum, M'ashington (USNM); Muse- a short ray at middle of its front edge and 2 long um fur Naturkunde cler I-Iumboldt Universitat, posterior rays, one along margin, other near su- Berlin (ZMI-IB); Museum & Institute of Zoology, ture; it may form 2 complete rings enclosing dark Polish Academy of Sciences, Warsaw (ZMPA); Zoo- spots. Front tibia of male abruptly widened beyond logicalMuseum, University of Copenhagen, Copen- miclclle. Length 6-6.5 mm, width 3.4-3.6mm. hagen (ZhfUC); Zoologische St,nntssammlung, Mu- Most specimens examined are labeled "Brazil". nich (ZSMC). Brazil: Sta. Catarina, Corupa, XII. 1944, A. Muller Genus Epopterus Chevrolat (AMN 1-1). Chevrolat 1844::380; Erichson 1847:181; Gerstaeclter 2. Epopterus annr~latusGukrin-Mkneville 1858:274; Cl~apuis1876: 123; Gorham 1890:129; Fig. 34 Stroheclier 1953:62. Epopterus ar~r~i~latusGuBrin-MQneville 1857:268, pi. Type species: Erotylr~socel1a.t~~ Olivier (Stroheclier 13., fig. 20. Holotype 6,Brazil (NHML). 1953:63). Very similar to and probably a color form of E. hfost species r3.5 - 5 mm long, although some ocelltr.tus, as noted by Csiki (1901:47). The figure in may slightly exceed 6 mm. Body outline broadly Guerin-Mkneville does not properly show the long cordate t,o oblong ant1 subparallel. Elytra variegat- caudal rays of the posterior elytral band. ed with black or brown and yellow or red markings, "Guyana", (FSCA). often wit,h a brontl tlarl; band across midclle. In two species, E. lir~eogrcl,l,tr.l,t~sGorham and E. clarn 3. Epopterus myops Gukrin-Mbneville Gorham, the pale elytra are marked with black points or st,reaks. The clorsum is pubescent, but E(popterr~s) trlyops Guitrin-MBneville 1857:268. Type often much abraded in museum material. material from Brazil?, Colombia?; repository un- Diagrams of color pattern only are shown for known. four species describetl from unique female t,ypes: E. Epopterus ocellatris nyops Csilci 1901:47. vicir~usPic, E. fcr,llcrx Gerstaecker, E, rubigir~osus Gerstaecker, and E. qr~nesitl~sGorham. All figures Originally described as tawny brown, elytra of elytra are intrended to show only pattern; any very dark, pronoturn lighter, venter and legs yel- realism is f~rtuit~ous.No illustrations are offered lowish, femora briefly fuscous at base. Pronotum 160 Volume 11, No. 2, June, 1997, INSECTA MUNDI with 4 black spots. Elytra finely punctate, black at 7. Epopterus partitus kirschi Strohecker, base and apex, with an anterior yellow band form- new status ing a circle near the scutellum and another pale Fig. 36 band at apical third. A smaller variety. Epopterus lzirscl~iStrohecker 1959:181, fig. 8. Holotype 6,Ecuador: Balzapamba (ZMHB). 4. Epopterus trimaculatus Gukrin-Mkneville More intensively colored than nominat,e form; Epopterus trit~iaculatt~sGuBrin-MQneville 1857:268. head, legs, middle of rnetasternum and first stern- Type material froni Colombia; repository unknown. ite black. Pronotum rufous with 4 black spots. Elytron with pale areas reduced to rather narrow Original descril~tionasfollows: Body and elytra bands, apex yellow. ~h~ aedeagus differs littlefrom black, shining, pronotum with anterior border yel- that of the nominate form Length 6-6.5 mm, width low. All antennomeres yellow, blackish at base. 3.5-3.7 mm. Legs black, knees ancl tarsi yellow. Elytron with a Colombia: Valle de Cauca, Anchicaya Dam, 70 large posterior yellow area, which is dentate on km E Buenaventura, 22.VII. 1970,H. &A. Howden, front edge and extends almost to apex, enclosing 3 266 (CMNC; FSCA). black spots, the miclclle one largest. Length 5.33 mm, width 3 mm. 8. Epopterus tigrinus Gerstaecker Fig. 37 5. Epopterus partitr'~partitr'~ Gerstaecker Epopterus tigrir~usGerstaecker 1858:279. Holotype 6, Fig. 35 Brazil (ZMHB).

Epopterus partitr~s Gerstaeclter 1858:277. Lectotype, Oblong, rather flat above. Venter, bases of Mexico: Yucatan (NIHML).A syntype Q is presum- femora and tibiae black. Pronotum black, sides ably in ZMPA. yellow with a small black spot near lateral margin. Oblong, elytra more gradually rounded caudad Elytron black, with an irregular yellow or orange than in E. ocellatus. Venter reddish yellow; thorac- band across base enclosing 2 dark spots. A broad ic sterna ancl legs may be dusky. Antenna1 base and apical pale band encloses a lateral dark spot and a pronotum recldish yellow, the latter with 4 black semicircular black spot on suture. Length 5.7-6.5 spots in a t'ransverse row. Elyt,ron reddish yellow mm, width 3-3.5 mm. with 2 black spot,s on base, smaller on umbo, larger Brazil: Sta. Catarina, Lagos, Fruhstorfer near scutellum and narrowly reaching elytral base; (ZMHB; FSCA); Nova Teutonia, F. Plaumann wicle blacli band across midtlle, U-excavate in front (GMFT; FSCA); Parana (FSCA). and V-excised at mitltlle of hind eclge. Black color extending forward along suture almost to scutel- 9. Epopterus geminus Gerstaecker lum and also cauctad, where it widens into a semi- Fig. 38 circular spot; lateral is an oval black spot. Length 6- Epopterusgetninus Gerstaecker 1858:279. Lectotype 6, 6.8 mm, wiclth 3.5-4 mm. Colombia (ZMHB).

6. Epopterus partitus maculosus Gorhanl Venter and legs black. Heact and antenna1base reddish yellow. Pronotum with front angles and Epopterus ocellntt~st~lacl~losrts Gorham 1890:129, pl. VII, fig. 19. sides yellow, disc with large trapezoidal black patch; Epopterus pnrtitus r~~ncr~los~ss:Arrow 1920: 48. Lecto- small black spot in pale area of each side. Elytron type 6,Nicaragua: Chontales, Janson, B. C. A., Col. mostly black; an irregular orange band across base VII, E. oceliatits 01. (NI-IML). encloses 2 black marks which reach base. A yellow area at apex encloses 2 oval black spots. Length 5.5 Differs from nominat,e form only in having the - 6 mm, width 3.3 mm. broacl black elytxal bancl interrupted at middle by a Venezuela: Caracas, Berg Avila, P. C. Vogl pale vitta, which, however, may be incomplete. (ZMHB; FSCA). Colombia: Santander del Norte, Reported by Gorham from Belize; Guatemala; Santiago, 700 m, 23.V. 1974, H. & A. Howden, 1 Nicaragua; Mexico: Tabasco. A considerable series pair; 30 km S Cucuta, 8.V. 1974,Howclens, 6 (CMNC; of paralectotypes are in NHML. FSCA). INSECTA MUNDI, Vol. 11, No. 2, June, 1997

10. Epopterus signaticollis Gerstaecker 12. Epopterus testudinarius Gorham Fig. 29, 30, 39 Fig. 31, 45

Epopterus sig~~aticollisGerstaecker 1858:280.Holotype Epopterus testudir~ariusGorham 1873:48, pl., fig. 7. 6,Colonlbia (ZMHB). Lectotype d,paralectotype Q, Brazil: Amazon, Bates Epopterus scalaris (in part) Gorham 1890:130. (NHML).

Oblong, venter yellow, hind part of metaster- Oblong. Antenna, legs, and venter black. Head num and first sternite black, sternites 2-4 dusky at and pronotum yellow, latter with large black patch middle. Head and antenna1 base yellow. Pronotum which may extend across ent,ire base, abruptly yellow, middle of front margin ancl 4 discal spots narrowed at middle, then continues to front mar- black. Elytron reddish yellow with a small dark gin. Scutellum black. Elytron yellow with small spot on umbo, an oval dark spot at mid-line of base, oblong black mark on shoulder, smaller round a broad multidentate dark band across middle and black spot near scut,ellum, and broad, irregularly 2 black spots before apex. Length 5.5 mm, width 3.5 margined black band across middle. Apex black, mm. between it and middle black band a deeply con- The description above was extractedfrom Ger- stricted yellow band. In a male from Villavicencio staecker. A male from Colombia: Villavicencio, has the pronotum is wholly black, the ent,ire side mar- the inner front spot narrowly connected to base, gin of elytron yellow. Length 6-6.5 mm, witlt,h 3.6- and t,he middle black band interr~pt~edat the side. 3.8 mm. A male from Panama: Bugaba (FSCA ex Janson) Colombia: hleta, Villavicencio, 13.VIII.1938, has venter ant1 legs entirely black. (2.1-I Seevers (FSCA). Colombia: Cundinamarca, Monterredondo, Schneble, 1 pair (GMFT); Meta, Villavicencio, 13. Epopterus comptus Gorliani 9.VIII.1938, C. I-I. Seevers, d(FSCA); Antioquia, Fig. 44 Puert,o Bcrrio, VIII. 1938. Seevers, @(l?SCA).Pan- Epopterus corr~ptusGorham 1890:130, pl. 7, fig. 200. Lectotype 6, 4 paralectotypes, Nicaragua: Chon- ama: Bugaba, 250-400 m, Champion, d(FSCA ex tales, Janson, Belt (NHML). Janson). Oblong. Venter black, paler caudad, legs yel- 11. Epopterus uacuus Gerstaecker low. Antenna yellow at base, antfennomeresVI-XI Fig. 40 black. Heacl and pronotum pitchy red, latter with basal 315 of sicle margin, 2 anterior and 3 elongate Epopterus vacli,~rsGerstaeclier 1868:283. Holotype 6, basal spots black. Elytra black with umbo yellow, French Guiana: Cayenne (MNIHN). pale color extending to suture as a ring enclosing Epopterzrs scalaris Gorham 1890:130, pl. VII, fig. 20. round black spot. Elytral apex extensively yellow Lectotype 6,Panama: Bugaba, Champion (NHML). with large clark spot enclosed. Length 5.8-6 mm, New synonym. width 3.5 mm. Nicaragua: Chontales, d probably from origi- Venter, femora and tibia1 bases black. Antenna nal collection; Q without locality data (FSCA ex stout, black, basal antennomeres (except 111) hard- Janson). ly longer than wide. I-lead dusky yellow. Pronotum with sides yellow, disc black, the dark color abrupt- 14. Epopterus undulatus Gubrin-Mbneville ly narrowed at mid-length ancl continued narrowly Fig. 41 forward to front margin. Elyt,ron black with a large Epopterus z~r~dztlatrtsGuQrin-MQneville 1857:269, 111. yellow patch on shoulder enclosing a small black 13, fig. 21. Type material fro111 Argentina: Buenos spot. The pale color does not reach suture, and its Aires, Dejean coll.; repository unknown. hind edge is t,ritlentate. Apex yellow, front edge of this pale area tridcnt,at,e. Length 6 mm. Oblong. Venter black, abdomen paler caudad. Legs yellow, basal half of middle and hind femora Although Gerstaecker cited a single female in and of all tibiae black. Antenna1 base and head the Thomson collection, t,he presumptive type is a yellow. Pronotum yellow with the following black male; the front tibia is less strongly modlfied than marks: an invertecl V with apex at midclle of front in the males of most species of the genus. margin, its lobate limbs extending to hind angles, a 162 Volume 11, No. 2, June, 1997, INSECTA MUNDI

