An Illustrated Descriptive Catalogue of the Genera Anidrytus Gerstaecker and Epopterus Chevrolat, with Descriptions of New Species (Coleoptera: Endomychidae)

158 Volume 11, No.2, June, 1997, INSECTA MUNDI

An illustrated descriptive catalogue of the genera Anidrytus Gerstaecker and Epopterus Chevrolat, with descriptions of new species (Coleoptera: Endomychidae)

H. F. Stroheckerl Research Associate, Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services 1911 SW 34th St. Gainesville, FL 32614-7100 U.S.A.

Abstract: The endomychid beetle genera Anidrytus Gerstaecker and Epopterus Chevrolat have been confused for many years. This paper discusses the similarities and differences of these genera, describes new species, makes some nomenclatural changes, and illustrates the genitalia for many species. New species described: Epopterus picticollis, E. biocu/atus, E. confusus, E. grad/is, E. /oretensis, E. alloma/us, E. arauacus, E. atriuelltris, E. crypticus, E. queclwanus, E. callerianus, E. /Zauollotatus, E. sublll(Jcu/atus, E. paruus, Allidrytus nimbiferus, A. pm'hi, A. compactus, A. gibbosus, A. circumdnctus, A. bechYlleorum, A. balesi, A. me:'dccmus, A. trinitatis, A. major, A. humerosus, A. grandis, A. cardiosoma. New synonymies: E. UCtC/UISGerstaecker (= E. scaloris Gorham); E. decempunctatus Gerstaecker (= E. bifasdatus Pic); E. uariegatus Erichson (= E. decomtus Kirsch); A. cOlltmctus Gerstaeclcer (= A. dolosus Gorham); A. he/uolus Gerstaecker (= Ephebus ignobilis Gorham); A. pamllelus Gerstaecker (= Ephebus depressus Gorham). Changes in nomenclatural status or combination: Epopterus hirshi Strohecker is reduced to a subspecies of E. partitus Gerstaecker; E. angustatus Strohecker is reduced to a subspecies of E. fasdatus (Fabricius); E. fuligl:/IOSUS Guerin-Meneville is transferred to Allidrytus.

The genera Anidrytus Gerstaecker andEpopter Mostprevious descriptions have been drawn on us Chevrolat share many common features, among body form and color pattern. Many similarities in them: body form ovate to oblong; dorsum pubes- these features are found among the species. Yet, cent, but much abraded in many specimens; anten- strong differences in color pattern are shown by nomeres III & IV subequaJ; antennomeres IX & X, specimens of Epopterus which, on the basis of especially X, mesally acuminate; spinasternum aedeagal structure, are conspecific. Some of the horizontal, extended beyond front coxae, broadly observed variation is due, no doubt, to differences spatulate with apex subtruncate. Males have protib- in maturity, but may also exist as color phases in ia curved, enlarged in apical third, often with a some species. I have used trinomials only where small tooth below; and abdominal sternite VI is names have already been published. visible. I consider the aedeagus the most reliable taxo- Gerstaecker (1858) distinguished the two gen- nomic character. This organ is strongly sclerotized, era on the basis of mandibular and maxillary struc- often asymmetrical and bizarre in form, which ture: mandibular apex in Anidrytus is entire, in differs in appearance with change of position, ei- Epopterus it is bifid; maxillary galea is corneous at ther by rotation or inclination. The camera lucida base in Anidrytus, in Epopterus it is entirely mem- drawings are of dextro-ventral aspect, the lesser branous. I doubt that Gerstaecker subjected all the curvature of the base is considered ventral. species which he treated to much analysis but, Study of the aedeagi suggests relationships rather, fell into the habit of assigning plainly col- between species which transgress the nominal ge- ored and mostly larger forms to Anidrytus and neric boundaries of these genera and suggests long those of strongly variegated pattern and mostly separated evolutionary lines not reflected in cur- smaller size to Epopterus. Later authors, notably rent nomenclature. Evaluation and action on these Gorham, followed him in this practice. ideas would be a long task, for which material and I have not so tested the many species now time left to me are inadequate. Therefore, I follow known but, after examining the holo- or lectotype of Gerstaecker's and Gorham's concepts in the assign- nearly all described forms, I think that a prelimi- ment of species. nary treatment may enable any future student of Because of interspecific similarities and in- the group to proceed without repeating my efforts, traspecific variations in external features, I found which have extended over many years. it impossible to construct a worthwhile key. I would be loathe to attach identity labels to most females

1 Dr. Strohecker finished this paper before his death, November 14, 1988. The manuscript has been changed little from its original form, and lacks certain details generally required for modern systematic papers and by Illsecta MUlldi. Even in its current state, it represents a step forward in the study of these genera and is here published for that reason. Paul E. Skelley, Florida State Collection of Arthropods, Florida Department of Agriculture & Consumer Services, Division of Plant Industry, 1911 SW 34th St., Gainesville, FL 32614·7100, U.S.A. INSECTA MUNDI, Vol. 11, No.2, June, 1997 159 unless in series 'with males, and I believe the most for the following nominal species: E. myops Guerin- reliable path to identification of males is by compar- Meneville and E. trimacu1atus Guerin-Meneville, ing figures of the aedeagi with the specimen. Each which I have been unable to locate; E. geniculatus figure is followed by a number in parentheses Pic, which is compared with E. picticollis n. sp.; E. which indicates the species number in the text. ryei Gorham, E. lineoguttatus Gorham, E. pan Material examined for this study is found in the therinus Gorham, and E. ocellatus maculosus following institu tions, the officers of which I thank Gorham. Habitus figures of the last four accompa- for their cooperation: American Museum of Natu- ny the original descriptions. ral History, New York (AMNI-I); California Acade- my of Sciences, San Francisco (CASC); Canadian 1. Epopterlls ocellatlls (Olivier) Museum of Nature, Ottawa (CMNC); Deutsches Fig. 24, 33 Entomoligisches Institut, Eberswalde Finow Erotylus ocellatus Olivier 1792:437. Type repository (DEI C); Field Museum of Natural History, Chicago unknown. Eumorphus ocellatus (Olivier) 1808: 1067, pI. 1, fig. 6. (FMNH); Florida State Collection of Arthropods, Epoptems ocellatus Gerstaecker 1858:275, pI. 3, fig. Gainesville (FSCA); Hungarian Natural History 43u, x; Strohecker 1953:pl. 3, fig. 30. Museum, Budapest (HNHM); Funcacion e Institu- to Miguel Lillo, Tucuman (IMLA); Instituto de Oblong, rather flat above, almost equally round- Zoologia Agricola, Universidad Central de Venezu- ed in front and in rear. Legs, head and antennal ela, Maracay (IZAV); Entomologisches Institut, base reddish yellow. Venter and pronotum reddish Museum G. Frey, Tutzing (MGF'!'); Museum brown, the latter with 4 black spots in a transverse d'Histoire Naturelle, Geneva (MHNG); Museum row. Elytron blackish brown with suture reddish. A National d'Histoire Naturelle, Paris (MNHN); Nat- yellow band on base of elytron encloses a dark ural History Museum, London (NHML); Staatli- humeral spot and a large dark ocellus, almost or ches Museum fur Tierkunde, Dresden (SMTD); U. quite touching base. A yellow pre-apical band emits S. National Museum, Washington (USNM); Muse- a short ray at middle of its front edge and 2 long um fur Naturkunde der Humboldt UniversiUit, posterior rays, one along margin, other near su- Berlin (ZMHB); Museum & Institute of Zoology, ture; it may form 2 complete rings enclosing dark Polish Academy of Sciences, Warsaw (ZMPA); Zoo- spots. Front tibia of male abruptly widened beyond 10gicalMuseum, University of Copenhagen, Copen- middle. Length 6-6.5 mm, width 3.4-3.6 mm. hagen (ZMUC); Zoologische Staatssammlung, Mu- Most specimens examined are labeled "Brazil". nich (ZSM C). Brazil: Sta. Catarina, Corupa, XII. 1944, A. Muller (AMNH). Genus Epopterlls Chevrolat Chevrolat 1844::380; Erichson 1847: 181; Gerstaecker 2. Epopterlls annlllatlls Guerin-Mimeville 1858:274; Chapuis 1876:123; Gorham 1890:129; Fig. 34 Strohecker 1953:62. Epoptems annulatus Guerin-Meneville 1857:268, pI. Type species: Erotylus ocellatus Olivier (Strohecker 13., fig. 20. Holotype 0', Brazil (NHML). 1953:63). Very similar to and probably a color form of E. Most species :3.5 - 5 mm long, although some ocellatus, as noted by Csiki (1901 :47). The figure in may slightly exceed 6 mm. Body outline broadly Guerin-Meneville does not properly show the long cordate to oblong and subparallel. Elytra variegat- caudal rays of posterior band. ed with black or brown and yellow or red markings, the elytral "Guyana", 0' (FSCA). often with a broad clark band across middle. In two species, E. lineoguUaf,us Gorham and E. clara 3. Epopterlls myops Guerin-Meneville Gorham, the pale elytra are marked with black points or streaks. The dorsum is pubescent, but E(poptems) myops Guerin-Meneville 1857:268. Type often much abraded in museum material. material from Brazil?, Colombia?; repository un- Diagrams of color pattern only are shown for known. four species described from unique female types: E. Epopterus ocellatus myops Csiki 1901:47. vicinus Pic, E. fallax Gerstaecker, E. rubiginosus Gerstaecker, and E. qlwesitus Gorham. All figures Originally described as tawny brown, elytra of elytra are intended to show only pattern; any very dark, pronotum lighter, venter and legs yel- realism is fortuitous. No illustrations are offered lowish, femora briefly fuscous at base. Pronotum 160 Volume 11, No.2, June, 1997, INSECTA MUNDI with 4 black spots. Elytra finely punctate, black at 7. Epopterus partitus Idrschi Strohecker, base and apex, with an anterior yellow band form- new status ing a circle near the scutellum and another pale Fig.3G band at apical third. A smaller variety. Epopterus hirschi Strohecker 1959: 181, fig. 8. Holotype cf', Ecuador: Balzapamba (ZMHB). 4. Epopterus trimaculatus Guerin-Mi:meville More intensively colored than nominate form; Epopterus trimaculatus Guerin-Meneville 1857:268. head, legs, middle of metasternum and first stern- Type material from Colombia; repository unknown. ite black. Pronotum rufous with 4 black spots. Elytron with pale areas reduced to rather narrow Original description as follows: Body and elytra bands, apex yellow. The aedeagus differs little from black, shining, pronotum with anterior border yel- that of the nominate form. Length G-G.5 mm, width low. All antennomeres yellow, blackish at base. 3.5-3.7 mm. Legs black, knees and tarsi yellow. Elytron with a Colombia: Valle de Cauca, Anchicaya Dam, 70 large posterior yellow area, which is dentate on km E Buenaventura, 22.VII.1970, H. & A. Howden, front edge and extends almost to apex, enclosing 3 2cf'cf' (CMNC; FSCA). black spots, the middle one largest. Length 5.33 mm, width :3 mm. 8. Epopterus tigrinus Gerstaecker Fig. 37 5. Epopterus partitus partitus Gerstaecker Epopterus tigrinus Gerstaecker 1858:279. Holotype cf', Fig. :35 Brazil (ZMHB).