small round spot on each side. These markings may especially the posterior one, which runs in undu- be obscured by general darkening. Elytron black lant fashion obliquely caudad and mesad. with narrow yellow band across base, enclosing black humeral spot and elongateparascutellar dark 18. Epopterus nigerrimus Strohecker mark; a narrow zigzag pale band at posterior quar- Fig. 70 ter, and lunat,e bar close to apex. Length 5-5.8 mm, Epopterus ~~igerrin~usStrohecker 1957:268, fig. 26. width 3-3.2 mm. Holotype 6, Brazil: Goyaz, Jatahy (HNHM). Argentina: TucumBn, TucumBn, Horco Molle, San Pedro de Colabao, Choromoro; Jujuy: Yuto; Narrowly oblong, subparallel. Deep black be- low and above, dorsum thickly punctate. Elytron Catamarca, El Alto. Bolivia: Sta. Cruz, El Cidral with 2 narrow red marks, the anterior Y-shaped, (IMLA). Argentina: Cortloba, Jesus Maria (GMFT). embracing the umbo, the posterior near apical Paraguay: Asuncion (I-INI-IM). third, transverse and undulate. Length 4.6-4.8 15. Epopterus zebra Strohecker mm, width 2.7 mm. Fig. 3, 69 Argentina: Cordoba, 266, Q (ZSMC; FSCA). Epopterus zebra Strohecker 1957:269, fig. 27. Holotype 6, Peru (HNHM). 19. Epopterus biocr~latusStrohecker, n. sp. Fig. 23, 55 Oblong. Upper surface reddish yellow. Prono- tum with a W-shaped mark and median basal spot Holotype 6, French Guiana: Cayenne (FSCA ex Jan- black. Elytron marked with black as follows: 2 son). round spot,s on base, an undulate band before Long-oval, elytra gradually narrowed from middle, another band just behind middle, and 2 anterior third. Venter and femoral bases clark brown. large coalescent spot,s before apex. Length 5.9 mm, Antenna1 base yellow, club black. Dorsum blackish width 3.3 mm. brown, elytron with anterior and posterior yellow 16. Epopterus picticollis Strohecker, n. sp. rings. Pronotum with sides subparallel basad, fair- Fig. 28, 76 ly strongly curved to front angles, hind angles very sharp, disc finely, sides coarsely and densely punc- Holot,ype 6,Peru: Huanuco, 25 km below Carpish, 2500 tate. Elytra more than 3X as long as pronotum. The m, 9.IX. 1946, F. Woytkowski, and allotype Q with anterior yellow area leaves a large dark humeral same data except 8.IX (AMNH). Paratypes: 29 Q, spot and completely encloses an oval dark spot near same locality, IX. 1948 (FSCA). scutellum. The posterior yellow band encloses a Oblong, subparallel. Venterblackish, distalpart dark lateral spot and is extended in 2 short rays of femora ferruginous. Antenna, except antenno- toward the scutellum. Length 5.3 mm, width 3 mm. mere I, black. I-Ieatl ant1 pronotum reddish brown, The single specimen has a previous determina- the lat,ter wilh a (lark marginal spot on each side tion "E. ocellntus". and 2 broad black marks ext,ending obliquely me- sac1 from hind angles; behind these marks, near 20. Epopterus cr~cr~llinusGorhani center, is a pair of black points. Elytron broadly red Fig. 48 at middle, base ancl apex suffused wit,h black. A narrow yellow band begins at side margin behind Epopterus cucl~llinusGorham 1873:49. Holotype 6, umbo and broadens mesally to form a large ring Brazil: Amazons (NHML). enclosing a large dark spot. At posterior third is a narrow undulate yellow band. Length 5 mm, width Described by Gorham as, "Evenly ovate and 3 mm. rather convex .... Elytra ovate, not much narrowed towards the apex, closely punctured .... rich pitchy- 17. Epopterus geniculatr~sPic red, with 3 orange-yellow markings, the 2 anterior Epopterus get~ic~~lat~~sPic 1931:9. Holotype 6,Bolivia irregular ancl transverse, the third .... an irregular (h4 NH N) . and not clearly defined spot near apex. Antennae pitchy-red, club black." Pronotum with 4 black Similar in size, form and markings to E. picti- spots. Length 5 mm. collis, which is possibly a color form of E. ger~iculn- Brazil: Ega, Bates, 266, Q (NHML); Amazons, tus. The yellow bands of elytron are narrower, 6 (SMTD). INSECTA MUNDI, Vol. 11, No. 2, June, 1997

21. Epopterus dilectus Gorliani 24. Epopterus confusus Stroliecker, n. sp. Fig. 25, 49 Fig. 77 Epopterus cucullinl~s:Strohecker 1957:267, fig. 21 (not Epoptcrus dilectl~sGorham 1875:16. Holotype 6,Ecua- of Gorham). Holotype 6,(Brazil?):Amazon (HNHM). dor: East Andes, Bucltley (NHML). Form oblong, elytra rather strongly convex. Rather narrowly ovate, similar to E. cuculli- Legs and venter brownish black, elytral epipleura nus. Antenna1 base reddish brown, club black. andsidesofprosternum yellow. Dorsum dark brown, Dorsum reddish to dark brown, pronotum with 4 elytron with 2 yellow patches, the anterior closely black spots. Elytron with a pale yellow fascia be- embracing the shoulder and bluntly extended me- hind umbo, inclined slightly caudacl and not reach- socaudad; the posterior crossing the caudal slope, ing suture; another, more irregular fascia across extending from side margin almost to suture. An- caudal slope. Length 4.5 mm. tenna slender, antenomeres 111-VI each about 3X as long as wide, antennomeres VII-VIII longer than 22. Epopterr~svicinus Pic wide, stalk yellowish brown, club black. Pronotum Fig. 14 with sides parallel basad, rather strongly rounded to front angles, finely margined, disc gently convex. Epopterus UC'CC'ILILS Pic 1931:s. Holotype Q, Bolivia Front tibia of male much widened distad, middle (M NH N) . and hind tibia straight. Length 4.9 mm, width 3 Oblong. Pronotum in part black. Elytra pitchy, mm. partly brown, wit,h anterior anciposterior unclulate 25. Epopterus longus Stroliecker and branched yello~vfasciae. Antenna red, club Fig. 67 black, legs mostly red. Length 5 mm. The clescript,ion above is a summary of Pic's Epopterus 1or~g~~sStrohecker1957:268, fig. 22. Holotype diagnosis. Specimens from Callanga, Peru, closely 6,Amazon (HNHM). resemble the holotype: dorsum blackish brown. Pronotum with 4 black spots in a transverse row; Elongate, subparallel. Elytron with 2 trans- these obscure in darker specimens. Figure 14 was verse yellow bands. In appearance much like E. drawn from a Callanga specimen. Its aedeagus is cucu.llir~usbut narrower and with posterior elytral identical to that of E. dilectus, and1 suspect that E. band more undulate. Length 4.5 mm, width 2.5 vicir~usis a color form of E. dilectus. mm. I have seen only the holotype. 23. Epopterr~snotatus Stroliecker Fig. 16, 71 26. Epopterus obliquus Strohecker Fig. 15, 72 Epopterus r~otatl~sStroheclier 1957:266, fig. 18. Holo- type 6,Amazon (HNI-IM). Epopterus obliqul~sStrohecker 1957;265, fig 14. Holo- type 6,Amazon sup.: Haituba (HNHM). The holot,ype and one other specimen subse- quently referred t,o this species are all I have seen; Subparallel in form but with elytra gradually both specimens are in somewhat immature state of narrowedfrom anterior third. Dorsum dark brown, coloration. Elyt.ra1 markings are similar to those of elytron with 3 yellow areas: a quadrate spot at E. vicir~r~sbutwitah yellow areas much wider. In the middle of base, a posthumera1 rounded spot,, and a holotype t,he posterior yellow bancl does not form narrow oblique band across post,erior t,hircl Length complete rings. Length 4.3-4.7 mm, width 2.5-3 4.4 mm, width 2.6 mm. mm. Only the holotype has been stuclied. Colombia: Amazonas, Leticia, 700ft, 9.VII. 1970, H.& A. Howden (CAlNC). 27. Epopterus fallax Gerstaecker Adclitional material is needed to determine the Fig. 21 relat,ionship of E. I)~C~ILILSand E. r~otcrtusto E. dilectus. similar it,^ of the aedeagi suggests con- Epopterus fnllax Gerstaecker 1858:287. Holotype 9, Amazon (ZMHB 21834). specificity. Volume 11, No. 2, June, 1997, INSECTA MUNDI

Form somewhat elongate, elytra widest near have been made from a closely similar example in mid-length. Venter black except front of proster- HNHM. I have seen no other material. num and distalsternites, bases offemora and tibiae fuscous, elytral epipleura yellow. Pronotum red- 30. Epopterus uernicatus Gerstaecker dish brown, head with a black fleck, pronotum with Fig. 26, 79 2 lunate cliscal spots ancl a marginal spot on each side black. Elytron reddish yellow on base andnear Epopterus uert~icatusGerstaecker 1858:285. Holotype apex, the extreme apex and a broad median band Q without locality data (NHML). dark brown. In the anterior yellow area are 2 dark Long-oval. Pronotum with sides convergent spots, the outer on the umbo and rectangular, the from base forward, little rounded, hind angles acu te. inner spot reaching the base and prolonged latero- Ely tra broadened from base in anterior third, grad- caudad as a slender streak. Two dark spots lie in the ually narrowed caudad, strongly convex. Venter, caudal pale area, the inner one touching the dark- legs, antennal base and pronotum reddish brown, ened suture. Length 4.8 mm, width 3 mm. pronotal disc infuscate. Elytra blackish brown. I have seen no additional examples. Behind umbo a yellow band extends mesad enclos- 28. Epopterus gracilis Strohecker, n. sp. ing a dark humeralspot and a larger oval dark spot. Fig. 32, 74 From this yellow band a narrow pale streak divides the dark disc (2 median black spots of Gerstaecker's Holotype 6 and allotype Q, Peru: Pucallpa, 2.X.1954, E. description). Rather narrow yellow bands enclose 2 I. Schlinger & E. S. Ross (CASC, holotype No. 13315). dark spots near apex, which is dark. Length 4.5 mm. Long-oval, elytra rather strongly convex. Ven- A male from Rio de Janeiro resembles the ter and legs reddish brown. Dorsum dark brown, holotype closely but is slightly larger (5 mm) and elytron with Lriramous yellow basal mark and lacks the discal pale streak. bilunat,e yellow band across apical slope. Antenna rather slender, antennomeres 111-VII each longer 31. Epopterus loretensis Strohecker, n. sp. than wide, antennomeres I-VII brown, antenno- Fig. 81 meres VIII-XI black. Pronotum with sides conver- Holotype 6,allotype Q Peru: Loreto, 20 km from Ucay- gent from base forward, little rounded to front ali on Rio Calleria, Colonia Calleria, 10-30.IX.1961, B. Malkin (FMNH). Paratypes: 39 Q with same angles, hind angles slightly divergent and acute, data as holotype (FMNH; FSCA). discalpunctures fine and well spaced, punctation of sides coarser, subrugose. Elytra 3.5X as long as Long-ovate. Venter infuscate, terminal sterni- pronotum, wiclest in anterior third, gradually nar- tes and legs reddish brown. Dorsum dark reddish rowed caudacl. Length 4 mm, width 2.5 mm. brown, elytron with yellow area at base enclosing dark humeral spot and ocellate medial spot, and an 29. Epopterus pictus (Perty) undulate yellow band across apical slope; band Fig. 22, 42 E~~dotttycl~r~spictl~s Per(,y 1832:11 1, pl. 22, fig. 11. Type emits 2 sharp rays in front, and its posterior border material from Brazil; repository unknown. excised by a fuscous spot; apex is pale. Pronotum Epopterus pictus: Gerstaeclter 1858:286. narrowed from base, its sides moderately curved to front angles, marginal bead rather wide and flat, Oval, elytra gradually tapering and strongly hind angles acute. Elytra 3.5X as long aspronotum, convex. Venter, legs, antennal base, headandprono- widened in anterior half, gradually narrowed tum reddish brown, pronotum with 4black spots in caudad. The abrupt widening of front tibia of male a transverse row. Gerst,aecker has described the is less than usual in the genus. Length 4.3-5 mm, elytron as reddish yellow with black-bordered brown width 2.7-3.5mm. markings: a rectangular shoulcler patch and mesad of this a large oval spot; a broad, irregularly mar- 32. Epopterus anomalus Strohecker, n. sp. ginedmedian band; behind this 2 large spots trans- Fig. 83 6, versely placetlantl triangular spot at apex. Length Holotype Ecuador: Pastaza, Ashuara, Rio Macuma a 10 k111 froni R. Morona, 300 m, 7-16.IX.1971, B. 4.4-4.8 mm. Malltin (FMNH). Allotype Q, Colombia: Putomayo, Unfortunately, I broke the aedeagus of the Rio San Miguel, Santa Rosa de Sucumbios, 400 m, Gerstaecker specimen in extracting it. Drawings 1-7.IX.1971, B. Malkin (FMNH). INSECTA MUNDI, Vol. 11, No. 2, June, 1997

3 pre-apical spots black; across middle of elytron is Oval. Venter, femora antl elytral epipleura black. a broad, black-bordered dark red band, Length 4 Dorsum dark reddish brown, elytron with apex and mm, width 2.6 mm. triramous humeral spot yellow. Pronotum relative- ly small, 2X as wide as long, sides parallel basad, 36. Epopterus atriventris Strohecker, n. sp. gently rounded in front, hind angles weakly acute. Fig. 80 Elytra 3.5X as long as pronotum, rapidly widened Holotype 6,Bolivia: Beni, 31.VII-2.VIII. 1960, B. Malkin from base in anterior third, gradually narrowed (FMNH). Allotype Q, Beni, 26.VII-4.VIII. 1960, B. Malkin (FMNH). Paratypes: 8, same data as holo- caudad. The anterior yellow patch covers theumbo, type; 6, Brazil: Matto Grosso, Corumb6 (FSCA). sends a ray across base and a shorter, blunter ray mesocaudad. Length 4.5 mm, width 2.9 mm. Oval, elytra rather strongly convex. Venter and 33. Epopterus aravacus Strohecker, n. sp. legs black, elytral epipleura yellow, clorsum red- Fig. 85 clish brown, elytron with 2 yellow areas. Pronotum narrowecl from base forward but sides well rounclcd Holotype 6,Venezuela: Aragua, 10 mi. nr. Cata, to front angles, hind angles weakly acute, disc 25.11.1971, H. & A. Howden (CMNC). finely and thickly, sides more coarsely and densely punctate. Elytra long-cordiform, about 4X as long Long-oval, similarly roundecl in front and be- as pronotum, base with yellow band enclosing ciark hind. Venter antl femoral bases black. Pronotum umbo andocellate median spot, middle 215 of elytron with disc black,shining, very finely punctate, sides dark reddish brown. A yellow band crosses apical yellow with a black spot on each side, which is more slope, its front edge deeply bilunate, its hind edge or less coalescent with black of disc. Elytron yellow, excised by a large round black spot. Apex pale with umbo and basal spot,, a broad median bancl, 2 brown. Length 4.1 mm, width 2.8 mm. pre-apical spots and extreme apex dark. Median 37. Epopterr~sdecempunctatus Gerstaecker bancl dark brown, its front edge twice (leeply incised Fig. 12, 13, 53, 86 by pale rays, hint1 margin similarly incised. Prono- tum 2.5X as wide as long, sides evenly rounded, Epopterus dccett~pr~r~ctatusGerstaecker 1858:290. Lec- anterior curvature not abrupt. Elytra 3.5X as long totype 6,? Venezuela: Puerto Cabello, Moritz as pronotum and but lit,tle wicler at base, gradually (ZMHB). Paralectotype Q without data (ZMHB). widened to about middle, thence evenly narrowecl Epopterus Oifasciatr~sPic 1931:9. Holotype 6,Venezu- caudad. ela (MNHN).(New synonym).

34. Epopterr~sreticulatr~s Strohecker Oval. Venter and legs rust-red, thoracic sterna Fig. 8, 66 and femoral bases darker. Antenna1 base red, club Epopterr~srcticrtlalr~s Strohecker 1957:264, fig. 12. Ho- black. Upper surface reddish yellow, elytron with 6, lotype Bolivia: Cochabamba, Germain (HNHM). brown or black markings: a rounded spot on shoul- der and, near mid-line, a narrowly oblong spot, Elliptical in outline. Yellow above with disc of obliquely placed and remote from base; a broad pronotum and 5 spots on elytron dark brown. The band across middle, it's front edge twice, its hind pronotal spot is transversely rectangular; the dark edge once excised; 3 spots of variable size before elytral markings consist of a roundedhumeralspot, apex. These spots tend to coalesce to form a dark an oblong spot near middle of base, 2 elongate band, leaving the apex and a pre-apical, strongly rectangles near middle, and a rounded or some- undulate band pale. Length 4.3, width 3 mm. what triangular spot before apex. Length 3.8 mm, Colombia: Antioquia, Puert,o Berrio (GMFT); width 2.5 mm. Norte de Santander, Cucuta, H. & A. Howden 35. Epopterus deltoider~sStroliecker (CMNC; FSCA). Venezuela: Bolivar, Suapure, Fig. 11, 61 Caura R., E. A. Iclages (FSCA); Lara, Sanare, Epopterus dcltoicle~csStrol~eclier1957:266, fig. 20. Holo- 7.XII.65, J. & B. BechynB; Aragua, Cata, 14.1X.65, type d,Anlazon SLID.:Haituba (HNHM). BechynB (IZAV; FSCA). Trinidad: Maracas, H. &A. I-Iow den (CMNC). Oval, elytra narro~vlycordate. Dorsum yellow, elytron with umba, a small oblong basal spot, and 166 Volume 11. No. 2. June. 1997. INSECTA MUNDI

38. Epopterus fasciatus fasciatus (Fabricius) 41. Epopterus cordatr~sStroliecker Fig. 1, 4G Fig. 10, 58 Tritorr~afasciaturt~ Fabricius 1801:570. hctotype 6, Epopterus cordat~~sStrohecker 1957:266, fig. 15. Holo- Am(erica) mer(ic1ionalis) (ZMUC). type 6, Amazon (DEIC). Epopterus fasciat~~s:Gerstaeclcer 1858:288 (not of Fab- ricius). Elytra long-cordate, flattened margin wider than usual. Brown above, elytron with a large Ovate. Venter and dorsum reddish to dark quadrate yellow spot on shoulder, continued as a brown. Elytron with yellow band across base, be- narrow band across base, and a transversely oval ginning at side margin, leaving umbo dark, and yellow spot before apex. Length 3.8-4.5 mm, width enclosing a large ocellate dark spot mesad. A yellow 2.5-2.8 mm. patch before allex encloses a large round dark spot, The type series comprised 15 specimens from which may be partly divided by a pale streak. t,he "upper Amazon". Brazil: Amazonas, Titirico; Length 3.8-4 mm, wiclth 2.4-2.5 mm. Boca Cauarubi; Maturaca, J.& B. Bechyn6. Vene- Brazil: Rio de Janeiro, 256, 2 Q Q (FSCA). zuela: 38 km El Dorado, F. Fernandez & C.J. Rosales; Ocano, Fernandez (IZAV; FSCA). 39. Epopterus fasciatus angustatus Strohecker, new status 42. Epopterus crypticus Stroliecker, n. sp. Fig. 2 Fig. 78 Epopterus ar~g~l.statt~sStroheclcer 1957:268, fig. 23. Ho- lotype 5, Amazon sup.: Haituba (HNHM). Holotype 6, (Brazil?): Amazon (HNHM).

Long-oval. Elytron dark reddish brown, with 2 Form long-oval, elytra strongly convex, venter yellowish whit,e crossbands edged with black. The and legs brownish black, elytral epipleura and anterior band is wide at side margin and ends sides of prosternum yellow. Dorsum brown, prono- narrowly near suture antlscutellum. Posterior band tum with 4 black points in tranverse row, elytron transversely rectangular. Length 4.6 mm, width with 2 black-bordered yellow patches, the anterior 2.6 mm. enclosing small dark humeral spot and bluntly Larger than the nominate form and somewhat prolonged mesocaudad; posterior patch on caudal narrower. The aecleagus is identical to that of E. slope, broadly undulate. Pronotum with sides grad- fasciatus lect,otype. ually convergent from base forward, gently round- ed to rather sharp front angles, hind angle weakly 40. Epopterus variegatus Erichson acute. Elytra more than 3X as long as pronotum, Fig. 6, 7, 43 gradually widened from base to near midlength, then gradually narrower caudad. Antennomere I11 Epopterus uariegatus Erichson 1847: 181. Holotype 6, Peru (ZMHB); Gerstaecker 1858:284. about 2X as long as wide, 7 and 8 hardly longer than Epopterc~sdecoratus Kirsch 1876: 114. . Holotype 6, wide. Front tibia of male much widened distad, Peru (SMTD). New synonym. middle and hind tibia lightly but distinctly curved. Epopterc~s~C~ILLI~~I~,LLS Pic, inedit. Length 4.5 mm, width 2.8 mm. Very similar in appearance to E. cortfusus, the Oval. Light reddish brown below and above, elytral pattern almost identical, but of more oval antenna1 base yellow. Elytron with dark markings form, with elytra more convex, and differing in as follows: a streak along suture, a humeralspot, an features described and illustrated. oval spot near mid-line not touching base, a broad, sinuate median band, a spot before apex; these 43. Epopterus eganus Gorliani markings dark reddish brown with narrow black Fig. 47 borders. Epopterus egarLus Gorham 1886:160, pl. 17, fig. 9. Lec- Description above extracted from original. Spec- totype 6, Brazil: Amazonas, Ega, H. W. Bates (NHML). Paralectotype 6, Ega (NHML). imens from southern Peru are marked as in figure 6, others from Iquitos and "Amazon" as in figure 7. Ovate. Pale chestnut red, elytron with 3 large Length 4-4.5 mm, wiclth 2.4-2.5 mm. yellowish white, black-bordered spots. Of the ante- Peru: Cuzco; Callanga; Marcapata. rior spots one lies against base and suture, the other behind umbo, the posterior spot is round and INSECTA MUNDI, Vol. 11, No. 2, June, 1997 167 on apical slope. The 2 anterior spots are narrowly 47. Epopterr~srubiginosr~s Gerstaecker connected in the lect,otype but quite separate in a Fig. 18 female from Ega (FSCA ex Janson). Length 5-5.25 Epopterus rubigir~osusGerstaecker 1858:291. Holotype mm, width 3.5 mm. 0, French Guiana: Cayenne (MNHN). 44. Epopterus lineogr~ttutusGorham Ovate or somewhat oblong. Reddish brown be- Epopterus lir~eogr~ttatr~sGorham 1886: 161, pl. 17, fig. 7. low and above, thoracic sterna infuscate. Antenna1 Holotype 6,Amazon, H. W. Bates (NHML). base ferruginous, its articles rather short and thick, club black. Pronotum with disc blackish, a vague Identical in structure, including aedeagus, to black spot in each marginal area, front and hind E. egal~usbut very differently marked. Gorham angles sharp. Length 4.7 mm. described the ely tral markings @lack)as follows: "a I have seen no other specimens which closely humeral small (lot, 2 elongate dashes near the resemble the holotype. Possibly the example here suture, 2 shorter ones in the middle of the disc, 2 described as E. bioculatus is the male of E. rubigi- still smaller near the margin above the middle, ancl nosu,s, but it is larger than the holotype, with elytra 1 small one near the apes. The dashes are not more rapidly tapering and differently marked. Se- placedregularly, but the 3 near t,he base form a sort ries of topotypes will be needed for clarification. of fascia, as do the 3 bclow the middle. The scutel- lum is pitchy black. Lengt,h G mm." 48. Epopterus batesi Strohecker Probably a color phase of E. egcrl~us. Fig. 68 Epoptcrus batesi Strohecker 1957:268, fig. 24. Holotype 45. Epopterus ephippiger Gorham 6,Amazon (HNHM). Fig. 54 Epoptcrus cph.ippigc?r Gorhanl 1886:160, pl. 17, fig. 8. Somewhat broadly ovate. Dorsum blackish Holoty~ed, Amazon, Bates (NI-IML). brown, elytron with 2 rather narrow crossbands, the anterior at basal quarter and extending from Ovate. Pitchy rccl bclow and above. Elytra paler side margin almost to suture, where it is narrowly than pronoturn, with a very large round black s1)ot rounded, the posterior band at caudal quarter of common to both. Length G mm, width 4.5 mm. elytron. Length 4.8 mm. I have secn only t.heholotyl~eof this distinctive- ly marked species. 49. Epopterr~sryei Gorham Epopter~~sryei Gorham 1873:50, pl. fig. 3. Holotype Q, 46. Epopterus quech.uanus Strohecker, n. sp. Amazons (NHML). Fig. 27, 75 Holotype 6,Peru: Junin, Tarma, Utcuyacu, 1600-3000 Antennomeres I1 andV-VIII bead-shaped,base m, 111-1918, F. Woytkowski (AMNH). Allotype Q red, club black. Dorsum blackish brown, elytron with same tlata except 12.111.1948 (AMNH). with 2 sharply defined wide yellow bands. The Paratypes: 40 9,Utcuyacu ancl Agua Dulce 111- anterior band embraces the shoulder, is recurved to 1948, Woytliowski (AMNI-I, FSCA). the base and there emits a broad blunt ray obliquely toward the suture; the posterior border of the band Oval. Venter and basal half of femora black, emits 2 short blunt rays. The posterior band, on distal sternit,es, tibiae and ends of femora re(l(1ish caudal slope is strongly undulate. Length 5 mm. brown. Dorsiim blackish brown, elytron with basal rings and apes yellow. Antenna reddish brown, 50. Epopterr~squaesitus Gorham club but little darker, most of basal antennomeres Fig. 20 longer than wide. Pronotal sides weakly conver- Epopterus quaesitus Gorhaln 1873:51. Holotype Q,Am- gent from base forward but rather strongly round- azons (NHML). ed to front angles, hind angles feebly acute, disc finely and some.ivhat clistantly punctate, punctures Similar to E. ryeibut smaller and with antenna1 of sides coarser and denser. Elytra long-cordate, base slenderer. The anterior yellow elytral band each wit,h a bioculat,e basal yellow band; apex lies behind umbo and widens mesad to form a broad extensively yellow with 1 or 2 black spots. Length arc open toward the suture. The posterior band is 5 mm, widt,h 3.5 mm. strongly undulate. Length 4.5 mm. 168 Volume 11, No. 2, June, 1997, INSECTA MUNDI