Epopterus partitus Gerstaecker 1858:277. Lectotype, Oblong, rather flat above. Venter, bases of Mexico: Yucatan (NHML). A syntype Q is presum- femora tibiae black. Pronotum black, sides ably in ZMPA. and yellow with a small black spot near lateral margin. Oblong, elytra more gradually rounded caudad Elytron black, with an irregular yellow or orange than in E. ocellatus. Venter reddish yellow; thorac- band across base enclosing 2 dark spots. A broad ic sterna and legs may be dusky. Antennal base and apical pale band encloses a lateral dark spot and a pronotum reddish yellow, the latter with 4 black semicircular black spot on suture. Length 5.7-G.5 spots in a transverse row. Elytron reddish yellow mm, width 3-3.5 mm. with 2 black spots on base, smaller on umbo, larger Brazil: Sta. Catarina, Lagos, Fruhstorfer near scu tellu m and narrow ly reaching elytral base; (ZMHB; FSCA); Nova Teutonia, F. Plaumann (GMFT; FSCA); Parana (FSCA). wiele black band across middle, U -excavate in front and V-excised at middle of hind edge. Black color extending forward along suture almost to scutel- 9. Epopterus geminus Gerstaecker lum and also caudad, where it widens into a semi- Fig. 38 circular spot; lateral is an oval black spot. Length G- Epopterus geminus Gerstaecker 1858:279. Lectotype cf', G.8 mm, width 3.5-4 mm. Colombia (ZMHB). 6. Epopterus maculosus partitus Gorham Venter and legs black. Head and antennal base reddish yellow. Pronotum with front angles and Epopterus ocellatus maculosus Gorham 1890:129, pI. VII, fig. 19. sides yellow, disc with large trapezoidal black patch; Epopterus partitus maculosus: Arrow 1920: 48. Lecto- small black spot in pale area of each side. Elytron type cf', Nicaragua: Chontales, Janson, B. C. A., Col. mostly black; an irregular orange band across base VII, E. ocellatus 01. (NHML). encloses 2 black marks which reach base. A yellow area at apex encloses 2 oval black spots. Length 5.5 Differs from nominate form only in having the - G mm, width :3.3 mm. broad black elytral band interrupted at middle by a Venezuela: Caracas, Berg Avila, P. C. Vogl pale vitta, which, however, may be incomplete. (ZMHB; FSCA). Colombia: Santander del Norte, Reported by Gorham from Belize; Guatemala; Santiago, 700 m, 23.V.1974, H. & A. Howden, 1 Nicaragua; Mexico: Tabasco. A considerable series pair; 30 km S Cucu ta, 8. V. 1974, Howelens, cf' (CMNC; of paralectotypes are in NHML. FSCA). INSECTA MUNDI, Vol. 11, No.2, June, 1997 161

10. Epopterus signaticollis Gerstaecker 12. Epopterus testudinarius Gorham Fig. 29, 30, 39 Fig. 31, 45

Epopterus signaticollis Gerstaecker 1858:280. Holotype Epopterus testudinarius Gorham 1873:48, pI., fig. 7. cJ', Colombia (ZMHB). Lectotype cJ',paralectotype Q, Brazil: Amazon, Bates Epopterus scalaris (in part) Gorham 1890: 130. (NHML).

Oblong, venter yellow, hind part of metaster- Oblong. Antenna, legs, and venter black. Head num and first sternite black, sternites 2-4 dusky at and pronotum yellow, latter with large black patch middle. Head and antennal base yellow. Pronotum which may extend across entire base, abruptly yellow, middle of fron t margin and 4 discal spots narrowed at middle, then continues to front mar- black. Elytron reddish yellow with a small dark gin. Scutellum black. Elytron yellow with small spot on umbo, an oval dark spot at mid-line of base, oblong black mark on shoulder, smaller round a broad multidentate dark band across middle and black spot near scutellum, and broad, irregularly 2 black spots before apex. Length 5.5 mm, width 3.5 margined black band across middle. Apex black, mm. between it and middle black band a deeply con- The description above was extracted from Ger- stricted yellow band. In a male from Villavicencio staecker. A male from Colombia: Villavicencio, has the pronotum is wholly black, the entire side mar- the inner front spot narrowly connected to base, gin of elytron yellow. Length 6-6.5 mm, width 3.6- and the middle black band interrupted at the side. 3.8mm. A male from Panama: Bugaba (FSCA ex Janson) Colombia: Meta, Villavicencio, 13.VIII.1938, has venter and legs en tirely black. C.H Seevers (FSCA). Colombia: Cundinamarca, Monterreclondo, Schneble, 1 pair (GMFT); Meta, Villavicencio, 13. Epopterus comptus Gorham 9.VIIL1938, C. H. Seevers; cJ'(FSCA); Antioquia, Fig. 44 Epopterns comptns Puerto Berrio, VIII. 1938. Seevers, cJ'(FSCA). Pan- Gorham 1890: 130, pI. 7, fig. 200. Lectotype cJ', 4 paralectotypes, Nicaragua: Chon- ama: Bugaba, 250-400 m, Champion, cJ'(FSCA ex tales, Janson, Belt (NHML). Janson). Oblong. Venter black, paler caudad, legs yel- 11. Epopterus vaCUllS Gerstaecker low. Antenna yellow at base, antennomeres VI-XI Fig. 40 black. Head and pronotum pitchy reel, latter with basal 3/5 of side margin, 2 anterior and3 elongate Epopterus vacl/,US Gerstaecker 1858:283. Holotype cJ', basal spots black. Elytra black with umbo yellow, French Guiana: Cayenne (MNHN). pale color extending to suture as a ring enclosing Epopterus scalaris Gorham 1890: 130, pI. VII, fig. 20. round black spot. Elytral apex extensively yellow Lectotype cJ', Panama: Bugaba, Champion (NHML). with large dark spot enclosed. Length 5.8-6 mm, New synonym. width 3.5 mm. Nicaragua: Chon tales , cJ' probably from origi- Venter, femora and tibial bases black. Antenna nal collection; Q without locality data (FSCA ex stout, black, basal antennomeres (except III) hard- Janson). ly longer than wide. Head dusky yellow. Pronotum with sides yellow, disc black, the dark color abrupt- 14. Epopterus llndulatus Guerin-Meneville ly narrowed at mid-length and continued narrowly Fig. 41 forward to front margin. Elytron black with a large Epopterus undulatus Guerin-Meneville 1857:269, pI. yellow patch on shoulder enclosing a small black 13, fig. 2l. Type material from Argentina: Buenos spot. The pale color does not reach suture, and its Aires, Dejean colI.; repository unknown. hind edge is tridentate. Apex yellow, front edge of this pale area tridentate. Length 6 mm. Oblong. Venter black, abdomen paler caudad. Although Gerstaecker cited a single female in Legs yellow, basal half of middle and hind femora the Thomson collection, the presumptive type is a and of all tibiae black. Antennal base and head male; the front tibia is less strongly modified than yellow. Pronotum yellow with the following black in the males of most species of the genus. marks: an inverted V with apex at middle offront margin, its lobate limbs extending to hind angles, a 162 Volume 11, No.2, JUlle, 1997, INSECTA MUNDI small round spot on each side. These markings may especially the posterior one, which runs in undu- be obscured by general darkening. Elytron black lant fashion obliquely caudad and mesad. with narrow yellow band across base, enclosing black humeral spot and elongate parascutellar dark 18. Epopterus nigerrimus Strohecker mark; a narrow zigzag pale band at posterior quar· Fig. 70 ter, and lunate bar close to apex. Length 5-5.8 mm, Epopterus nigerrimus Strohecker 1957:268, fig. 26. width 3-3.2 mm. Holotype 0', Brazil: Goyaz, Jatahy (HNHM). Argentina: Tucuman, Tucuman, Horco Molle, San Pedro de Colabao, Choromoro; Jujuy: Yuto; Narrowly oblong, subparallel. Deep black below and above, dorsum thickly punctate. Elytron Catamarca, EI Alto. Bolivia: Sta. Cruz, EI Cidral (IMLA). Argentina: Cordoba, Jesus Maria (GMFT). with 2 narrow red marks, the anterior Y-shaped, Paraguay: Asuncion (HNHM). embracing the umbo, the posterior near apical third, transverse and undulate. Length 4.6-4.8 15. Epopterus zebra Strohecker mm, width 2.7 mm. Fig. 3, 69 Argentina: Cordoba, 20'0', 9 (ZSMC; FSCA). Epopterus zebra Strohecker 1957:269, fig. 27. Holotype 0', Peru (HNHM). 19. Epopterus bioculatus Strohecker, ll. sp. Fig. 23, 55 Oblong. Upper surface reddish yellow. Prono- tum with a W-shaped mark and median basal spot Holotype 0', French Guiana: Cayenne (FSCA ex Jan- son). black. Elytron marked with black as follows: 2 round spots on base, an undulate band before Long-oval, elytra gradually narrowed from middle, another band just behind middle, and 2 an terior third. Ven ter an d femoral bases dark brown. large coalescent spots before apex. Length 5.9 mm, Antennal base yellow, club black. Dorsum blackish width 3.3 mm. brown, elytron with anterior and posterior yellow 16. Epopterus picticollis Strohecker, ll. sp. rings. Pronotum with sides subparallel basad, fair- Fig. 28, 76 ly strongly curved to fron t angles, hind angles very sharp, disc finely, sides coarsely and densely punc- Holotype 0', Peru: Huanuco, 25 km below Carpish, 2500 tate. Elytra more than 3X as long as pronotum. The m, 9.IX.1946, F. Woytkowski, ancl allotype 9 with anterior yellow area leaves a large dark humeral same data except 8.IX (AMNH). Paratypes: 299, spot and completely encloses an oval dark spot near same locality, IX.1948 (FSCA). scutellum. The posterior yellow band encloses a Oblong, subparallel. Venter blackish, distal part dark lateral spot and is extended in 2 short rays of femora ferruginous. Antenna, except antenno- toward the scutellum. Length 5.3 mm, width 3 mm. mere I, black. I-lead and pronotum reddish brown, The single specimen has a previous determina- the latter with a clark marginal spot on each side tion "E. ocellatus". and 2 broad black marks extending obliquely mesad from hind angles; behind these marks, near 20. Epopterus cucullinus Gorham center, is a pair of black points. Elytron broadly red Fig. 48 at middle, base and apex suffused with black. A narrow yellow band begins at side margin behind Epopterus cucullinus Gorham 1873:49. Holotype 0', umbo and broadens mesally to form a large ring Brazil: Amazons (NHML). enclosing a large dark spot. At posterior third is a narrow undulate yellow band. Length 5 mm, width Described by Gorham as, "Evenly ovate and 3mm. rather convex .... Elytra ovate, not much narrowed towards the apex, closely punctured .... rich pitchy- 17. Epopterus geniculatus Pic red, with 3 orange-yellow markings, the 2 anterior Epopterus geniculatus Pic 1931:9. Holotype 0', Bolivia irregular and transverse, the third .... an irregular (MNHN). and not clearly defined spot near apex. Antennae pitchy-red, club black." Pronotum with 4 black Similar in size, form and markings to E. picti spots. Length 5 mm. collis, which is possibly a color form of E. genicula Brazil: Ega, Bates, 20'0', 9 (NHML); Amazons, tus. The yellow bands of elytron are narrower, 0' (SMTD). INSECTA MUNDI, Vol. 11, No.2, June, 1997 163

21. Epopterus dilectus Gorham 24. Epopterus confusus Strohecker, n. sp. Fig. 25, 48 Fig. 77 Epopterus cucullinus: Strohecker 1957:267, fig. 21 (not Epopterus dilectus Gorham 1875:16. Holotype 6', Ecua- of Gorham). Holotype 6', (Brazil?): Amazon (HNHM). dor: East Aneles, Buckley (NHML). Form oblong, elytra rather strongly convex. Rather narrowly ovate, similar to E. cuculli Legs and venter brownish black, elytral epipleura nus. Antennal base reddish brown, club black. and sides ofprostern urn yellow. Dorsum dark brown, Dorsum reddish to dark brown, pronotum with 4 elytron with 2 yellow patches, the anterior closely black spots. Elytron with a pale yellow fascia be- embracing the shoulder and bluntly extended me- hind umbo, inclined slightly caudad and not reach- socaudad; the posterior crossing the caudal slope, ing suture; another, more irregular fascia across extending from side margin almost to suture. An- caudal slope. Length 4.5 mm. tenna slender, antenomeres III-VI each abou t 3X as long as wide, antennomeres VII-VIII longer than 22. Epopterus vicinus Pic wide, stalk yellowish brown, club black. Pronotum Fig. 14 with sides parallel basad, rather strongly rounded to front angles, finely margined, disc gently convex. Epopterus vicinus Pic 1931:9. Holotype Q Bolivia (MNHN). Front tibia of male much widened distad, middle and hind tibia straight. Length 4.8 mm, width 3 Oblong. Pronotum in part black. Elytra pitchy, mm. partly brown, with an terior and posterior undulate 25. Epopterus longus Strohecker and branched yellow fasciae. Antenna red, club Fig. 67 black, legs mostly reel. Length 5 mm. The description above is a summary of Pic's Epopterus longus Strohecker 1957:268, fig. 22. Holotype diagnosis. Specimens from Callanga, Peru, closely 6', Amazon (HNHM). resemble the holotype: dorsum blackish brown. Pronotum with 4 black spots in a transverse row; Elongate, subparallel. Elytron with 2 trans- these obscure in darker specimens. Figure 14 was verse yellow bands. In appearance much like E. drawn from a Callanga specimen. Its aedeagus is cucullinus bu t narrower and with posterior elytral identical to that of E. dilectus, and I suspect thatE. band more undulate. Length 4.5 mm, width 2.5 vicinus is a color form of E. dilectus. mm. I have seen only the holotype. 23. Epopterus notatus Strohecker Fig. 16, 71 26. Epopterus obliqUllS Strohecker Fig. 15,72 Epopterus notatns Strohecker 1957:266, fig. 18. Holo- type 6', Amazon (HNI-IM). Epopterus obliquus Strohecker 1957;265, fig 14. Holo- type 6', Amazon sup.: Haituba (HNHM). The holotype and one other specimen subse- quently referred to this species are all I have seen; Subparallel in form but with elytra gradually both specimens are in somewhat immature state of narrowed from anterior third. Dorsum dark brown, coloration. Elytral markings are similar to those of elytron with 3 yellow areas: a quadrate spot at E. vicinus but with yellow areas much wider. In the middle of base, a posthumeral rounded spot, and a holotype the posterior yellow band does not form narrow oblique band across posterior third. Length complete rings. Length 4.:3-4.7 mm, width 2.5-3 4.4 mm, width 2.6 mm. mm. Only the holotype has been studied. Colombia: Amazonas, Leticia, 700ft, 8. VII. 1870, H.& A. Howden (CMNC). 27. Epopterus fallax Gerstaecker Additional material is needed to determine the Fig. 21 relationship of E. vicinus and E. notatus to E. dilectus. Similarity of the aedeagi suggests con· Epopterus fallax Gerstaecker 1858:287. Holotype Q specificity. Amazon (ZMHB 21834). 164 Volume 11, No.2, June, 1997, INSECTA MUNDI

Form somewhat elongate, elytra widest near have been made from a closely similar example in mid-length. Venter black except front of proster- HNHM. I have seen no other material. num and distal sternites, bases offemora and tibiae fuscous, elytral epipleura yellow. Pronotum red- 30. Epopterus vernicatus Gerstaecker dish brown, head with a black fleck, pronotum with Fig. 26, 79 2 lunate discal spots and a marginal spot on each side black. Elytron reddish yellow on base and near Epopterus uernicatus Gerstaecker 1858:285. Holotype Q without locality data (NHML). apex, the extreme apex and a broad median band brown. In the anterior yellow area are 2 dark dark Long-oval. Pronotum with sides convergent spots, the outer on the umbo and rectangular, the from base forward, little rounded, hind angles acu teo inner spot reaching the base and prolonged latero- Elytra broadened from base in anterior third, grad- a slender streak. Two dark spots lie in the caudadas ually narrowed caudad, strongly convex. Venter, caudal pale area, the inner one touching the dark- legs, antennal base and pronotum reddish brown, ened suture. Length 4.8 mm, width 3 mm. pronotal disc infuscate. Elytra blackish brown. I have seen no additional examples. Behind umbo a yellow band extends mesad enclos- 28. Epopterus gracilis Strohecker, n. sp. ing a dark humeral spot and a larger oval dark spot. Fig. 32, 74 From this yellow band a narrow pale streak divides the dark disc (2 median black spots ofGerstaecker's Holotype 0' ancl allotype Q, Peru: Pucallpa, 2.X.1954, E. description). Rather narrow yellow bands enclose 2 I. Schlinger & E. S. Ross (CASC, holotype No.13315). dark spots near apex, which is dark. Length 4.5 mm. Long-oval, elytra rather strongly convex. Ven- A male from Rio de Janeiro resembles the ter and legs reddish brown. Dorsum dark brown, holotype closely but is slightly larger (5 mm) and elytron with triramous yellow basal mark and lacks the discal pale streak. bilunate yellow band across apical slope. Antenna rather slender, antennomeres III-VII each longer 31. Epopterus loretensis Strohecker, n. sp. than wide, antennomeres I-VII brown, antenno- Fig. 81 meres VIII-XI black. Pronotum with sides conver- Holotype 0', allotype Q Peru: Loreto, 20 km from Ucay- ali on Rio Calle ria, Colonia Calleria, 10-30.IX.1961, gent from base forward, little rounded to front B. Malkin (FMNH). Paratypes: 3 Q Q with same angles, hind angles slightly divergent and acute, data as holotype (FMNH; FSCA). discal punctures fine and well spaced, punctation of sides coarser, subrugose. Elytra 3.5X as long as Long-ovate. Venter infuscate, terminal sterni- pronotum, widest in anterior third, gradually nar- tes and legs reddish brown. Dorsum dark reddish rowed cauclad. Length 4 mm, wiclth 2.5 mm. brown, elytron with yellow area at base enclosing 29. Epopterus pictus (Perty) dark humeral spot and ocellate medial spot, and an undulate yellow band across apical slope; band Fig. 22, 42 emits 2 sharp rays in front, and its posterior border Endolnychns pictus Perty 1832: Ill, pI. 22, fig. 11. Type material from Brazil; repository unknown. excised by a fuscous spot; apex is pale. Pronotum Epopterus pictus: Gerstaecker 1858:286. narrowed from base, its sides moderately curved to front angles, marginal bead rather wide and flat, Oval, elytra gradually tapering and strongly hind angles acute. Elytra 3.5X as long as pronotum, convex. Venter, legs, antennal base, headandprono- widened in anterior half, gradually narrowed tum reddish brown, pronotum with 4 black spots in caudad. The abrupt widening offront tibia of male a transverse row. Gerstaecker has described the is less than usual in the genus. Length 4.3-5 mm, elytron as reddish yellow with black-bordered brown width 2.7-3.5 mm. markings: a rectangular shoulder patch and mesad 32. Epopterus anomalus of this a large oval spot; a broad, irregularly mar- Strohecker, n. sp. gined mechan band; behind this 2 large spots trans- Fig. 83 Holotype 0', Ecuador: Pastaza, Ashuara, Rio Macuma versely placed and a triangular spot at apex. Length 10 km from R. Morona, 300 m, 7-16.IX.1971, B. 4.4-4.8 mm. Malkin (FMNH). Allotype Q, Colombia: Putomayo, Unfortunately, I broke the aedeagus of the Rio San Miguel, Santa Rosa de Sucumbios, 400 m, Gerstaecker specimen in extracting it. Drawings 1-7.lX.1971, B. Malkin (FMNH). INSECTA MUNDI, Vol. 11, No.2, June, 1997 165

3 pre-apical spots black; across middle of elytron is Oval. Venter, femora and elytral epipleura black. a broad, black-bordered dark red band, Length 4 Dorsum dark reddish brown, elytron with apex and mm, width 2.G mm. triramous humeral spot yellow. Pronotum relatively small, 2X as wide as long, sides parallel basad, 36. Epopterus atriventris Strohecker, n. sp. gently rounded in front, hind angles weakly acute. Fig. 80 Elytra 3.5X as long as pronotum, rapidly widened Holotype cr,Bolivia: Beni, 31.VII-2.VIII.1960, B. Malkin from base in an terior third, gradually narrowed (FMNH). Allotype Q, Beni, 26.VII-4,VIII.1960, B. Malkin (FMNH). Paratypes: cr, same data as holo- caudad. The anterior yellow patch covers the umbo, type; cr, Brazil: Matto Grosso, Corumba (FSCA). sends a ray across base and a shorter, blunter ray mesocaudad. Length 4.5 mm, width 2.9 mm. Oval, elytra rather strongly convex. Venter and 33. Epopterus aravacus Strohecker, n. sp. legs black, elytral epipleura yellow, dorsum reddish brown, elytron with 2 yellow areas. Pronotum Fig. 8.5 narrowed from base forward but sides well rounded Holotype cr, Venezuela: Aragua, 10 mi. nr. Cata, to front angles, hind angles weakly acute, disc 25.II.1971, I-I. & A. Howden (CMNC). finely and thickly, sides more coarsely and densely punctate. Elytra long-cordiform, about 4X as long Long-oval, similarly rounded in front and be- as pronotum, base with yellow band enclosing dark hind. Venter and femoral bases black. Pronotum umbo andocellate median spot, middle 2/5 of elytron with disc black,shining, very finely punctate, sides dark reddish brown. A yellow band crosses apical yellow with a black spot on each side, which is more slope, its front edge deeply bilunate, its hind edge or less coalescen t with black of disc. Elytron yellow, excised by a large round black spot. Apex pale with umbo and basal spot, a broad median band 2 brown. Length 4,1 mm, width 2.8 mm. pre-apical spots and extre~eapex dark. Medi~n37. Epopterus decempunctatus Gerstaecker band dark brown, its front edge twice deeply incised Fig. 12, 13, 53, 8G by pale rays, hind margin similarly incised. Prono- tum 2.5X as wiele as long, sides evenly rounded, Epopterus decempnnctatus Gerstaecker 1858:290, Lec- anterior curvature not abrupt. Elytra 3.5X as long totype cr, ? Venezuela: Puerto Cabello, Moritz as pronotum and but little wider at base, gradually (ZMHB). Para lectotype Q without data (ZMHB), widened to about middle, thence evenly narrowed Epopterus bifasciatus Pic 1931:9, Holotype cr, Venezu- caudad. ela (MNHN),(New synonym).