51. Epopterus similis Strohecker 700 ft, 10.XII. 1970 and 13.11-2.111.1974, H. & A. Fig. 4, 5, 63 I-Iowden (CMNC; FSCA). Epopterus similis Stroheclter 1957:265, fig. 13. Holo- type d, Amazon sup.: Haituba (HNHM). 54. Epopterus cyclicus Strohecker Fig. 9, 62 Ovate. Light to dark reddish brown, elytron Epopterus cyclicus Strohecker 1957:268, fig. 25. Holo- with 2 rounded or somewhat rectangular, black type Peru: Janjui (HNHM). bordered, pale yellow spot,s. Of these the anterior lies just behind umbo and is distant from suture. Short-oval. Pronotum ferrugineous. Elytron The posterior spot is similar to the anterior one in dark brown with 2 large yellow marks, the anterior, size and shape and lies on posterior slope. Remarks just behind shoulder, rectangular with its anter- on variation in pattern were made in original de- omesal angle narrowly extended along scutellum scription. Spccimensfrom Ecuador and Peru (Lore- and reflexed laterad along base; the posterior mark to; Huanuco) have the basal elytralmark triramous is a ring, narrowly closed along margin. Length 4.2 and the posterior mark an undulant band but agree mm, width 2.9 mm. in all structural features, including aecleagus, with Peru: San Martin, Janjui, 1600 ft, 5.XII. 1946, the holotype. Lengt,h 4.5-5 mm, width 3-3.5 mm. J. C. Pallister (AMNH; FSCA). Peru: Loreto, Pucallpa; Huanuco, TingoMaria, J.C. Pallister. Ecuntlor: Pastaza, Ashuara, Rio 55. Epopterus callerianus Stroliecker, n. sp. Macuma 10 km from R. Moronca, B. Malkin (FMNH; Fig.- 82 Holotype 6 and allotype Q, Ecuador: Pastaza, Ashuara, FSCA). Rio Macuma 10 km from R. Morona, 300 m, 7- In antcnnnal stxucturc this taxon resembles E. 16.VII. 1971, B. Malkin (FMNH). qr~aesitus,and I am inclined to think that it is a color form of that species. Broadly oval but elytra somewhat long-cordi- form. Venter and femoral bases infuscate, tibiae, 52. Epopterus transversus Stroliecker femoral apices and elytral epipleura ferruginous. Fig. 56, 59 Dorsum reddish brown, elytron with biocellate yel- Epoptercts tran.soersrts St,rolleclter 1957:264, fig. 10, A, B. Holotyl~ed, Bolivia: Cochabamba (HNHM). low band across base and an undulate pale band across apical slope. Pronotum narrowed from base Ovate, elytxa broatlly cordat,e. Elyt,ron mostly forward, marginal bead wide and flat, hind angles black buk \vit,h base and apes extensively, suture weakly acute. Length 3.7-4 mm, width 2.5-2.7 mm. narrowly yellow. The basal pale area leaves the 56. Epopterus flauolineatus Strohecker umbo dark and encloses 1-1 large black spot which Fig. 60 joins t,hc broad black middle area of elytron. Ap- Epopterus flavolit~eatusStrohecker 1957:266, fig. 16. proximately the posterior quarter of elytron is pale, Holotype 6,(Brazil?): Amazon (HNHM). enclosing 1 or 2 black sl~ot~s.Length 4.4-4.7 mm, widt,h 2.6-2.8 mm. Oval. Elytra broadly cordate. Legs and venter blackish brown. Dorsum brown, elytron with 2 53. Epopterus nigrocinctus Strohecker yellow cross bands, the anterior behind umbo, ex- Fig. 57 tending from side margin almost to suture, the Epopter~ts~~igrocir~ctri~s Strohecker 1957:264, fig. 11. Holotype d,Amazon (HNHM). similar posterior band on caudal slope. The mar- gins of the bands are almost straight. Length 3.5 Short,-oval,elytra broatlly cordate. Dorsum dark mm, width 2.5 mm. brown, elytron with 2 broad, black-bordered yellow Only the small type series has been seen. bands, one behind umbo, the other on caudal slope. 57. Epopterus flavonotatus Strohecker, n. sp. These bands touch the side margin but do not reach Fig. 65 the suture. Length 4 mm, width 2.6 mm. Epopterus flavot~otatusPic, inedit. Brazil: Amazonas, Boca Cauaburi, 31.111.1964, Epopterus fasciatus: Gerstaecker 1858:288 (not of Fab- J. & B. Bechyn6; Titirico, 13.IV, Bechynes; Matura- ricius). Holotype 6,French Guiana: Cayenne (FSCA ca 7.IV. 1964; Uapes, 23.111.1964, Bechynes (IZAV; ex Janson). Paratypes: Para (ZMHB). Brazil: Belem, FSCA). Venezuela: Amazonas, Cucui, 24.111.1964, XII-1955, G. & B. Frey, 2dd (GMFT; FSCA); Hai- Bechynes (IZAV). Colombia: Amazonas, Leticia, tuba, 266 (HNHM; FSCA). INSECTA MUNDI, Vol. 11, No. 2, June, 1997

Oval, elytra strongly convex. Some examples, 60. Epopterus mimeticus Stroliecker including the Pic specimen apparently have not Fig. 64 attained definitive coloration. In these, the dorsum Epopterus ~r~irt~eticusStrohecker 1959: 181, fig. 5. Holo- is generally pale brown, elytron with a yellow band type 6,Amazon (ZMHB). across base and curving caudad along scutellum and suture. Behind this band is a cliffuse black area Short-oval, similar in form to E. clara. Elytron enclosing 2 yellow spot,s transversely placed. Apex yellow with with 2 dark basal flecks and a large yellow and in front of this pale area are 2 rather black patch at mid-length, which encloses a yellow large rounded yellow spots. spot. Length 4 mm, width 2.5 mm. The pattern which I consider the mature one is similar to figure 19. Legs and venter, inclueling 61. Epopterus dives Gorham Fig. 50 elytral epipleura, dark brown. The distalexpansion Epopterus dives Gorham 1874:443. Holotype 6,Ama- of front tibia of male is rather gradual and moder- zons (NHML). ate. Length 4.3 mm, width 2.8 mm. Short-oval. Dorsum brown, elytron with 2 yel- 58. Epopterus submaculatus Strohecker, n. sp. Fig.87 low marks, the anterior triramous, the posterior Epopterus sr~i)tt~a.cc~latc~sPic inetlit. EIolotype 6,Boliv- undulate, narrowed mesacl and almost reaching ia, Pic coll. (MNI-IN). suture. The anterior yellow bar extends forward along side margin and int,ernal to umbo, emits a Oblong in outline, strongly convex. Legs, an- spur to elytral base, thus enclosing a dark humeral tennal base and prosternum yellow; other parts of spot; from origin of this spur a pale bar deflects venter dusky, becoming paler caudad. Pronotum obliquely caudad, its end a little enlarged and far with disc and hind angles black, front angles and from suture. Length 3.5-3.8 mm, width 2.6 mm. sides broadly yellow. Elytron black with a broacl U. Amazon: Yurimaguas (FSCA ex Janson). transverse yellow bancl behind umbo, reaching 62. Epopterus parvus Stroliecker, n. sp. suture and t,here expancled slight,ly cauclacl and Fig. 19, 73 forward to scut,ellum. Distal third of elytron has 2 Holotype 5 and allotype Q, Brazil: Para (FSCA ex oval yellow spots transversely placed and behind Janson). Paratypes: 50 '2, Para (FSCA). these a large round yellow spot which leaves apex, suture ancl side margin narrowly black. Front Libia Short-oval, elytra cordiform, tapering from of male curvecl at base, clistally enlarged with anterior third. Venter and legs brown. Antennal proximal encl of enlargement mesally angulate. base brownish yellow, club black. Dorsum pale to Lengt,h 3.8 mm, cviclth 2.4 mm. clark brown, elytron with 2 yellow marks: a band at 59. Epopterus clara Gorham base which leaves a clark humeral spot and encloses Fig. 51 an ocellate dark spot near base and scutellum; Epopterus clara Gorham 1874:443. Holotype Q, Ama- apical third yellow, enclosing 1 or 2 black spots, the zons (NHML). single spot due to fusion of the two. Length 3.5-4 mm, width 2.3-2.8mm. Short-oval,conves. Elytxa widest in basal third, 63. Epopterus histrio Gerstaecker gently tapering, apes rather sharply roundecl. Fig. 17, 52 Pronotum and scutellum reddish t,o dark brown. Epopterus histrio Gerstaecker 1858:292. Lectotype Q , Elytron brownish pello~vwith G or 7 black mark- French Guiana: Cayenne (MNHN). ings: a humeral spot ~vhichdoes not reach base and mesad of this and oblique oval spot, 3 patches Short-ovate, elytra cordate, widest in basal transversely placed near middle, the lateral spot third. Venter and legs brownish black. Antennal elongate ancl close to margin, the middle spot larg- base dark red, club black. Head and pronotum dark est and long-oval, the inner spot subquadrate and brown. Elytron black at sides; center of disc ancl slightly behind the others, 2 spots on apical slope apex reddish brown. A yellow band encloses a clark which may be fusecl into an arcuate band. Length humeral spot, turns forward to run along base and 3.8-4 mm, width 2.6-2.8mm. is then deflected along scutellum; the hind border Brazil: ParA; Rio de Janeiro; S5o Paulo (FSCA). of this band emit,s a sharp ray. The band may form 170 Volume 11. No. 2. June. 1997. INSECTA MUNDI