34. Epopterus reticulatus Strohecker Oval. Venter and legs rust-red, thoracic sterna Fig. 8, GG and femoral bases darker. Antennal base red, club Epoptems reticula/,ns Strohecker 1957:264, fig. 12. Ho- black. Upper surface reddish yellow, elytron with lotype cr, Bolivia: Cochabamba, Germain (HNHM), brown or black markings: a rounded spot on shoulder and, near mid-line, a narrowly oblong spot, Elliptical in outline, Yellow above with disc of obliquely placed and remote from base; a broad pronotum and 5 spots on elytron dark brown. The band across middle, its front edge twice, its hind pronotal spot is transversely rectangular; the dark edge once excised; 3 spots of variable size before elytral markings consist of a rounded humeral spot, apex. These spots tend to coalesce to form a dark an oblong spot near middle of base, 2 elongate band, leaving the apex and a pre-apical, strongly rectangles near middle, and a rounded or some- undulate band pale. Length 4.3, width 3 mm. what triangular spot before apex. Length 3.8 mm, Colombia: Antioquia, Puerto Benio (GMFT); width 2.5 mm. Norte de Santander, Cucuta, H. & A. Howden 35. Epopterus deltoideus Strohecker (CMNC; FSCA). Venezuela: Bolivar, Suapure, Fig. 11, Gl Caura R., E. A. Klages (FSCA); Lara, Sanare, Epoptems deltoideus Strohecker 1957:266, fig, 20, Holo- 7.XII,G5, J. & B. Bechyne; Aragua, Cata, 14.IX.G5, type cr, Amazon sup,: I-Iaituba (HNHM), Bechyne (lZAV; FSCA), Trinidad: Maracas, H. & A. Howden (CMNC). Oval, elytra narrowly corda teo Dorsum yellow, elytron with umbo, a small oblong basal spot, and 166 Volume 11, No.2, June, 1997, INSECTA MUNDI

38. Epopterus fasciatus fasciatus (Fabricius) 41. Epopterus cordatus Strohecker Fig. 1,46 Fig. 10,58 Tritoma fasciatum Fabricius 1801:570. Lectotype 0', Epopterus cordatns Strohecker 1957:266, fig. 15. Holo- Am(erica) mer(idionalis) (ZMUC). type 0', Amazon (DEIC). Epopterus fasciatlLs: Gerstaecker 1858:288 (not of Fab- ricius). Elytra long-cordate, flattened margin wider than usual. Brown above, elytron with a large Ovate. Venter and dorsum reddish to dark quadrate yellow spot on shoulder, continued as a brown. Elytron with yellow band across base, be· narrow band across base, and a transversely oval ginning at side margin, leaving umbo dark, and yellow spot before apex. Length 3.8-4.5 mm, width enclosing a large ocellate dark spot mesad. A yellow 2.5-2.8 mm. patch before apex encloses a large round dark spot, The type series comprised 15 specimens from which may be partly divided by a pale streak. the "upper Amazon". Brazil: Amazonas, Titirico; Length 3.8-4 mm, width 2.4-2.5 mm. Boca Cauarubi; Maturaca, J.& B. Bechyne. Vene- Brazil: Rio de Janeiro, 20'0', 2 Q Q (FSCA). zuela: 38 km EI Dorado, F. Fernandez & C.J. 39. Epopterus fasciatus angustatus Rosales; Ocano, Fernandez (IZAV; FSCA). Strohecker, new status 42. Epopterus crypticus Strohecker, n. sp. Fig. 2 Fig. 78 Epopterus angnstatns Strohecker 1957:268, fig. 23. Ho- lotype 0', Amazon sup.: Haituba (HNHM). Holotype 0', (Brazil?): Amazon (HNHM).

Long-oval. Elytron dark reddish brown, with 2 Form long-oval, elytra strongly convex, venter yellowish white crossbands edged with black. The and legs brownish black, elytral epipleura and anterior band is wide at side margin and ends sides of prostern um yellow. Dorsum brown, prono- narrowly near su ture and scu tellum. Posterior band tum with 4 black points in tranverse row, elytron transversely rectangular. Length 4.6 mm, width with 2 black-bordered yellow patches, the anterior 2.6mm. enclosing small dark humeral spot and bluntly Larger than the nominate form and somewhat prolonged mesocaudad; posterior patch on caudal narrower. The aedeagus is identical to that of E. slope, broadly undulate. Pronotum with sides grad- fasciatu,s lectotype. ually convergent from base forward, gently round- 40. Epopterus variegatus Erichson ed to rather sharp front angles, hind angle weakly acu teo Elytra more than 3X as long as pronotum, Fig. 6, 7, 43 gradually widened from base to near midlength, Epopterus variegatus Erichson 1847: 181. Holotype 0', then gradually narrower caudad. Antennomere III Peru (ZMHB); Gerstaecker 1858:284. about2X as long as wide, 7 and8 hardly longer than Epopterus decoratns Kirsch 1876: 114 .. Holotype 0', wide. Front tibia of male much widened distad, Peru (SMTD). New synonym. middle and hind tibia lightly but distinctly curved. Epopterns pernvianns Pic, inedit. Length 4.5 mm, width 2.8 mm. Very similar in appearance to E. confu,su,s, the Oval. Light reddish brown below and above, elytral pattern almost identical, but of more oval antennal base yellow. Elytron with dark markings form, with elytra more convex, and differing in as follows: a streak along suture, a humeral spot, an features described and illustrated. oval spot near mid-line not touching base, a broad, sinuate median band, a spot before apex; these 43. Epopterus eganus Gorham markings dark reddish brown with narrow black Fig. 47 borders. Epopterus eganus Gorham 1886: 160, pI. 17, fig. 9. Lec- Description above extracted from original. Spec- totype 0', Brazil: Amazonas, Ega, H. W. Bates (NHML). Paralectotype 0', Ega (NHML). imens from southern Peru are marked as in figure 6, others from Iquitos and "Amazon" as in figure 7. Ovate. Pale chestnut red, elytron with 3 large Length 4-4.5 mm, width 2.4-2.5 mm. yellowish white, black-bordered spots. Of the ante- Peru: Cuzco; Callanga; Marcapata. rior spots one lies against base and suture, the other behind umbo, the posterior spot is round and INSECTA MUNDI, Vol. 11, No.2, June, 1997 167 on apical slope. The 2 anterior spots are narrowly 47. Epopterus rubiginosus Gerstaecker connected in the lectotype but quite separate in a Fig. 18 female from Ega (FSCA ex Janson). Length 5-5.25 Epopterus rubiginosns Gerstaecker 1858:291. Holotype mm, width 3.5 mm. Q, French Guiana: Cayenne (MNHN).

44. Epoptel'us lineoguttatus Gorham Ovate or somewhat oblong. Reddish brown be- Epopterus lineognttatus Gorham 1886: 161, pI. 17, fig. 7. low and above, thoracic sterna infuscate. Antennal Holotype 0', Amazon, H. W. Bates (NHML). base ferruginous, its articles rather short and thick, club black. Pronotum with disc blackish, a vague Identical in structure, including aedeagus, to black spot in each marginal area, front and hind E. eganus but very differently marked. Gorham angles sharp. Length 4.7 mm. described the elytral markings (black) as follows: "a I have seen no other specimens which closely humeral small dot, 2 elongate dashes near the resemble the holotype. Possibly the example here suture, 2 shorter ones in the middle of the disc, 2 described as E. biaculatus is the male of E. rubigi still smaller near the margin above the middle, and nasus, but it is larger than the holotype, with elytra 1 small one near the apex. The dashes are not more rapidly tapering and differently marked. Se- placed regularly, bu t the :3 near the base form a sort ries of topotypes will be needed for clarification. offascia, as do the :3 below the middle. The scutellum is pitchy black. Length G mm." 48. Epopterus batesi Strohecker Probably a color phase of E. eganus. Fig. G8 Epopterns batesi Strohecker 1957:268, fig. 24. Holotype 45. Epopterus ephippiger Gorham 0', Amazon (HNHM). Fig. 54 Epoptems ephippiger Gorham 1886:160, pI. 17, fig. 8. Somewhat broadly ovate. Dorsum blackish Holotype 0', Amazon, Bates (NHML). brown, elytron with 2 rather narrow crossbancls, the anterior at basal quarter and extending from Ovate. Pitchy red below and above. Elytra paler side margin almost to suture, where it is narrowly than pronotum, with a very large round black spot rounded, the posterior band at caudal quarter of common to both. Length G mm, width 4.5 mm. elytron. Length 4.8 mm. I have seen on ly the holotype of this distinctive- ly marked species. 49. Epopterus ryei Gorham Epopterus ryei Gorham 1873:50, pI. fig. 3. Holotype Q, 46. Epopterus quechuanus Strohecker, n. sp. Amazons (NHML). Fig. 27, 75 Holotype 0', Peru: J unin, Tarma, Utcuyacu, 1600-3000 Antennomeres II and V-VIII bead-shaped, base m, III-1948, F. Woytkowski (AMNH). Allotype Q red, club black. Dorsum blackish brown, elytron with same data except 12. III.1948 (AMNH). with 2 sharply defined wide yellow bands. The Paratypes: 4 Q Q, Utcuyacu and Agua Dulce III- anterior band embraces the shoulder, is recurved to 1948, Woytkowski (AMNH, FSCA). the base and there emits a broad blunt ray obliquely Oval. Venter and basal half of femora black, toward the suture; the posterior border of the band distal sternites, tibiae and ends of femora reddish emits 2 short blunt rays. The posterior band, on brown. Dorsum blackish brown, elytron with basal caudal slope is strongly undulate. Length 5 mm. rings and apex yellow. Antenna reddish brown, 50. Epopterus quaesitus Gorham club but little darker, most of basal antennomeres Fig. 20 longer than wide. Pronotal sides weakly conver- Epopterus quaesitns Gorham 1873:51. Holotype Q, Am- gent from base forward but rather strongly round- azons (NHML). ed to front angles, hind angles feebly acute, disc finely and somewhat distantly punctate, punctures Similar toE. ryeibutsmaller and with antennal of sides coarser and denser. Elytra long-cordate, base slenderer. The anterior yellow elytral band each with a bioculate basal yellow band; apex lies behind umbo and widens mesad to form a broad extensively yellow with 1 or 2 black spots. Length arc open toward the suture. The posterior band is 5 mm, width :3.5 mm. strongly undulate. Length 4.5 mm. 168 Volume 11, No.2, June, 1997, INSECTA MUNDI