a complete ring mesad to enclose an oval dark spot. ferred to Arcidrytus by Csiki or me: A. ju,uer~cus,A. Another yellow band of irregular shape crosses the ci~~ctiper~nis,A. fuligil~osus, and A. spadiceus. apical third of elytron, it,s hind edge excised by a black spot. This band may also completely encircle 65. Anidrytz~sbipunctatus Gerstaecker the dark spot. Length 3.6-4 mm, width 2.4-2.8 mm. Fig. 92 French Guiana: Cayenne (FSCA). Brazil: Ama- Ar~idrytusDip~~r~ctalus Gerstaecker 1858:257. Lectotype pa, Rio Carnot; Serra Lombartl, J. & B. Bechyn6 6,Brazil (MNHN); paralectotype, Brazil (ZMHB). (IZAV; FSCA); Par6 (FSCA). Long-ovate, bright to dark rust-red with yellow 64. Epopterus pantherinr~sGorliani pubescence; spinasternum, meso- and metaster- Fig. 84 num, basal h.alf of tibiae and base of middle and Epopterus par~tl~erir~r~sGorham 1889: 131, pl. 7, fig. 22. hind femora blackish brown. Antennomeres I-II Lectotype Q, Type1 sp. figured1 David, Chiriqui, red, the others black. Length 9-9.5 mm. Champion/ B. C. A., Col. VIIl Epopterus pantheri- Pronotum more than 2X as wide as long, nar- nus Gorh. (NHML). rowed from base forward, front angles short and rounded, hind angles weakly acute, disc with 2 Oblong-ovate, strongly convex. Antenna black, dark flecks, irregularly punctate. Elytra elongate, antennomeres 1-11 reddish. Tibia1 base and scutel- wider at base than pronotum, gradually widened in lum black. Pronotum redtlish brown, front margin, anterior third then gradually convergent clistad. I basal half of side margin ant1 base narrowly black. have seen only the types. Elytron with gale markings as follows: a broadly triramous band covering most of basal third, 2 66. Anidrytris plagiatus Gerstaecker transversely placed median spots which may be Fig. 93 united, 2 spots on apical slope and 1 at apex. These Ar~idryl~~splagiatusGerstaecker 1858:259. Holot,ype 6, pale areas cover most of elytron, leaving clarker French Guiana: Cayenne (MNHN). areas quite narrow. Length 5.25-6 mm. I have recently examined a pair from Barro Form oval, elytra raoher strongly convex. Ven- Coloraclo Islancl sent by I-Ienk Wolda. ter and legs rust-red. Antennomeres I-V red, the others black. Pronotum 2.5X as kvicle as long, clisc Genus Anidrytr~sGerstaecker with rectangular black patch, sides subparallel Gerstaecker 1858:256; Cl~apuis 1876:122; Gorham basad, abruptly roundecl in fron t, front angles short, 1889: 125; Stroheclter 1953:64. hind angles produced ancl acute. Length 6.5 mm. Type species: Ar~idrytrssbip~o~ctatrss Gerstaecker (Stro- Front tibia of male with inner edge widely hecker 1953:64). excavate beyond middle, then abruptly widened. Specimens of most species are 5.5-9mm long. In 67. Anidrytus nimbiferr~sStroliecker, n. sp. several species, however (Nos. 74-84, 95), length Fig. 94 may be less than 5 mm. Coloration more somber Holotype 6,Colombia: Amazonas, Leticia, 700 ft, than in E~~opterus,often entirely brown or black 11.VII.1970, H. &A. Howden (CMNC); paratype 6, with paler color when present confined to sides and Brazil; Amazonas, Bates (NHML). apex of elytron. Rarely a pale sutural stripe is present. Alcidrytus vnrieg~~1~t~s(Gu6rin-M6neville) Long-oval, elytra rather strongly convex. Ven- of southern Brazil is, as its name indicates, strik- ter, legs, head and antennomeres I-II rust-red, ingly bicolored, while A. decorcr.t,us Arrow and A. antennomeres III-XI shiningblack. Pronotum with hurnerosrrs n.sp. may have elytral base extensively rectangular black patch at middle, sides broadly, yellow or reddish. Apuzzling maculate specimen is front margin narrowly red. Elytra black with base, noted untler A. porcrrs Strohecker. Body outline is including scutellum, side margin and apex red. broadly ovate to oblong or even long parallel. Sides of pronotum convergent from base forward, No illustrations are presentee1 for A. 1citidu.lar- front angles prominent and weakly acute, hind ills Gerstaecker, A. Irilctris Gerstaecker,A. chalnlpi- angles produced and decidedly acute, lateral sulci olti (Gorham), A. quadripr~~cctntusGorham, A. linear with fovea at base. Elytra more than 3X as rrlargir~a.tus(Fabricius), and 4 species described by long as pronotum (mid-line) and as wide at base, GuBrin-M6neville under E')ipocus but were trans- slightly widened to midlength then gradually con- INSECTA MUNDI, Vol. 11, No. 2, June, 1997 vergent caudad. Front tibia of male as in A. plagia- 71. Anidrytus ovatulus Gerstaecker tus. Length 8.3 mm, width 4.9 mm. Fig. 98 Ar~idrytuso~att~lus Gerstaecker 1858:262. Lectotype 6, 68. Anidrytus variegatus (Gubrin-Mbneville) Colombia (ZMHB). Fig. 134 Oval, strongly convex. Rather dark brown, an- E(pipocr~s) unriegatirs GuBrin-MBneville 1857:266, pl. tennomeres 111-XI black, elytral epipleura and tib- 13, fig. 19. Holotype Q ?, Brazil; repository un- iae yellowish brown. Similar to A, col~trnctusbut of known. somewhat longer form. Length G mm, width 3.9 Anidrytus sit~gt~larisGerstaecker 1858:260. Holotype mm. 6,Brazil (MNIIN); Csiki 1901:35 (syn.). Venezuela: Maracay, P. Vogl (ZMHB; FSCA). Short-oval, elytxa cordiform, convex. Prono- 72. Anidrytus parki Strohecker, n. sp. tum narrowed from base forward, reddish yellow Fig. 133 with 3 discal spots and basal half of marginal bead Holotype 6 and allotype Q, Panama: Barro Colorado black. Elytra blackish brown with 2 large pale Island, 28.VII.1973, 0. Park (FMNH). markings: the anterior a broad undulant band behincl umbo, the posterior a large triangular, Oval. Largely rust-red, the distal antennomer- patch near apex. These marks may be connected by es darker but not black. Antenna stout, antenno- a pale band close to suture. Length 6 mm. mere I11 less than 2s as long as wide, antennomeres The type of A. st~lgzilarisis callow, so soft that IV-V subequal to 111, antennomeres VI-VIII each the aedeagus coultl not, be e~tract~ed.Rio cle Janeiro, about as wide as long. Pronotum with sides parallel 6,2 Q Q (NHML). basad, well rounded to front angles, hind angles rectangular, disc very finely punctate, punctures 69. Anidrytus nitidularius Gerstaecker at sides closer and denser. Lateral sulci not quite Ar~idrytust~itidi~lci~rii~s Gerstaecker 1858:262. Holotype reaching mitldle. Elytra not quite as long as prono- (sex ?), Mexico, presumably in Dohrn coll. (ZMPA). tum, rather strongly convex, widest before middle Short, oval. Blackish brown, antennomeres I- and gradually narrowed caudad, reddish black in holotype with umbo, side margins and apex red; V, front angles of pronotum, clytral apex, venter allotype black above, sides of pronotum, elytral and legs red. Length 6 mm. umbo, side margins narrowly and apex red. Length Antennomeres 111-Ilrequal,antennomeresVII- 6-6.5 mm, width 3.7 mm. VIII globular. Pronotum rather narrow, hardly 2.X Front tibia ofmale has angulateflange on inner as wide as long, sitlcs subparallel basad. Elytra etlge. widest in basal third, thence gratlually convergent. Last abdominal sternite (1ecl)ly semicircularly ex- 73. Anidrytus porclis Strohecker cised. I have not recognizccl this species in the Fig. 129 material studied. At~idryli~sporcus Strohecker 1957:269,fig. 28. Holotype 6,Amazon (HNHM). 70. Anidrytlis contractus Gerstaecker Fig. 97 Short-ovalstronglyconvex, sides of ely tra much Ar~idrytusCOIL~,~CLC~,~LS Gerstaecker 1858:263. Holotype rounded. Elytral disc dark brown, base and sides Q , Costa Rica (ZMI-IB 2 1818). reddish brown. Pronotum ant1 antennomeres I-VII Ar~idr~ytusdolosi~s Gorham 1889:127, pi. 7, fig. 18. Lec- totype ancl 5 paralcctot,yl>es, Panama: Volcan c-le recldish brown. In male front tibia is dist,ally wid- Chiriqui, Charnl~ion(NIIML). New synonym. ened ancl bidentate, hind tibia with a short sharp toot,h on inner edge near apex. Length 5.7 mm, Short-oval, strongly convex. Brown t,o blackish wiclth 3.8 mm. brown, antennomeres I-V or 1-VI recldish. Notable Colombia: Amazonas, Leticia, 700 ft, 19- features of the species are the short broad form and 25.11.1972, H. & A. Hotvden. One male of this series the very deep lat,eral sulci of 1)ronotum. Length 5.7- is smaller (5 mm) and has dorsum reddish brown, G mm, width 4-4.2 mm. elytron with 2 large, black-bordered, transversely Costa Rica: Turrialba, A. I-Ieyne (ZMHB; FSCA), oval yellow bands, but is structurely identical, Panama: Barro Colorado Island (CASC; FMNH). including aedeagus, with typical specimens. 172 Volume 11, No. 2, June, 1997, INSECTA MUNDI

74. Anidrytus unicolor Kirsch Short-oblong. Ferruginous to dark brown be- Fig. 128 low and above, antennomeres I-V or I-VI rufous, Ar~idryttss~sr~icolor I

coloured specimens the five basal joints and the margins narrowly red. Pubescence long and gray. extreme tip of the antennae, the sides of the protho- Length 5.5 mm. rax, the shoulders and extremities of the elytra, and the legs are reddish and the remaining parts 84. Anidrytus gibbosus Strohecker, n. sp. very dark. It is a rather small, short and broad Fig. 121 insect, not very convex and rather strongly and Holotype 6, Colombia: Valle del Cauca, 70 km E closely punctured. Long. 4.5-5 mm." (Arrow). Buenaventura, 1200 ft, 17.11.1970, H. Howden Front tibia of male slightly dilated toward tip, (CMNC). with an acute tooth at distal third of inner edge. Venter, legs, head andantennomeres 1-111bright Some specimens of the type series have the red. Dorsum deep black, marginal bead of prono- dorsum entirely black. A series taken by Borys turn reddish. Pubescence long and gray. Pronotum Malkin (FMNI-I) in NE Matto Grosso shows the narrowed from base, sides little curved. Elytra same range of coloration. In males of this series the highly convex, broad cordiform. Front tibia of male apophysis of the aedeagus forms a sharper angle bowed, widened in distal half. Length 5.8 mm, with the shaft t,han in type examples, but I am width 4.2 mm. reluctant to regard this as a specific differen tiation, This may represent the male sex and mature especially since the Malkin series exhibits some coloration ofA. cll.a,rrt,pionibut until topotypic males variation in this feature. of that species become available I think it better to 82. Anidrytr~scompactus Stroliecker, n. sp. record it as distinct. Fig. 124 85. Anidrytus pilosus Arrow Holotype 6, Peru: Monson Valley, Tingo Maria, 21.X.1924, E. I. Schlinger & E. S. Ross (CASC, Fig. 116 holotype no. 1:3:314). Allotype 3,same data as 11010- Ar~idryti~spilosus Arrow 1920:47. Lectotype 6 ancl a type except 9.XII.54 (CASC). Paratypes: 2d6,Tin- paralectotype Q, S Brazil: Espirito Santo, Schlnidt go Maria, 9.XII.54 (CASC; FSCA);d, Tingo Maria, (NHML). 2000ft. (600 m), 20.X. 1956, J. C. Pallister (AMNI-I); Colombia: Putomayo, Santa Rosa, headwaters Rio Form (male) broadly ovate, strongly convex. San Miguel, 10-23.X.1970, B. Malkin & P. Buchan- Dorsum brownish yellow with long pale yellow an, 266, 2QQ (FMNH; FSCA). pubescence. Pronotum wit,h sides gently curved in front, divergent basad, hind angles acute. Elytra Short-oblong, compact, strongly convex. Ven- broadly cordiform. Length 5.5 mm. ter black, legs clark reddish brown, femoral bases Arrow described the female as narrower t,han fuscous. Antennomeres 1-11 dark red, 3-1 1 black. male with the pronotal sides strongly rounded in Dorsum entirely blaclr, finely and closely punctate, front and parallel toward base. pubescence fine antl gray. Pronotum with sides strongly rounded to front angles, parallel basacl, 86. Anidrytusparallelus Gerstaecker hind angles rectangular, lateral sulci deep, base Fig. 1 17 strongly sinuate. Elytra widened from base to mid- Ar~idr.yt~sparallelus Gerstaecker 1858:266. Holotype 6 dle, gradually narrowed caudacl, apices abruptly (not Q!), Brazil (ZMHB 21820). rounded. Length 6 mm, width 4 mm. E~IL~DLLsdepress~~s Gorham 1875:17. Holotype Q , Bra- zil: Rio de Janeiro (NHML). New synonym. 83. Anidrytus championi (Gorhani) Ar~idrylusdepressus: Arrow 1920:46. Systaechea cl~a.ttlpiot~iGorham 1889:133, pl. 8, fig. 3. Holotype Q, Panama: Volcan de Chiriqui, Champi- Elongate-oblong. Dark redclish brown, thoracic on (NHML). sterna and first sternite fuscous, legs rust-red. Arridrylu~schartlpio~~i Arrow 1920:53. Antennomeres VIII-X darker t'han others. Front tibia of male somewhat widenecl toward apex. Short-oval, subhemispheric. Venter antl legs A very good photograph of the holotype of E. red. Ant,ennomeres I-V red, VI-XI black. Pronoturn depressus has been compared with the single type narrowedfrom base to front with little curvature of ofA, parallelus and shows the two to be much alike. sides, front ancl hind angles acut,e, base bisinuate, The original descriptions are also very similar. The disc bluish black at base, sides and front margin type of A. parallelus is 7 mm long, that of depressus red. Elytra highly convex, bluish black with side 6.3 mm. 174 Volume 11. No. 2. June. 1997, INSECTA MUNDI

87. Anidrytus stenus Strohecker A male in the Janson material (FSCA) agrees Fig. 118 well with the lectotype and has Gorham's label Arridrylus sterit~sStrohecber 1979:716.Holotype a", Bra- "lugubris". Another male is structurally identical to zil: Sta. Catarina, Nova Teutonia (MHNG). the first but has elytral umbo and apex reddish Elongate-oblong. Dark brown with gray pubes- yellow. Front tibia with distal third dilated, flat- cence, pronotal disc vaguely darker. Antenna slen- tened and pubescent beneath. der, antennomeres I-V red, others deep black but tip of antennomere XI pale. Pronotum with sides 91. Anidrytus ephippium Gerstaecker strongly rounded from middle to front angles, a Fig. 103 little convergent basad, hind angles feebly acute or rectangular. Elytra with sides almost exactly par- Ar~idrytusepl~ippiurr~ Gerstaecker 1858:264. Holotype allel from behind umbo to apical third. Front tibia Q, Surinam: Paramaribo, presumably in Dohrn of male evenly and weakly widened distad. Length coll. (ZMPA). 7 mm, width 3.4 mm. Ar~idr.ylus fallaciosus Gorham 1873:47. Holotype Q, French Guiana: Cayenne (NHML);Arrow 1920:47 The elongate form and bizarre aedeagi of this (syn.1. species and A. parallelus indicate divergence of generic rank. Oblong, subparallel. Prosternum, elyt?ralepi- 88. Anidrytus atratulr~sGerstaecker pleura andlegs brownish yellow, meso- andmetast- Fig. 122 ernum fuscous. Antennomeres I-V, head, sides of Ar~idryti~satrat~tltss Gerstaeclier 1858:273.Holotype Q , pronotum ancl scutellum rust-red. Elyt,ra black, S Brazil (ZMHB 21824). shoulder, side margin narrowly atapex extensively red. Front tibia of male clavately widened to tip, Oblong, subparallel. Deep black above with with a slight angulation of inner edge near apex. brown pubescence, elytral shoulder and apex clark Length 6-6.5 mm. red, surface coarsely and densely punctate. Length A pair from Cayenne, det. Gorham (NHML);, 5 mm. Cayenne, dct. Gorham (FSCA ex Janson). A specimen with label "Braz." (FSCA ex Jan- 92. Anidrytus marginatus (Fabricius) son) agrees well with the holotype. This male spec- Er~dorti.yc1ii~srtiargir~atus Fabricius 1801:504. Holotype imen also carries the notation by Gorham "n. sp.? Q,Am(erica) mer(idionalis), Schmid (ZMUC). n.? fallaciosus Gorh." 89. Anidrytus hilaris Gerstaecker Oblong, subparallel. Under surface, legs and elytral epipleura rufous, metasternum darker. Head Ar~idrytt~sl~ilaris Gerstaeclier 1858:271. Holotype Q, S Brazil Zello (ZMHB 21822). and antennomeres I-VII clark red, antennomeres VIII-XI black, antennomere XI apex reddish, an- Oblong, subparallel. Antennomeres IV-XI shin- tennomeres IX-X strongly transverse and mesally ing black, the insect otherwise shining rust-red, acuminate. Pronotum with sides parallel basad, dorsum with yellow pubescence. Length 5 mm. strongly rounded in front, hind angles feebly acute, I have seen no additional examples. disc weakly convex, finely and densely punctate, black, sides and front angles rufous. Elytra about 90. Anidrytus lugubris Gerstaecker 3X as long as pronotum, moderately convex, finely Fig. 104 and thickly punctate, black with umbo, side margin Ar~idrytusZi~gl~bris Gerstaeclier 1858:270.Lectotype Q, narrowly ancl apex extensively rufous. Length 6.3 Colombia (ZMHB 21821). Paralectotypes: Q, Co- mm, width 3.6 mm. lombia (ZMHB);Q, Colombia (NHML). I have seen no other specimens which agree entirely with the holotype, which resembles A. Broadly oblong, subparallel but elytra rather ephippiurn closely but differs markedly in struc- abruptly rounded to apex. Deep black with anten- ture of antenna1 club. nomeres 1-V, elytral epipleura ancl lower face of tibiae brownish yellow; tips of miclclle ancl hind 93. Anidrytus cinctipennis (Gukrin- femora pale yellow. Length 5.2-6mm, witlth 3.4-3.6 Meneville) mm. E(pipocus) cir~ctiper~r~isGuBrin-MQneville 1857:267. Holotype, sex ?, French Guiana: Cayenne, coll. De- jean; repository unknown. INSECTA MUNDI, Vol. 11, No. 2, June, 1997

Described by Gubrin as black, oval, rather black, strongly and rather densely punctate, sides shining. Heatl, sides of pronotum and elytra above red, rather rugosely punctate, red coloration ex- and below, basal half of antenna, and legs tawny panded atfront angles. Elytra more than 3X as long yellow. Pronotum ancl elytra punctate, tomentose. as pronotum, gradually and evenly widened to mid- Length 6 mm, width 3.5 mm. length, andsimilarly narrowedcaudad, apex abrupt- The name E. cir~ctiper~r~iswas treated as a ly rounded; base, sides and apex red, disc black, synonym of A. irlargir~atusby Gerstaecker. Per- densely punctured and with short tawny pubes- haps it should be usecl for the species described by cence. Venter red, sternite 1 ancllegsreddish black. Gerstaecker as A. epliippiuin but lack of a type and Front tibia of male thickened distad but without the brevity of Gubrin's diagnosis make me unwill- tooth; a small sixth sternite is visible. Length 4.5 ing to indicate such synonymy mm, width 2.5 mm. Unusual among the smaller forms of the genus 94. Anidrytus circumcinctr~sStrohecker, n. sp. in its contrasting coloration. Fig. 106 Holotype d, Guatemala: Sac., Reunion, 4000 ft, 96. Anidrytus batesi Strohecker, n. sp. 22.VIII.1947, C. & P. Vaurie (AMNH). Fig. 105 Holotype d, Brazil: Ega (NHML). Parat,ypes:d, Ama- Oblong, subl~arallel.Venter and legs rust-red. zon, Bates (NHML);@,Amazon (FSCA ex Janson). Head, antennomeres I-VI and sides of pronotum red, pronotal disc black. Elytxon black wit,h base, Broaclly oblong, moderately convex. Venter red- umbo, side margin, apex and suture red. Antenna dish black, legs and elytral epipleura rufous. Dor- stout, antennomcres 11-IVsomewhat elongate, sub- sum black, sides of pronotum vaguely reddish, sicle equal, antennomeres V-VIII harclly longer than margins and apex of elytra red, surface finely and wide. Pronotum with sides parallel in basal half, densely punctate, pubescence coppery. Antenna strongly rounded in front, hint1 angles rectangular, rather slender, recl at base, black beyond, antenno- lateral sulci foveatcly wider at base. Elytra nearly meres 111-IV subequal, antennomeres V-VIII secu- 4X as long as pronotum, slight,ly widenedfrom base larly shorter, antennomere IX sharply acuminate in anterior t,hircl, then parallcl for much of their internally but its medial sicle is equal to its distal length, apical rounding rather abrupt. Front tibia width. Pronotum strongly transverse, sides paral- of male clavately wiclened distacl, without distinct lel basad, much rounded in front. Elytra with sides tooth. Length 5.5 mm, wiclth 3.3 mm. subparallel but continuousIy curved, apical round- ing rather blunt. Front tibia of male curved atbase, 95. Anidrytr~sbech,yneorum Strohecker, n. sp. much widened near tip with low angulate tooth Fig. 107 below; hind tibia broadly curved. Length 6.5-7mm, Holotype 6,Venezuela: Aragua, El Limon, 450 m, width 4.5 mm. 6.V.1962, F. Fernanclez. Allotype Q, same data as holotype (IZAV). ParaLypes: 4dd with same data 97. Anidrytris amazonicris Gorhani as holotype; El Limon, 450 111, 25.V. 1977, J. Clavijo, Fig. 101 Q; 25 antl 27.V.1965, 2Q Q, Fernandez; Atridr.ytus an~nzor~icusGorhanl 1873:46, pl. fig. 8. Holo- 22.VII. 1971, Q , Fernanclez; Barinas, Reserva For- type Q, Brazil: Amazonas, Bates (NHML). estal Caparo-Camp Cachicamos, 100 m,dQ,J. Sal- cedo, F. Zambrano; Portugitesa Est,. Exp. San Nico- Broadly oblong, subparallel but elytra with las, 56 km de Guanare, 180 111, I-2.IV. 1968,d, C. J. Rosales. sides continuously curved, rather bluntly rounded to apex. Venter and legs red, hind coxae darker. Small for the genus, outline subparallel. Anten- Antenna1 base red, club black. Pronotum red, sides na black, tip of lastjoint rufescent, antennomere I11 parallel basad, much rounded in front. Scutellum slightly more t,han 2S as long as \vide, antennomer- red. Elytra black with umbo and apex red; side es IV-V each longer than \vide, antennomeres VI- margin may be narrowly red. Front tibia of male VIII quadrat,e. Pronot,um not quit,e 2S as wide as angulately widened near apex, with small tooth long (mid-line), front angles roundctl, sides feebly below. Length (male) 7.8 mm, wiclth 4.8 mm. rounded and weakly convergent to base, hind an- Brazil: Fonte Boa, Bates, d (NHML). gles rectangular, lateral sulci long ancl cleep, disc