51. Epopterus similis Strohecker 700 ft, 1O.XII.1970 and 13.II-2.III.1974, H. & A. Fig. 4, 5, 63 Howden (CMNC; FSCA). Epopterus similis Strohecker 1957:265, fig. 13. Holo- type cJ', Amazon sup.: Haituba (HNHM). 54. Epopterus cyclicus Strohecker Fig. 9, 62 Ovate. Light to dark reddish brown, elytron Epopterus cyclicus Strohecker 1957:268, fig. 25. Holo- with 2 rounded or somewhat rectangular, black type Peru: Janjui (HNHM). bordered, pale yellow spots. Of these the anterior lies just behind umbo and is distant from suture. Short-oval. Pronotum ferrugineous. Elytron The posterior spot is similar to the anterior one in dark brown with 2 large yellow marks, the anterior, size and shape and lies on posterior slope. Remarks just behind shoulder, rectangular with its anter- on variation in pattern were made in original de- omesal angle narrowly extended along scutellum scription. Specimens from Ecuador and Peru (Lore- and reflexed laterad along base; the posterior mark to; Huan uco) have the basal elytral mark triramous is a ring, narrowly closed along margin. Length 4.2 and the posterior mark an undulant band but agree mm, width 2.9 mm. in all structural features, including aedeagus, with Peru: San Martin, Janjui, 1600 ft, 5.XII.1946, the holotype. Length 4.5-5 mm, width 3-3.5 mm. J. C. Pallister (AMNH; FSCA). Peru: Loreto, Pucallpa; Huanuco, TingoMaria, 55. Epopterus callerianus Strohecker, n. sp. J.C. Pallister. Ecuador: Pastaza, Ashuara, Rio Fig. 82 Macuma 10 km from R. Moronca, B. Malkin (FMNH; Holotype cJ' and allotype Q, Ecuador: Pastaza, Ashuara, FSCA). Rio Macuma 10 km from R. Morona, 300 m, 7- In antcnnnal structure this taxon resembles E. 16.VII.1971, B. Malkin (FMNH). qzwesitus, and I am inclined to think that it is a color form of that species. Broadly oval but elytra somewhat long-cordiform. Venter and femoral bases infuscate, tibiae, 52. Epopterus transversus Strohecker femoral apices and elytral epipleura ferruginous. Fig. 56, 59 Dorsum reddish brown, elytron with biocellate yel- Epopterus transversus Strohecker 1957:264, fig. 10, A, B. Holotype cJ', Bolivia: Cochabamba (HNHM). low band across base and an undulate pale band across apical slope. Pronotum narrowed from base Ovate, eIytra broadly cordate. Elytron mostly forward, marginal bead wide and flat, hind angles black but with base and apex extensively, suture weakly acute. Length 3.7-4 mm, width 2.5-2.7 mm. narrowly yellow. The basal pale area leaves the 56. Epopterus flavolineatus Strohecker umbo dark and encloses a large black spot which Fig. GO joins the broad black middle area of elytron. Ap- Epopterus flavolineatus Strohecker 1957:266, fig. 16. proximately the posterior quarter of elytron is pale, Holotype cJ', (Brazil?): Amazon (HNHM). enclosing 1 or 2 black spots. Length 4.4-4.7 mm, width 2.6-2.8 mm. Oval. Elytra broadly cordate. Legs and venter blackish brown. Dorsum brown, elytron with 2 53. Epopterus nigrocinctus Strohecker yellow cross bands, the anterior behind umbo, ex- Fig. 57 tending from side margin almost to suture, the Epopterns nigrocinctns Strohecker 1957:264, fig. II. similar posterior band on caudal slope. The mar- Holotype 0', Amazon (HNHM). gins of the bands are almost straight. Length 3.5 Short-oval, elytra broadly cordate. Dorsum dark mm, width 2.5 mm. brown, elytron with 2 broad, black-bordered yellow Only the small type series has been seen. bands, one behind umbo, the other on caudal slope. 57. Epopterus flavonotatus Strohecker, n. sp. These bands touch the side margin but do not reach Fig. 65 the suture. Length 4 mm, width 2.6 mm. Epopterus flavonotatus Pic, inedit. Brazil: Amazonas, Boca Cauaburi, 3UII.I964, Epopterus fasciatus: Gerstaecker 1858: 288 (not of Fab- J. & B. Bechyne; Titirico, 13.IV, Bechynes; Matura- ricius). Holotype cJ', French Guiana: Cayenne (FSCA ca 7.IV.19G4; Uapes, 23.III.19G4, Bechynes (IZAV; ex Janson). Paratypes: Para (ZMHB). Brazil: Belem, FSCA). Venezuela: Amazonas, Cucui, 24. III. 1964, XII-1955, G. & B. Frey, 2cJ'cJ' (GMFT; FSCA); Hai- Bechynes (IZA V). Colombia: Amazonas, Leticia, tuba, 2cJ'cJ' (HNHM; FSCA). INSECTA MUNDI, Vol. 11, No.2, June, 1997 169

Oval, elytra strongly convex. Some examples, 60. Epopterus mimeticus Strohecker including the Pic specimen apparently have not Fig. G4 attained definitive coloration. In these, the dorsum Epopterus mimeticns Strohecker 1959: 181, fig. 5. Holo- is generally pale brown, elytron with a yellow band type 0', Amazon (ZMHB). across base and curving caudad along scutellum and suture. Behind this band is a diffuse black area Short-oval, similar in form to E. clara. Elytron enclosing 2 yellow spots transversely placed. Apex yellow with with 2 dark basal flecks and a large yellow and in front of this pale area are 2 rather black patch at mid-length, which encloses a yellow large rounded yellow spots. spot. Length 4 mm, width 2.5 mm. The pattern which I consider the mature one is 61. Epopterus dives similar to figure 19. Legs and venter, including Gorham Fig. 50 elytral epipleura, dark brown. The distal expansion Epopterus dives Gorham 1874:443. Holotype 6', Ama- of front tibia of male is rather gradual and moder- zons (NHML). ate. Length 4.3 mm, width 2.8 mm. 2 58. Epopterus submaculatus Strohecker, n. sp. Short-oval. Dorsum brown, elytron with yel- Fig.87 low marks, the anterior triramous, the posterior Epopterns sltlimacltlatlls Pic ineclit. Holotype 0', Boliv- undulate, narrowed mesad and almost reaching ia, Pic coli. (MNI-IN). suture. The anterior yellow bar extends forward along side margin and internal to umbo, emits a Oblong in outline, strongly convex. Legs, an- spur to elytral base, thus enclosing a dark humeral tennal base and prosternum yellow; other parts of spot; from origin of this spur a pale bar deflects venter dusky, becoming paler caudad. Pronotum obliquely caudad, its end a little enlarged and far with disc and hind angles black, front angles and from suture. Length 3.5-3.8 mm, width 2.G mm. sides broadly yellow. Elytron black with a broad U. Amazon: Yurimaguas (FSCA ex Janson). transverse yellow band behind umbo, reaching 62. Epopterus parvus Strohecker, n. sp. suture and there expanded slightly caudad and Fig. 19,73 forward to scutellum. Distal third of elytron has 2 Holotype 0' ancl allotype 9, Brazil: Para (FSCA ex oval yellow spots transversely placed and behind Janson). Paratypes: 599, Para (FSCA). these a large round yellow spot which leaves apex, suture and side margin narrowly black. Front tibia Short-oval, elytra cordiform, tapering from of male curved at base, distally enlarged with anterior third. Venter and legs brown. Antennal proximal end of enlargement mesally angulate. base brownish yellow, club black. Dorsum pale to Length 3.8 mm, width 2.4 mm. dark brown, elytron with 2 yellow marks: a band at 59. Epopterus clara Gorham base which leaves a dark humeral spot and encloses Fig. 51 an ocellate dark spot near base and scutellum; Epopterus clara Gorham 1874:443. Holotype 9, Ama- apical third yellow, enclosing 1 or 2 black spots, the zons (NHML). single spot due to fusion of the two. Length 3.5-4 mm, width 2.3-2.8 mm. Short-oval, convex. Elytra widest in basal third, 63. Epopterus histrio Gerstaecker gently tapering, apex rather sharply rounded. Fig. 17, 52 Pronotum and scutellum reddish to dark brown. Epopterus his trio Gerstaecker 1858:292. Lectotype 9 Elytron brownish yellow with G or 7 black mark- French Guiana: Cayenne (MNHN). ings: a humeral spot which does not reach base and mesad of this and oblique oval spot, 3 patches Short-ovate, elytra cordate, widest in basal transversely placed near middle, the lateral spot third. Venter and legs brownish black. Antennal elongate and close to margin, the middle spot larg- base dark red, club black. Head and pronotum dark est and long-oval, the inner spot subquadrate and brown. Elytron black at sides; center of disc and slightly behind the others, 2 spots on apical slope apex reddish brown. A yellow band encloses a dark which may be fused into an arcuate band. Length humeral spot, turns forward to run along base and 3.8-4 mm, width 2.G-2.8 mm. is then deflected along scutellum; the hind border Brazil: Para; Rio de Janeiro; Sao Paulo (FSCA). of this band emits a sharp ray. The band may form 170 Volume 11, No.2, June, 1997, INSECTA MUNDI a complete ring mesad to enclose an oval dark spot. ferred to Anidrytus by Csiki or me: A. juveneus, A. Another yellow band of irregular shape crosses the einetipennis, A. fuliginosus, and A. spadieeus. apical third of elytron, its hind edge excised by a black spot. This band may also completely encircle 65. Anidrytlls bipunctatus Gerstaecker the dark spot. Length 3.6-4 mm, width 2.4-2.8 mm. Fig. 92 French Guiana: Cayenne (FSCA). Brazil: Ama- Anidrytus bipunctalus Gerstaecker 1858:257. Lectotype 0', Brazil (MNHN); paralectotype, Brazil (ZMHB). pa, Rio Carnot; Serra Lombard, J. & B. Bechyne (lZAV; FSCA); Para (FSCA). Long-ovate, bright to dark rust-red with yellow 64. Epopterlls pantherinlls Gorham pubescence; spinasternum, meso- and metaster- Fig. 84 num, basal half of tibiae and base of middle and Epopterus pantherinns Gorham 1889: 131, pI. 7, fig. 22. hind femora blackish brown. Antennomeres I-II Lectotype Q, Type/ sp. figured/ David, Chiriqui, red, the others black. Length 9-9.5 mm. Champion/ B. C. A., Col. VIII Epopterus pantheri- Pronotum more than 2X as wide as long, nar- nus Gorh. (NHML). rowed from base forward, front angles short and rounded, hind angles weakly acute, disc with 2 Oblong-ovate, strongly convex. Antenna black, dark flecks, irregularly punctate. Elytra elongate, antennomeres I-II reddish. Tibial base and scutel- wider at base than pronotum, gradually widened in lum black. Pronotum reddish brown, front margin, anterior third then gradually convergent distad. I basal half of side margin and base narrowly black. have seen only the types. Elytron with pale markings as follows: a broadly triramous band covering most of basal third, 2 66. Anidrytus plagiatus Gerstaecker transversely placed median spots which may be Fig. 93 united, 2 spots on apical slope and 1 at apex. These Anidrytus plagiatns Gerstaecker 1858:259. Holot.ype 0', pale areas cover most of elytron, leaving darker French Guiana: Cayenne (MNHN). areas quit.e narrow. Length 5.25-6 mm. I have recen tly examined a pair from Barro Form oval, elytra rather strongly convex. Ven- Colorado Island sent by Henk Wolda. ter and legs rust-red. Antennomeres I-V red, the others black. Pronotum 2.5X as wide as long, disc Genus Anidrytus Gerstaecker with rectangular black patch, sides subparallel Gerstaecker 1858:256; Chapuis 1876: 122; Gorham basad, abrup tly rounded in fron t, front angles short, 1889: 125; Strohecker 1953:64. hind angles produced and acute. Length 6.5 mm. Type species: Anidrytlls bipunctatlls Gerstaecker (Stro- Front tibia of male with inner edge widely hecker 1953:64). excavate beyond middle, then abruptly widened. Specimens of most species are 5.5-9 mm long. In 67. Anidrytus nimbiferlls Strohecker, n. sp. several species, however (Nos. 74-84, 95), length Fig. 94 may be less t.han 5 mm. Coloration more somber Holotype 0', Colombia: Amazonas, Leticia, 700 ft, than in Epopterus, often entirely brown or black 11.VII.1970, H. & A. Howden (CMNC); paratype 0', with paler color when present. confined to sides and Brazil; Amazonas, Bat.es (NHML). apex of elytron. Rarely a pale sutural stripe is present. Anidrytus uariegatus (Guerin-Meneville) Long-oval, elytra rather strongly convex. Ven- of southern Brazil is, as its name indicates, strik- ter, legs, head and antennomeres I-II rust-red, ingly bicolored, while A. deeoratus Arrow and A. antennomeres III -XI shining black. Pronotum with humerosus n.sp. may have elytral base extensively rectangular black patch at middle, sides broadly, yellow or reddish. A puzzling maculate specimen is front margin narrowly red. Elytra black with base, noted under A. poreus Strohecker. Body outline is including scutellum, side margin and apex red. broadly ovate to oblong or even long parallel. Sides of pronotum convergent from base forward, No illustrations are presented for A. nitidular front angles prominent and weakly acute, hind ius Gerstaecker, A. hilaris Gerstaecker, A. ehampi angles produced and decidedly acute, lateral sulci oni (Gorham), A. quadripunetatus Gorham, A. linear with fovea at base. Elytra more than 3X as marginatus (Fabricius), and4 species described by long as pronotum (mid·line) and as wide at base, Guerin-M6neville under Epipoeus but were trans- slightly widened to midlength then gradually con· INSECTA MUNDI, Vol. 11, No.2, June, 1997 171 vergent caudad. Front tibia of male as in A. plagia 71. Anidrytus ouatulus Gerstaecker tus. Length 8.3 mm, width 4.9 mm. Fig. 98 Anidrytus ovatulus Gerstaecker 1858:262. Lectotype cr, 68. Anidrytus uariegatus (Guerin-Meneville) Colombia (ZMHB). Fig. 134 Oval, strongly convex. Rather dark brown, an- E(pipoClts) variegatns Guerin-Meneville 1857:266, pI. tennomeres III-XI black, elytral epipleura and tib- 13, fig. 19. Holotype Q ?, Brazil; repository un- iae yellowish brown. Similar to A. contractus but of known. somewhat longer form. Length 6 mm, width 3.9 Anidrytns singularis Gerstaecker 1858:260. Holotype mm. cr, Brazil (MNHN); Csiki 1901:35 (syn.). Venezuela: Maracay, P. Vogl (ZMHB; FSCA). Short-oval, elytra cordiform, convex. Prono- 72. Anidrytus parhi Strohecker, n. sp. tum narrowed from base forward, reddish yellow Fig. 133 with 3 discal spots and basal half of marginal bead Holotype cr and allotype Q, Panama: Barro Colorado black. Elytra blackish brown with 2 large pale Island, 28.VII.1973, O. Park (FMNH). markings: the anterior a broad undulant band behind umbo, the posterior a large triangular, Oval. Largely rust-red, the distal antennomer- patch near apex. These marks may be connected by es darker but not black. Antenna stout, antenno- a pale band close to su ture. Length 6 mm. mere III less than 2x as long as wide, antennomeres The type of A. singlliaris is callow, so soft that IV-V subequal to III, antennomeres VI-VIII each the aedeagus cou ld not be extracted. Rio de Janeiro, aboutas wide as long. Pronotum with sides parallel cr, 2 Q Q (NHML). basad, well rounded to front angles, hind angles rectangular, disc very finely punctate, punctures 69. Anidrytus nitidularius Gerstaecker at sides closer and denser. Lateral sulci not quite Anidrytus nitidularius Gerstaecker 1858:262. Holotype reaching middle. Elytra not quite as long as prono- (sex ?), Mexico, presumably in Dohrn colI. (ZMPA). tum, rather strongly convex, widest before middle and gradually narrowed caudad, reddish black in Short, oval. Blackish brown, antennomeres I- holotype with umbo, side margins and apex red; V, front angles of pronotum, elytral apex, venter allotype black above, sides of pronotum, elytral and legs red. Length 6 mm. umbo, side margins narrow ly and apex red. Length Antennomeres III-IV equal, antennomeres VII- 6-6.5 mm, width 3.7 mm. VIII globular. Pronotum rather narrow, hardly 2X Front tibia of male has angulate flange on inner as wide as long, sides subparallel basad. Elytra edge. widest in basal third, thence gradually convergent. Last abdominal sternite deeply semicircularly ex- 73. Anidrytus porcus Strohecker cised. I have not recognized this species in the Fig. 129 material studied. Anidrytus porcus Strohecker 1957:269, fig. 28. Holotype cr, Amazon (HNHM). 70. Anidrytus contractus Gerstaecker Fig. 97 Short-oval strongly convex, sides of elytra much Anidrytus con/,rac/'ns Gerstaecker 1858:263. Holotype rounded. Elytral disc dark brown, base and sides Q, Costa Rica (ZMHB 21818). Anidrytus dolosns Gorham 1889: 127, pI. 7, fig. 18. Lec- reddish brown. Pronotum and antennomeres I-VII totype and 5 paralectotypes, Panama: Volcan de reddish brown. In male front tibia is distally wid- Chiriqui, Champion (NHML). New synonym. ened and bidentate, hind tibia with a short sharp tooth on inner edge near apex. Length 5.7 mm, Short-oval, strongly convex. Brown to blackish width 3.8 mm. brown, antennomeres I-V or I-VI reddish. Notable Colombia: Amazonas, Leticia, 700 ft, 19- features of the species are the short broad form and 25.11.1972, H. & A. Howden. One male of this series the very deep lateral sulci of pronotum. Length 5.7- is smaller (5 mm) and has dorsum reddish brown, 6 mm, width 4-4.2 mm. elytron with 2 large, black-bordered, transversely Costa Rica: Turrialba, A. Heyne (ZMHB; FSCA), oval yellow bands, but is structurely identical, Panama: Barro Colorado Island (CASC; FMNH). including aedeagus, with typical specimens. 172 Volume 11, No.2, June, 1997, INSECTA MUNDI

74. Anidrytus unicolor Kirsch Short-oblong. Ferruginous to dark brown be- Fig. 128 low and above, antennomeres I-V or I-VI rufous, Anidrytus unicolor Kirsch 1876: Ill. Holotype Q, Peru the others black. In some examples of lighter color (SMTD). (less mature) the entire antennal base may be pale. Kirsch has described the type as follows: short- Front tibia ofmale gradually widened to apex, hind ovate, moderately convex, pubescence yellowish tibia enlarged and somewhat flattened in distal gray, dorsum and antenna light brown, front of half. Length 4.5-5 mm, width 2.9-3.3 mm. head, side margin of elytra and venter dull yellow, I have examined many specimens from Venezu- metastern urn and femora brownish. Pronotum with ela' Surinam, Guyana. sides parallel from base to beyond middle then strongly rounded to front angles. Elytra hardly 78. Anidrytus concolor Kirsch wider at base than pronotum, widened in basal Fig. 120 third, thence gradually narrowing. Length 4 mm, Anidrytlls concolor Kirsch 1876: 112. Holotype Q, Peru width 2.5 mm. (SMTD). Two pairs from Peru (Dept. Huanuco) conform well to Kirsch's diagnosis except for color; the type Exceedingly similar to A. helvolus, so similar is probably immature. In these 4 examples the that I have considered the placement of the name dorsum is dark brown with marginal area of elytra concolor in synonymy. The slight difference in yellowish brown and an tennal club is black. Front aedeagi, however, seems to be consistent. tibia of male curved, gradually enlarged to apex, I have examined many examples from Peru, hind tibia clavately widened with low broad tooth at Ecuador, Colombia (Putomayo) and Brazil (lga- distal third of inner edge. Length 4.2-4.7 mm, width rape). 2.7-3 mm (AMNH; FSCA). 79. Anidrytus gerstaecheri Kirsch 75. Anidrytus glaber Kirsch Fig. 130 Fig. 125 Anidrytus gerstaecheri Kirsch 1876: 112. Holotype Q, Peru (SMTD). Andrytusglaber Kirsch 1876: 113. Lectotype Q with type label, Peru (SMTD). Two pairs (SMTD) are probably syntypes (paralectotypes). Short-ovate. Reddish to dark browm. Antenna rather short, antennomeres I-V red, antennomeres Short-oblong, dark reddish brown, antenno- VI-XI black. Front tibia of male curved, gradually meres black VI-XI. Front tibia of male widened to enlarged to tip. Length 3.7 mm, width 2.5 mm. apex. Length 3.8-4 mm, width 2.8 mm. Structurally similar, including aedeagus, to Peru: Marcapata (FSCA). Epopterus dives, but lacks the elytral markings of that species. 76. Anidrytus consobrinus Strohecker Fig. 12G 80. Anidrytus tenuipes Strohecker Anidrytus consobrinns Strohecker 1957:270, fig. 32. Fig. 127 Holotype el, Amazon (HNHM). Anidrytlls tenllipes Strohecker 1957:271, fig. 33. Holo- type el, Bolivia: Cochabamba (HNHM). Short-oblong. Dark reddish brown below and above, antennomeres VII-XI black. Front tibia of Oblong, rather narrow. Dark ferruginous with male gradually widened distad, with a small sharp sparse brassy pubescence, antennomeres VII-XI tooth at distal third. Length 4.5 mm, width 3 mm. black. Front tibia of male enlarged distad, some- Brazil: Mission Cauaburi am Kanal Maturaca, what knob-shaped at apex, hind tibia long (1.2 mm) 24.II-4.III.19G4; Uaupes, Rio Negro, 18.I-3.II.19G4, and slender. Length 4.8 mm, width 2.7 mm. C. Lindemann (ZSMC). Amazon sup.: Haituba (HNHM; FSCA). 77. Anidrytus heluolus Gerstaecker 81. Anidrytus [usc us Arrow Fig. 119 Fig. 123 Anidrytlls helvolzts Gerst.aecker 1858:272. Lectotype el, Anidrytlls fllSCllS Arrow 1920:47. Lectotype el and 31 Venezuela: Orinoco (ZMHB). paralectotypes, Brazil: Pernambuco (NHML). Epheblls ignobilis Gorham 1875: 17. New synonym. Holotype Q, French Guiana: Cayenne (NHML). "A considerable series of this .... vary in colour from light brown to nearly black, but in fully INSECTA MUNDI, Vol. 11, No.2, June, 1997 173

coloured specimens the five basal joints and the margins narrowly red. Pubescence long and gray. extreme tip of the antennae, the sides of the protho- Length 5.5 mm. rax, the shoulders and extremities of the elytra, and the legs are reddish and the remaining parts 84. Anidrytlls gibboSliS Strohecker, n. sp. very dark. It is a rather small, short and broad Fig. 121 insect, not very convex and rather strongly and Holotype 0', Colombia: Valle del Cauca, 70 km E closely punctured. Long. 4.5-5 mm." (Arrow). Buenaventura, 1200 ft, 17.r1.1970, H. Howden (CMNC). Front tibia of male slightly dilated toward tip, with an acute tooth at distal third of inner edge. Ven ter, legs, head and antennomeres I -III brigh t Some specimens of the type series have the red. Dorsum deep black, marginal bead of prono- dorsum entirely black. A series taken by Borys tum reddish. Pubescence long and gray. Pronotum Malkin (FMNH) in NE Matto Grosso shows the narrowed from base, sides little curved. Elytra same range of coloration. I n males of this series the highly convex, broad cordiform. Front tibia of male apophysis of the aedeagus forms a sharper angle bowed, widened in distal half. Length 5.8 mm, with the shaft than in type examples, but I am width 4.2 mm. reluctant to regard this as a specific differentiation, This may represent the male sex and mature especially since the Malkin series exhibits some coloration ofA. championi but until topotypic males variation in this feature. of that species become available I think it better to 82. Anidrytlls compactlls Strohecker, n. sp. record it as distinct. Fig. 124 85. Anidrytlls piloSliS Arrow Holotype 0', Peru: Monson Valley, Tingo Maria, Fig. 11G 21.X.1924, E. I. Schlinger & E. S. Ross (CASC, Anidrytus pilosus 0' holot.ype no. 1:3:314). Allot.ype Y, same dat.a as holo- Arrow 1920:47. Lectot.ype and a type except 9.XrI.54 (CASC). Paratypes: 20'0", Tin- paralectotype 9, S Brazil: Espirito Santo, Schmidt go Maria, 9.XrI.54 (CASC; FSCA);O', Tingo Maria, (NHML). 2000 ft. (600 m), 20.X. 1956, J. C. Pallister (AMNH); Colombia: Putomayo, Santa Rosa, headwaters Rio Form (male) broadly ovate, strongly convex. San Miguel, 10-23.X.1970, B. Malkin & P. Buchan- Dorsum brownish yellow with long pale yellow an, 20'0', 299 (FMNH; FSCA). pubescence. Pronotum with sides gently curved in front, divergent basad, hind angles acute. Elytra Short-oblong, compact, strongly convex. Ven- broadly cordiform. Length 5.5 mm. ter black, legs dark reddish brown, femoral bases Arrow described the female as narrower than fuscous. Antennomeres I-II dark red, 3-11 black. male with the pronotal sides strongly rounded in Dorsum entirely black, finely and closely punctate, front and parallel toward base. pubescence fine and gray. Pronotum with sides strongly rounded to front angles, parallel basad, 86. Anidrytlls paralleills Gerstaecker hind angles rectangular, lateral sulci deep, base Fig. 117 strongly sinuate. Elytra widened from base to mid- Anidrytl1s parallell1s Gerstaecker 1858:266. Holotype 0' dle, gradually narrowed caudad, apices abruptly (not 9 !), Brazil (ZMHB 21820). Ephebus depressus Gorham 1875: 17. Holotype 9, Bra- rounded. Length G mm, width 4 mm. zil: Rio de Janeiro (NHML). New synonym. 83. Anidrytlls championi (Gorham) Anidrytl1s depressl1s; Arrow 1920:46. Systaechea championi Gorham 1889: 133, pI. 8, fig. 3. Holotype 9, Panama: Volcan de Chiriqui, Champi- Elongate-oblong. Dark reddish brown, thoracic on (NHML). sterna and first sternite fuscous, legs rust-red. Anidrytus championi Arrow 1920:53. Antennomeres VIII-X darker than others. Front tibia of male somewhat widened toward apex. Short-oval, subhemispheric. Venter and legs A very good photograph of the holotype of E. red. Antennomeres I-V red, VI-Xl black. Pronotum depressus has been compared with the single type narrowed from base to fron t with little curvature of of A. parallelus and shows the two to be much alike. sides, front and hind angles acute, base bisinuate, The original descriptions are also very similar. The disc bluish black at base, sides and front margin type of A. parallelus is 7 mm long, thatof depressus red. Elytra highly convex, bluish black with side G.3mm. 174 Volume 11, No.2, June, 1997, INSECTA MUNDI