INSECTA MUNDI, Vol. 11, No. 2, June, 1997

at anterior end of lateral sulci, finely and rather 5500 ft (1760 m), on fungus, A. D. Mitchell,o", 2Q Q sparsely punctate. Antenna black, antennomere I (FMNH). rufous. Elytra a little wider atbase than pronotum, widened in basal third, thence subparallel, rather 107. Anidrytus quadripunctatr~sGorham bluntly rounded to apes. Length 5.6-7.5 mm. A~lidr~ylr~squadripur~ctatus Gorham 1886: 159. Holotype Q, Brazil: Sta. Catarina, Blumenau, probably in Csiki (1901:35) placecl A. bisigi~ntusin synon- Oberthiir coll. (MNHN); location not confirmed. ymy under A. ju,uei~cus,probably on the basis of a remark by Gerstaecker. Arrow (1920) suggested Oblong or somewhat ovate. Venter red, legs red the conspecificity of A. Disig.~~crlrrsand A. a,l~gcc.stu- wit,h bases of tibiae slightly infuscate. Antenna lus and my stucly of the types su~portshis observa- with first 4 or 5 antennomeres rufous, the others tion. Although I damaged the aecleagus ofA. bisig- black. Dorsum rust-red with coppery pubescence, r~atusin extracting it, I have no doubts about the pronotum with 4 rat,her large round black spots, structural identity of the 2 holotypes. one just medial to front end of each lateral sulcus, 105. Anidrytus trinitatis Stroliecker, 11. sp. the others lateral to and a little in front of discal Fig. 90 pair. Length 7.5-8 mm, width 4.3 mm. Holotype 6, W.I.; Trinidad, Morne Bleu, 2700 ft. (830 I have seen no males. Brazil: Parana, Caviuna, m) 6.VIII.1969, I-I. & A. Howden (CMNC). VI. 1944, A. Mueller, 2 Q Q (AMNH; FSCA). Paratypes:d, Venezuela: Barinas, Reserva Forestal Ticoporo, 230 in, 22-28.V.1968, M. Gelber & J. 108. Anidrytus major Stroliecker, n. sp. Salceclo (IZAV);d,Carabobo, I-Iac. Montera Montal- Fig. 88 ban, 1300 In, 18.IX.1968,J. & B. Bechyn6 (FSCA). Holotype 6, Brazil (NHML).

Broadly oblong, moderately convex. Venter dark Long-oblong, subparallel, elytra rather strong- brown, legs anti elytral el~ipleurareddish brown. ly convex. Venter black, legs and elytral el~ipleura Antenna black, antennomere I red. Dorsum red- rufous, femoral bases dusky. Antenna slender, all dish brown with short, oblique coppery pubescence. basal antennomeres longer than wide, antenno- Pronotum with sides parallel basatl, well roundecl mere IX longer than wide, its medio-distal angle to front angles, hind angles rect,angular, lateral somewhat acute but short; antennomeres I-VI ru- sulci linear, reaching midclle of disc, which is but fous, ot,hers black. Head rufous. Pronotum 2X as little elevated, finely punctale, with a small black wide as long, red with rectangular black patch spot at front end of each sulcus. Elytra about 3X as between lateral sulci, which are long and rather long as pronoturn, sides subparallel but continuosly wide, roundly espanclecl at base; punctation of disc rounded and rather abruptly convergent to apex. rather open, that of sides dense; sides parallel Length 8.2 mm, width 4.9 mm. toward base, well rounded to front angles. Scutel- Very similar in all features to A. u~~guslulus,of lum black. Elytra more than 3X as long as prono- which it may yet prove to be a subspecies. tum, black, with base, side margin and apex red, finely and densely punctate. Front tibia of male 106. Anidrytr~sgr~atenzalae Arrow enlarged distad, with small tooth below. Length 8 Fig. 91 mm, width 4.G mm. Ar~idrytz~sgz~at(?~r~ulc~eArrow1920:46. Holotype 6,Gua- temala: San Joachin (NHML). 109. Anidrytus humilis Gorham Ar~idr.ytusliq~~c>facLus (in part) Gorham 1889: 126. Fig. 111 Ar~idr.ytus? sp. Gorham 1889:128. Al~idrytushurrcilis Gorham 1873:48. Lectotype 6 and paralectotype Q , Nicaragua: Chontales, Janson Long-oblong, elytra about 4X as long as prono- (NHML). tum. Rust-recl, antennomeresVI-XI black. Gorham noted the type specimen as having disc ofpronotum Oblong, subparallel, moclerately convex, Red- infuscate but this may be an artifact. Length 8 mm, dish black with golden pubescence. Antennomeres width 4.5 mm. I-V red, others black. Pronotrum 2X as wide as long, Honduras: Morazon, ridge below La h4anzanita "narrowed from base to middle, and thence more and C. Uyuca, 5 I;m S\Y Suyapa, 5400 ft (1660 m), suddenly t,o the apical angles; hind angles acute...." slab pile, 5.WII. 1948, T. I-I. I-Iubbell, 2 @d(MMZ; Ely tra subparallelfor two thirds their length, grad- FSCA). Guatemala: Zacapa, Sierra de las Minas, 178 Volume 11, No. 2, June, 1997, INSECTA MUNDI ually narrowed to apex. Front tibia of male with recumbent. Antennomeres I-V dark red, others small tooth at distal quarter. Length 7.5 mm. black, antennomeres VII-VIII each longer than wide, antennomeres IX-X strongly transverse, 110. Anidrytus decoratus Arrow medially acuminate. In the specimens from Coroico Fig. 112 the elytral base is broadly red, enclosing a large A~~idryt~~sdepressus Gorham 1889: 128. Holotype 6, black spot. Length 5.7-7.2 mm, width 3.5-4 mm. Guatemala: El Tumbador, Charnpion (NHML). Not In allfeatures, including aedeagus, this species of Gorham 1875. Ar~idrytusdecoratus Arrow 1920:46 (r~onler~t~ov.). is similar to A. hurrreralis. Ar~idrytushu:nerosus is slightly more elongate and antennomere VIII is not Structurally similar, including aedeagus, to A. subglobose as in A. humera.lis and A. rrrargir~a,tus. hu,rnilis. Dorsum black, front angles of pronotum, 113. Anidrytus arrowi Strohecker base and apex of elytra reddish orange. The pale Fig. 109 area of pronotum extends obliquely basad to reach A~~idrytusarrowi Strohecker 1957:269, fig. 31a. Holo- hind angles. The pale areas of elytra are extensive, type 6,Brazil: Espirito Santo, Staudinger (HNHM). that of base encloses a large black spot. Length 7.5 mm, width 4.2 mm. Oblong, subparallel, moderately convex. Upper Guatemala: Zapate, Escuintla, 8.VII. 1948, R. surface ferruginous, disc of pronotum and elytra D. Mitchell, Q (FhfNI-I); Moca, 31 .VIII. 1947, C. & vaguely infuscate. Antennomeres I-V rufous, oth- P. Vaurie, 2 Q Q (AnfNI-I; FSCA).; label indecipher- ers black. Length 6.1 mm, width 3.3. mm. able, 6 (SMTD). 114. Anidrytus latus (Chevrolat) 111. Anidrytr~shumeralis Arrow Fig. 108 Fig. 110 Lycoperdir~ulata Chevrolat 1844:317, pl. 50, fig. 11. Ar~idryti~sI~(~t~~.eraIis Arrow 1920:47. Lectotype 6,Equa- Ar~idrytuslatr~s: Gerstaecker 1858:265. dor, Bucltley (NHML). Paralectotypes;6, Equador, Buckley; Q , Equatlor, Macas, Bucltley; Q , I have seen no type material and my determina- Colombia; Q,Peru (NMML). A syntype 6 is refer- tion of the species is based on the Gerstaecker able to the species nest described. "tvne".- - Rather narrowly oblong and but mocleratly Broaclly oblong, subparallel. Venter ferrugi- convex. Reddish brown, sides of pronotum, elytral nous. Antenna ratherstout, ant,ennomereVIII short, apex, legs and venter rufous, meso- and metaster- transverse, antennomeres IX-X strongly transverse num darker. Dorsum with short, rather sparse and medially acuminate, antennomeres I-Vrufous, coppery pubescence. Antenna fairly stout, anten- the others black. Pronotum black, sides reddish. nomeres VI-VIII subglobose, antennomeres I-V Elytra black, umbo and apex rufous. Front tibia of rufous, others black. Pronot,um with sides strongly male widened distad, wit,h avery small tooth before rounded to the short front angles, parallel basad to apex. Length 5.5-7 mm, width 3-4 mm. the rectangular hind angles, lateral sulci wide, Peru: Loreto, Estiron, Rio Ampiyacu, 13.XI- rather short. Front tibia of male gradually widened 9.XII. 1961, 13. Malkin, U(FhfN1-I). from base, with an angular expansion at apical Very similar to A. rncrrgirlcrtzrs (Fabr.). third, hind tibia incurved at apex. Length 6-7 mm, 112. Anidrytus h,umerosr~sStrohecker, 11. sp. width 3.1-3.6 mm. Fig. 113 A male from Rio de Janeiro (FSCA) conforms Holotype 6 ancl allotype Q,Peru: Loreto, Estiron, Rio closely to the Gerstaecker "type" but has ent,ire Ampiyacu, 13.XI-9.XII.1961, B. Malkin (FMNH). prothorax, meso- and metasternum and first ster- Paratypes: 366, 3 Q Q with same data as holotype; nite quite black, legs and elytral epipleura rufous. 266, Loreto, 20 km from Ucayali on Rio Calle,ria, Colonia Galleria, 10.IX-15.X. 196 1, B. Malkin 115. Anidrytus pardalinis Strollecker (FMNH; FSCA);@, Peru (NHML). Bolivia: Coroico, Fig. 131 6 Q (FSCA) . Ar~idrytuspardali~~us Strohecker 1943:388, fig. 12f, g. Holotype 6, Costa Rica, F. Nevermann (USNM Oblong, subparallel. Venter ancl legs brown. 55888). Dorsum blaclc with head, front angles ofpronotum, elytral umbo amcl apex recl; pubescence fine, gray, INSECTA MUNDI, Vol. 11, No. 2, June, 1997