87. Anidrytlls stenllS Strohecker A male in the Janson material (FSCA) agrees Fig. 118 well with the lectotype and has Gorham's label Anidrytus stenns Strohecker 1979:716. Holotype 0', Bra- "lugubris". Another male is structurally identical to zil: Sta. Catarina, Nova Teutonia (MHNG). the first bu t has elytral umbo and apex reddish Elongate-oblong. Dark brown with gray pubes- yellow. Front tibia with distal third dilated, flat- cence, pronotal disc vaguely darker. Antenna slen- tened and pubescent beneath. der, antennomeres I-V red, others deep black but tip of antennomere XI pale. Pronotum with sides 91. Anidrytlls ephippillm Gerstaecker strongly rounded from middle to front angles, a Fig. 103 little convergent basad, hind angles feebly acute or rectangular. Elytra with sides almost exactly par- Anidrytus ephippinm Gerstaecker 1858:264. Holotype allel from behind umbo to apical third. Front tibia Q, Surinam: Paramaribo, presumably in Dohrn of male evenly and weakly widened distad. Length colI. (ZMPA). Anidrytus fallaciosns 7 mm, width 3.4 mm. Gorham 1873:47. Holotype Q, French Guiana: Cayenne (NHML); Arrow 1920:47 The elongate form and bizarre aedeagi of this (syn.). species and A. parallelus indicate divergence of generic rank. Oblong, subparallel. Prosternum, elytral epi- 88. Anidrytlls atratllills Gerstaecker pleura and legs brownish yellow, meso- and metast- Fig. 122 ernum fuscous. Antennomeres I-V, head, sides of Anidrytus atratnlns Gerstaecker 1858:273. Holotype Q, pronotum and scutellum rust-red. Elytra black, S Brazil (ZMHB 21824). shoulder, side margin narrow ly at apex extensively red. Front tibia of male clavately widened to tip, Oblong, subparallel. Deep black above with with a slight angulation of inner edge near apex. brown pubescence, elytral shoulder and apex dark Length 6-6.5 mm. red, surface coarsely and densely punctate. Length A pair from Cayenne, det. Gorham (NHML);, 5mm. Cayenne, det. Gorham (FSCA ex Janson). A specimen with label "Braz." (FSCA ex Jan- 92. Anidrytlls marginatlls son) agrees well with the holotype. This male spec- (Fabricius) Endomychns marginatus Fabricius 1801:504. Holotype imen also carries the notation by Gorham "n. sp.? Q, Am(erica) mer(idionalis), Schmid (ZMUC). n.? fallaciosus Gorh." 89. Anidrytlls hilaris Gerstaecker Oblong, subparallel. Under surface, legs and Anidrytns hilaris Gerstaecker 1858:271. Holotype Q, S elytral epipleura rufous, metasternum darker. Head Brazil Zello (ZMHB 21822). and antennomeres I-VII dark red, antennomeres VIII-XI black, antennomere XI apex reddish, an- Oblong, subparallel. Antennomeres IV-XI shin- tennomeres IX-X strongly transverse and mesally ing black, the insect otherwise shining rust-red, acuminate. Pronotum with sides parallel basad, dorsum with yellow pubescence. Length 5 mm. strongly rounded in front, hind angles feebly acute, I have seen no additional examples. disc weakly convex, finely and densely punctate, black, sides and front angles rufous. Elytra about 90. Anidrytlls lllgllbris Gerstaecker 3X as long as pronotum, moderately convex, finely Fig. 104 and thickly punctate, black with umbo, side margin Anidrytns lngubris Gerstaecker 1858:270. Lectotype Q, narrowly and apex extensively rufous. Length 6.3 Colombia (ZMHB 21821). Paralectotypes: Q, Co- mm, width 3.6 mm. lombia (ZMHB); Q , Colombia (NHML). I have seen no other specimens which agree entirely with the holotype, which resembles A. Broadly oblong, subparallel but elytra rather ephippium closely but differs markedly in struc- abruptly rounded to apex. Deep black with anten- ture of antennal club. nomeres I-V, elytral epipleura and lower face of tibiae brownish yellow; tips of middle and hind 93. Anidrytlls cinctipennis (Guerin- femora pale yellow. Length 5.2-6 mm, width 3.4-3.6 Meneville) mm. E(pipocus) cinctipennis Guerin-Meneville 1857:267. Holotype, sex ?, French Guiana: Cayenne, colI. De- jean; repository unknown. INSECTA MUNDI, Vol. 11, No.2, June, 1997 175

Described by Guerin as black, oval, rather black, strongly and rather densely punctate, sides shining. Head, sides of pro no tum and elytra above red, rather rugosely punctate, red coloration ex- and below, basal half of antenna, and legs tawny panded atfront angles. Elytra more than 3X as long yellow. Pronotum and elytra punctate, tomentose. as pronotum, gradually and evenly widened to mid- Length 6 mm, width 3.5 mm. length, and similarly narrowed caudad, apex abrupt- The name E. cinctipennis was treated as a ly rounded; base, sides and apex red, disc black, synonym of A. marginatus by Gerstaecker. Per- densely punctured and with short tawny pubes- haps it should be used for the species described by cence. Venter red, sternite 1 and legs reddish black. Gerstaecker as A. ephippizun bu t lack of a type and Front tibia of male thickened distad but without the brevity of Guerin's diagnosis make me unwill- tooth; a small sixth sternite is visible. Length 4.5 ing to indicate such synonymy. mm, width 2.5 mm. Unusual among the smaller forms of the genus 94. AnidrytlJ.s circlJ.mcinctus Strohecker, n. sp. in its contrasting coloration. Fig. 106 Holotype 0', Guatemala: Sac., Reunion, 4000 ft, 96. AnidrytlJ.s batesi Strohecker, n. sp. 22.VIII.1947, C. & P. Vaurie (AMNH). Fig. 105 Holotype 0', Brazil: Ega (NHML). Paratypes:O', Ama- Oblong, subparallel. Venter and legs rust-red. zon, Bates (NHML);O', Amazon (FSCA ex Janson). Head, antennomeres I-VI and sides of pronotum red, pronotal elisc black. Elytron black with base, Broadly oblong, moderately convex. Venter red- umbo, side margin, apex and suture reel. Antenna dish black, legs and elytral epipleura rufous. Dor- stout, antennomeres II-IV somewhat elongate, sub- sum black, sides ofpronotum vaguely reddish, side equal, antennomeres V-VIII hardly longer than margins and apex of elytra red, surface finely and wide. Pronotum with sides parallel in basal half, densely punctate, pubescence coppery. Antenna strongly rounded in front, hind angles rectangular, rather slender, red at base, black beyond, antenno- lateral sulci foveately wider at base. Elytra nearly meres III-IV subequal, antennomeres V-VIII secu- 4X as long as pronotum, slightly widened from base larly shorter, antennomere IX sharply acuminate in anterior third, then parallel for much of their internally but its medial side is equal to its distal length, apical rounding rather abrupt. Front tibia width. Pronotum strongly transverse, sides paral- of male clavately widened distad, without distinct lel basad, much rounded in front. Elytra with sides tooth. Length 5.5 mm, width 3.3 mm. subparallel but continuously curved, apical rounding rather blunt. Front tibia ofmale curved at base, 95. AnidrytlJ.s bechyneorlJ.m Strohecker, n. sp. much widened near tip with low angulate tooth Fig. 107 below; hind tibia broadly curved. Length 6.5-7 mm, Holotype 0', Venezuela: Aragua, EI Limon, 450 m, width 4.5 mm. 6.V.1962, F. Fernandez. Allotype 0, same data as holotype (IZAV). ParaLypes: 40'0' with same data 97. Anidrytus amazonicus Gorham as holotype; EI Limon, 450 tn, 25.V.1977, J. Clavijo, Fig. 101 25 27. V. 1965, 0; and 20 0, Fernandez; Anidrytus amazonicus Gorham 1873:46, pI. fig. 8. Holo- 22. VII.1971, 0, Fernandez; Barinas, Reserva For- type 0, Brazil: Amazonas, Bates (NHML). estal Caparo-Camp Cachicamos, 100 m,O'O, J. Sal- cedo, F. Zambrano; Portuguesa Est. Exp. San Nico- las, 56 km de Guanare, 180 m, 1-2.IV.1968,O', C. J. Broadly oblong, subparallel but elytra with Rosales. sides continuously curved, rather bluntly rounded to apex. Venter and legs red, hind coxae darker. Small for the gen us, ou tline su bparallel. An ten- Antennal base red, club black. Pronotum red, sides na black, tip oflastjoint rufescent, antennomere III parallel basad, much rounded in front. Scutellum slightly more than 2X as long as wide, antennomer- red. Elytra black with umbo and apex red; side es IV-Veach longer than wide, antennomeres VI- margin may be narrowly red. Front tibia of male VIII quadrate. Pronotum not quite 2X as wide as angulately widened near apex, with small tooth long (mid-line), front angles rounded, sides feebly below. Length (male) 7.8 mm, width 4.8 mm. rounded and weakly convergent to base, hind an- Brazil: Fonte Boa, Bates, 0' (NHML). gles rectangular, lateral sulci long and deep, disc 176 Volume 11, No.2, June, 1997, INSECTA MUNDI