Oblong or somewhat oval, rather strongly con- 118. Anidrytus nigropiceus Gorham vex. General color deep rust-red, dorsum with Fig. 115 short, declivent, coppery pubescence. Pronotum with 2 large quadrate black patches on base. Elytra At~idrytusnigropiceus Gorham 1889: 126. Lectotype 0' with base, most of suture and 2 round spots black, and 1 paralectotype 6,Panama: Bugaba, Champion the anterior spot near mid-length, the posterior (NHML). spot smaller ancl on apical slope. Front tibia of male gradually widened in distalhalf, with a sharp tooth Broadly oblong, subparallel. Dark brown be- on inner edge. Length 8 mm, width 4 mm. low. Dorsum black. Antennomeres I-Vand margins of pronotum and elytra reddish black. Lateral sulci 116. Anidrytus grandis Strohecker, n. sp. of pronotum very deep, almost reaching middle of Fig. 132 disc. Front tibia of male angulately widened beyond Holotype d,French Guiana: Cayenne, with Gorham's middle. Length 7 mm, width 4.6 mm. label A. tr~argil~alz~s(FSCA ex Janson). 119. Anidrytus fuliginosus (GuQrin- Long-oval, rather stxongly convex. Meso- and MQneville),new combination metasternum and first 2 sternites black; other E(pipocus) fuligir~osusGuBrin-MQneville 1857:266. Type parts of venter, including legs and epipleura ru- (sex ?), Colombia: Cartagena, Dejean Coll. fous. Dorsum black wit,h sicles of pronotum broadly, elytral base, side margin and apex narrowly red. Bocly blackish brown, antenna, legs, elytral Antennomeres 111-V subequal, each hardly 2X as epipleura, sides of sterna and abdomen paler brown. long as wide, antennomeres VI-VIII similar to each Head, pronotum and elytra black, a little shining, other, slightly shorter than antennomere IV, an- punctured, a little pubescent. Length 6 mm, width tennomere IS about as wide at apes as long, its 3.66 mm. Very close to (A. nigricar~s)in form. mediodistal angle very short,. Pronot,um relat>ively I have not recognized this insect among the small, 2X as \vide as long, sides parallel basad, material examined. It can hardly be the same as the rounclecl to obtuse front, angles, marginal bead specimens described by Gerstaecker as El)il)ocu.s rather wide in front,, evanescent at rectangular fuligil~osus,for which a new name has been pro- hind angles, lateral sulci long ancl wide, slightly posed (Strohecker 1977:323). convergent. Elytra 4X as long as pronotum, wid- 120. Anidrytus spadicer~s(Gukrin-MQneville) ened from base in anterior third thence gradually narrowed caudad, densely andsubrugosely. Length E(pipocus) spadiceus GuBrin-MBneville 1857:266. Type 8 mm, width 4.7 mm. (sex ?), Colombia, Dejean coll. Anidrytus spadiceus: Csiki 1901:35. 117. Anidrytus cardisoma Strohecker, n. sp. Fig. 114 Body black, mouthparts, legs and elytral epi- Holotype 6,Bolivia: Coroico (FSCA). pleura yellowish brown. Pronotum and elytra very finely punctured, pubescent. Length 5.5mm, width Oval, moderately convex, elytra weakly flat- 3.25 mm. Very close to A. fz~ligi7~osu,s,~)ossibly a tened. Sides of pronoturn, elytral e~ipleura,legs variety. and distal sternites rust-red. Thoracic sterna and This brief diagnosis has not been sufficient for first sternite black. Dorsum generally black, head me to identify specimens as this species. and sides of pronotum reddish. Antenna long, an- tennomeres I-IVdark red, antennomeresv-XI black, Acknowledgnlents anntennomeres 111-IV subequal, each more than I (Paul E. Skelley) thank R. H. Arnett, M. C. 2X as long as wide, ant,ennomeres V-VIII secularly Thomas, R. E. Woodruff, Florida State Collection of shorter but each longer than wide. Pronotum par- Arthropods; R. A. B. Leschen, Michigan State Uni- allel to rect,angular hint1 angles, rounded to obtuse versity; and S. A. Slipinski, Polish Academy of front angles, lateral sulci, reaching midclle of clisc. Sciences, for their valuable editorial comments. Elytra 3Xas long as pronoturn, cordiform, widest in Florida Department of Agri~ult~ureand Consumer basal third, rather rapiclly narrowed t,o apex. Front Services, Division of Plant Industry, Entomology tibia of male gradually but not greatly widened Contribution No. 840. distad, hind tibia weakly curved. Length 7 mm, width 4.3 mm. 180 Volume 11, No. 2, June, 1997, INSECTA MUNDI

References GuBrin-Mkneville, F. E. 1857. MatQriaux pour une Arrow, G. J. 1920. A contribution to the classification monographie des ColQoptBresdu groupe des Eumor- of coleopt,erous family Endomychidae. Transactions phides et plus specialement du genre Eurr~orphus. of the Entomological Society of London 1920: 1-83, 1 Archives entomologiques I:237-280, 1 p1. PI. Kirsch, T. 1876. Beitrage zur Icenntnis der peruanis- Chapuis, F. 1876. IILJ. T. Lacordaire. Histoire naturel- chen Kaferfsuna, auf Dr. Abendroth's Sammlung le des Insectes, Genera tles ColQoptBres, Vol. XII. basirt, Sechtes Stuck. Deutsche Entonloligische Paris, 374 pp. Zeitschrift XX:81-133. Chevrolat, L. A. A. 1844a. IILF. E. GuBrin-MQneville. Olivier, A. G. 1791-1792. I~LEncyclopQdie MQthodique. Iconographic du RBgne Animal cle Cuvier. Vol. 7. Dictionnaire des Insectes. Vol. VI. Paris. Paris, 576 pp., 104 111s. Olivier, A. G. 1808. Entomologie ou histoire naturelle Chevrolat, L. A. A. 1844b. Dictionnaire universe1 des Insectes. Pars VI. Paris, pp. 613-1 104, 41 pls. $Histoire naturelle. Vol. V. Perty, J. A. M. 1832. Delectus animalium articula- Csiki, E. 1901. Catalogus Endomychidarum. Ter- torum, quae in itinere per Brasiliam-collegerunt m6szetrAjzi Fuzetelr XXIV: 1-53. Spix et Martius. Fasc. 2:61-124, 40 pls. Erichson,W. F. 1847. Conspectus insectorum coleopter- Pic, M. 1931. NouveautBs diverses. Melanges Exotico- orum quae in Republics Peruana observata sunt. entomologiques 58. Moulins, 36 pp. Archiv fiir Naturgeschichte XIII:67-185. Strohecker, H. F. 1943. Some fungus beetles of the Fabricius, J. C. 1801. Systema eleut,heratorum. Vol. I, family Endolnychidae in the U.S. National Muse- I

Figs. 1-16. Elytral pattern. 1) Epopterus fusciutus fusciutus - lectotype, 2) E. fusciutus urtgustutus - holotype,3) E. zebra - holotype, 4) E. sirnilis - holotype, 5) E. sirrlilis - Ecuador, 6) E. uuriegutus - Peru:Callanga, 7) E. uuriegutus - Amazon, 8) E. reticr~lutus- holotype, 9) E. cyclicus - holotype, 10) E. cordutris - holotn~e,11) E. deltoider~s- holotype, 12) E, decerr~pr~~r~ctatr~s- Trinidad, 13) E. decerr~pur~ctutns - Colombia: Cucuta, 14) E. ? uicirtrrs - Peru: Cnllanga, 15) 15. obliqur~s- holotype, 16) E. rtotatus - holotype. 182 Volume 11, No. 2, June, 1997, INSECTA MUNDI

Figs. 17-32. Elytrnlpattern. 17) Epopterus llistrio- topotype, 18) E. rubigitrosus- holotype, 19) E.parvus-paratype, 20) E. quaesitus - holotype, 21) E. fctllax - holotype, 22) E.pictussensu Gerstaecker - Brazil, 23) E. bioculatus - holotype, 24) E. ocellatussensu Gerstaecker -Brazil,25) E. dilectus - holotype, 26) E. vertlicutus -holotype, 27) E. quecltuattus -holotype, 28) E.picticollis - holotype, 29) E. sigt~aticollis - Panama, 30) E. sigrtctticollis - Colombia, 31) E. testudittarius - Colombia, 32) E. grwilis - holotype. INSECTA MUNDI, Vol. 11, No. 2, June, 1997 183

Figs. 33-57.Aedeagus, clestro-ventral aspect. 33) Epopterusocellatussensu Gerstaecker -Brazil: Corupa, 34) E. ar~t~ulatt~s-Guyana, 35) E. partitus pcrrl,ilus - Honcluras, 3G) E. partitus kirscl~i- holotype, 37) E. tigrirrus - holotype, 38) E. gerrlirrus - holrtype, 39) E. sigtraticollis - Colombia: Villavicetlcio, 40) E. vact~us- holotype, 41) E. ut~dulatus-Argentina: TucumBn, 42) E. pictus sensu Gerstaecker - Brazil, 43) E. vcrriegc~tus- holotype, 44) E. cott~ptus- topotype (syntype?), 45) E. testudirturius - lectotype, 46) E. fasciatus fasciatus - lectotype, 47) E. egcrrlus - lectot~l~e,48) E. ct~c~~llirrus- holotype, 49) E. dilectus - holotype, 50) E. dives - holotype, 51) E. clara - Brazil: ParB, 52) E. Iiistrio - Brazil: AmapB, 53) E. decetr~pt~.r~ctatus- Colomnbia: Puerto Berrio, 54) E. ephippiger - holotype, 55) E. bioculattrs - holotype, 56) E. rtigrocitrctt~s- holot~l~e,57) E, trarrsversns - holotype. 184 Volume 11, No. 2, June, 1997, INSECTA MUNDI

Figs. 58-78. Aetlengus, dextro-ventral nspect. 58) Epopterus cordatus - holotype, 59) E. rcigrocircctus - Colombia: Leticia, 60) E. flauolircecrtus - holotype, 61) Endeltoideus - holotype, 62) E. cyclicus - holotype, 63) E. siniilis - holotype, 64) E. ntinieticus - holotype, 65) E. flauorcotatus - holotype, 66) E. reticulatus - holotype, 67) E. lorzgus - holotype, 68) E. batesi - holotype, 69) E. zebra - holotype, 70) E. rligerrincus - holotype, 71) E. rtotatus - holotype, 72) E. obliquus - holotype, 73) E. paruus - holotype, 74) E. gracilis - holotype, 75) E. queclcuarcus - holotype, 76) E. picticollis - holotype, 77) E. corcfusus - holotype, 78) E. crypticus - holotype. INSECTA MUNDI, Vol. 11, No. 2, June, 1997 185

Figs. 79-87. Aetleagus, dextro-vental aspect. 79) Epopterus 2 verrticutr~s- Brazil: Rio de Janeiro, 80) E. ntriverttris - holotype, 81) E. loretertsis - holotype, 81)E. cccrllericrr~r~s- holotype, 83) E. ur~or~zulr~s- holotype, 84) E. pnrttherirrr~s- Panama, 85) E. aravacrLs - holotype, 86) E. decer~z~~ur~ctc~lrrs- lectotype, 87) E. subr~~ucrrlntus- holotype. 186 Volume 11, No. 2, June, 1997, INSECTA MUNDI

Figs. 88-10G. Aedeagus, destro-ventral aspect. 88) Ar~idrytusnrcIjor - holotype, 89) A. juvertcus ? - holotype of A. ar~gr~stulus,90) A. trirlitcrtis - halotype, 91) A. guaterrlcrlc~- holotype, 92) A. bipr~rlctatus- holotype, 93) A. plcrgiutr~s- holotype, 94) A. rtinlbiferr~s- holotype, 95) A. liqr~efc~tus,9G) A. trrexiccrrirts - holotype, 97) A. corctructus - Costa Rica: Turrialba, 98) A. ovatulus - lectotype, 99) A. rligricarls - Rio de Janeiro, 100) A. kirsclti - holotype, 101) A. urrluzor~icus- Brazil: Fonte Boa, 102) A. cupreutus - holotype, 103) A. ephippiurn - French Guinna, det. Gorhnm, 104) A. lrtgubris - no locality, det. Gorham, 105) A. batesi - holotype, 106) A. circurncinctus - holotype. INSECTA MUNDI, Vol. 11, No. 2, June, 1997 187

Figs. 107-127. Aetleagus, dextro-ventral aspect. 107) Artidrytus becl~yr~eorurr~- holotype, 108) A. latus - Gerstaecker "type", 109) A. arrotoi - holotype, 110) A. lrrrrr~erc~li~- lectotype, 111) A. 11~11tilis- lectotype, 112) A. decorcztt~s- holotype, 113) A. Iturrterosus - holotype, 114) A. cardisorr~n- holotype, 115) A. rligropiceus - lectotype, 116) A. pilosus - lectotype, 117) A. parallelus - holotype, 118) A. stertus - holotype, 119) A. Itelvolus - Venezuela, comp, lectotype, 120) A. cor~color- Peru, comp. holotype, 121) A. gibbosus - holotype, 122) A. atruttrlus - Brazil, comp. holotyl~e,123) A. ~ILSCILS - lectotype, 124) A. corrt~~c~tus- holotype, 125) A. glaber - Peru, comp. holotype, 126) A. cotasobrir~us- holotype, 127) A. tertnipes - holotype. 188 Volume 11, No. 2, June, 1997, INSECTA MUNDI

Figs. 128-134. Aetleagus, tlestro-ventral aspect. 128) Ailidrytus urzicolor - Peru, comp. holotype, 129) A. porcus - holotype, 130) A. gerstueclter~i- lectotyl~e,131) A. pcrl.dalitic~.s- holotype, 132) A. grurldis - holotype, 133) A. parki - holotype, 134) A. oariegntus - Rio de Janeiro.