98. Anidrytus cupreatlls Strohecker except pale apex of antennomere XI. Front tibia of Fig. 102 male angulately expanded near apex, with small Anidrytus cupreatns Strohecker 1957:269, fig. 29. Holo- tooth below. Length 6.5-7.4 mm, width 4-4.3 mm. type cf, Amazon sup. (HNHM). Peru: Huanuco, Terma, Utcuyacu, 18.III.1948, Very similar in structure to A. amazonicus but F. Woytkowski,cf, 300 (AMNH; FSCA); Diviso- of somewhat narrower, more parallel form. Anten- ria, 27.IX.1946, Woytkowski, (AMNH; FSCA). nomeres I-III rufous, others black. Body and legs ferruginous with coppery pubescence above. Front 101. Anidrytus lique/actlls Gorham tibia of male angulately dilated near apex with Fig. 95 small tooth below, middle tibia with row of stout Anidrytus liquefactus Gorham 1873:47. Holotype cf, setae on inner edge, hind tibia curved. Length 7-7.4 Colombia (NHML). mm, width 4.2-4.4 mm. Peru: Estiron, Rio Ampiacu, 15-22.V.19G6, B. Oblong, elytra obovate, dark ferruginous, shin- Malkin, cf (FMNH). ing, antennomeres I-IV rufous, others black. Prono- tum 2X as wide as long, sides parallel basad, strong- 99. Anidrytus nigricans (Guerin-Mimeville) ly rounded in front. Elytra 1.5X as wide as prono- Fig. 99 tum, evenly rounded from shoulder to apex, strong- E(pipocus) nigricans Guerin-Meneville 1857:265. Holo- ly but not very thickly punctate and with depressed type (sex ?), Brazil, Dejean colI.; repository un- gray pubescence. All tibiae of male bent, front ones known. angulately widened near apex. Length 7-7.6 mm. Anidrytus nigricans Gerstaecker 1858:267. Holotype Q, Panama: Bugaba, Champion, cf (NHML). Co- Brazil (MNHN). lombia: Muzo, cf (ZMHB). Although Guerin's description is too brief for 102. Anidrytus mexicanus Strohecker, n. sp. usefulness, Csiki (1901::3.5) attributed the species Fig. 96 name to him. According to Gerstaecker's diagnosis Holotype cf Mexico: Chiapas, San Quintin, 700 ft, rain the species is oblong but with elytra somewhat forest, 5-20.11. 1966, G. F. Baker & D. R. Whitehead attenuate caudad. Venter brown, legs and elytral (FSCA). epipleura rust-red. An tennomeres I-IV red, antennomeres V-XI black. Dorsum black, elytral tips Entirely similar to A. liquefactus except in reddish. Length 6.7 mm. some features of the aedeagus andin having anten- As I have identified it on the basis of aedeagus, nomeres I-V rufous. It may only be a color form of this species varies in size, color and to some extent A. liquefactus. Length 7.7 mm, width 4.5 mm. in elytral outline. Some specimens, evidently callow, have sides of pro not urn , base, side margin and 103. Anidrytusjuuencus (Guerin-Meneville) E(pipocus) juvencus apex of elytra ferruginous. Other specimens are Guerin-Meneville 1857:266. Evi- dently described from 2 or more examples from entirely deep black above, while others from Para- Colombia in colI. Dejean. Location unknown. guay are black with elytral umbo and apex dark Anidrytus juvencus: Csiki 1901:35. red. Length 5.5-6.8 mm, width :3.5-4 mm. Brazil: Rio de Janeiro, various collections; Sao Tawny yellow, venter and legs paler. Antenna Paulo; Corumba (FSCA). Paraguay: San Pedro, black with antennomeres I-III yellow. Middle of Corumbe, R. Golbach (IMLA; FSCA); way between pronotum and scutellum dusky. Dorsum finely Asuncion and Puerto P. Stroessner, Mahunka punctate, with yellowish pubescence. Length 6-8 (HNHM; FSCA). Argentina: Jujuy, Tierras de Lam- mm, width 4-4.5 mm. See No. 104. brisca, F. Monr6s (IMLA); Salta, Tartaga, Golbach (IMLA); Cotomayo, Golbach (IMLA; FSCA). 104. Anidrytus angustulus Gerstaecker Fig. 89 100. Anidrytus hirschi Strohecker Anidrytus angustulus Gerstaecker 1858:269. Holotype Fig. 100 cf, Colombia (NHML); Arrow 1920:45. Anidrytus hirschi Strohecker 1957:269, fig. 30. Holotype Anidrytus bisignatus Gerstaecker 1858:268. Holotype cf, Peru (HNHM). cf, Colombia (NHML).

Oblong, subparallel. Dark ferruginous below Broadly oblong, moderately convex. Rust-red, and above. Antennomeres I-II red, others black, sides of pro not urn paler, its disc with 2 black spots INSECTA MUNDI, Vol. 11, No.2, June, 1997 177 at anterior end of lateral sulci, finely and rather 5500 ft (17GO m), on fungus, A. D. Mitchell,cr, 29 9 sparsely punctate. Antenna black, antennomere I (FMNH). rufous. Elytra a little wider at base than pronotum, widened in basal third, thence subparallel, rather 107. Anidrytus quadripunctatus Gorham bluntly rounded to apex. Length 5.G-7.5 mm. Anidrytus quadripunctatus Gorham 1886: 159. Holotype Csiki (1901::35) placed A. bisignatus in synon- 9, Brazil: Sta. Catarina, Blumenau, probably in Oberthur colI. (MNHN); location not confirmed. ymy under A. juvenclJ,s, probably on the basis of a remark by Gerstaecker. Arrow (1920) suggested Oblong or somewhat ovate. Venter red, legs red A. bisignatus A. angustu the conspecificity of and with bases of tibiae slightly infuscate. Antenna Ius and my study of the types supports his observa- with first 4 or 5 antennomeres rufous, the others A. bisig tion. Although I damaged the aedeagus of black. Dorsum rust-red with coppery pubescence, natus in extracting it, I have no doubts about the pronotum with 4 rather large round black spots, structural identity of the 2 holotypes. one just medial to front end of each lateral sulcus, the to a little of dis cal 105. Anidrytus trinitatis Strohecker, 11. sp. others lateral and in front Fig. 90 pair. Length 7.5-8 mm, width 4.3 mm. Holotype cr, W. 1.; Trinidad, Morne Bleu, 2700 ft. (830 I have seen no males. Brazil: Parana, Caviuna, m) 6.VIII.19G9, H. & A. Howden (CMNC). VI. 1944, A. Mueller, 2 99 (AMNH; FSCA). Paratypes:cr, Venezuela: Barinas, Reserva Forestal Ticoporo, 230 m, 22-28. V.19G8, M. Gelber & J. 108. Anidrytus major Strohecker, n. sp. Salcedo (IZAV);cr, Carabobo, Hac. Montera Montal- Fig. 88 ban, 1300 m, 18.IX.19G8, J. & B. Bechyne (FSCA). Holotype cr, Brazil (NHML).

Broadly oblong, moderately convex. Venter dark Long-oblong, subparallel, elytra rather strong- brown, legs and elytral epipleura reddish brown. ly convex. Venter black, legs and elytral epipleura Antenna black, antennomere I red. Dorsum red- rufous, femoral bases dusky. Antenna slender, all dish brown wi th short, oblique coppery pubescence. basal antennomeres longer than wide, antenno- Pronotum with sides parallel basad, well rounded mere IX longer than wide, its medio-distal angle to front angles, hind angles rectangular, lateral somewhat acute but short; antennomeres I-VI ru- sulci linear, reaching middle of disc, which is but fous, others black. Head rufous. Pronotum 2X as little elevated, finely punctate, with a small black wide as long, red with rectangular black patch spot at front end of each sulcus. Elytra about 3X as between lateral sulci, which are long and rather long as pronotum, sides subparallel but continuosly wide, roundly expanded at base; punctation of disc rounded and rather abruptly convergent to apex. rather open, that of sides dense; sides parallel Length 8.2 mm, width 4.9 mm. toward base, well rounded to front angles. Scutel- Very similar in all features toA. angustuIus, of lum black. Elytra more than 3X as long as prono- which it may yet prove to be a subspecies. tum, black, with base, side margin and apex red, finely and densely punctate. Front tibia of male 106. Anidrytus guatemalae Arrow enlarged distad, with small tooth below. Length 8 Fig. 91 mm, width 4.G mm. Anidrytlls gllatemalae Arrow 1920:46. Holotype cr, Gua- temala: San Joachin (NHML). 109. Anidrytus humilis Gorham Anidrytlls liquefactlls (in part) Gorham 1889: 12G. Fig. III Anidrytlls ? sp. Gorham 1889: 128. Anidrytus humilis Gorham 1873:48. Lectotype cr and paralectotype 9, Nicaragua: Chontales, Janson Long-oblong, elytra about 4X as long as prono- (NHML). tum. Rust-red, antennomeres VI-XI black. Gorham noted the type specimen as having disc ofpronotum Oblong, subparallel, moderately convex, Red- infuscate but this may be an artifact. Length 8 mm, dish black with golden pubescence. Antennomeres width 4.5 mm. I-V red, others black. Pronotum 2X as wide as long, Honduras: Morazon, ridge below LaManzanita "narrowed from base to middle, and thence more and C. Uyuca, 5 km SW Suyapa, 5400 ft (1GGOm), suddenly to the apical angles; hind angles acute .... " slab pile, 5.VIII.1948, 1'. H. Hubbell, 2 crcr(MMZ; Elytra subparallel for two thirds their length, grad- FSCA). Guatemala: Zacapa, Sierra de las Minas, 178 Volume 11, No.2, June, 1997, INSECTA MUNDI ually narrowed to apex. Front tibia of male with recumbent. Antennomeres I-V dark red, others small tooth at distal quarter. Length 7.5 mm. black, antennomeres VII-VIII each longer than wide, antennomeres IX-X strongly transverse, 110. Anidrytus decoratus Arrow medially acuminate. In the specimens from Coroico Fig. 112 the elytral base is broadly red, enclosing a large Anidrytus depressns Gorham 1889: 128. Holotype 0', black spot. Length 5.7-7.2 mm, width 3.5-4 mm. Guatemala: EI Tumbador, Champion (NHML). Not In all features, including aedeagus, this species of Gorham 1875. humeralis. Anidrytus humerosus Anidrytus decoratns Arrow 1920:46 (nomen nov.). is similar to A. is slightly more elongate and antennomere VIII is not A. humeralis marginatus. Structurally similar, including aedeagus, to A. subglobose as in and A. hwnilis. Dorsum black, front angles of pronotum, 113. Anidrytus arrowi Strohecker base and apex of elytra reddish orange. The pale Fig. 109 area of pronotum extends obliquely basad to reach Anidrytns arrowi Strohecker 1957:269, fig. 31a. Holo- hind angles. The pale areas of elytra are extensive, type 0', Brazil: Espirito Santo, Staudinger (HNHM). that of base encloses a large black spot. Length 7.5 mm, width 4.2 mm. Oblong, subparallel, moderately convex. Upper Guatemala: Zapate, Escuintla, S.VII.194S, R. surface ferruginous, disc of pronotum and elytra D. Mitchell, Q (FMNH); Moca, 31.VIII.1947, C. & vaguely infuscate. Antennomeres I-V rufous, oth- P. Vaurie, 2 Q Q (MiNH; FSCA).; label indecipher- ers black. Length G.1 mm, width 3.3. mm. able, 0' (SMTD). 114. Anidrytus latus (Chevrolat) 111. Anidrytus humeralis Arrow Fig. lOS Fig. 110 Lycoperdina lata Chevrolat 1844:317, pI. 50, fig. 11. Anidrytus lWl1/.eralis Arrow 1920:47. Lectotype 0', Equa- Anidrytns latns: Gerstaecker 1858:265. dol', Buckley (NHML). Paralectotypes;O', Equador, Buckley; Q, Equador, Macas, Buckley; Q , I have seen no type material and my determina- Colombia; Q, Peru (NHML). A syntype 0' is refer- tion of the species is based on the Gerstaecker able to the species next. described. "type". Rather narrowly oblong and but moderatly Broadly oblong, subparallel. Venter ferrugi- convex. Reddish brown, sides of pronotum, elytral nous. Antenna rather stout, antennomere VIII short, apex, legs and venter rufous, meso- and metaster- transverse, antennomeres IX-X strongly transverse num darker. Dorsum with short, rather sparse and medially acuminate, antennomeres I-Vrufous, coppery pubescence. Antenna fairly stout, anten- the others black. Pronotum black, sides reddish. nomeres VI-VIII subglobose, antennomeres I-V Elytra black, umbo and apex rufous. Front tibia of rufous, others black. Pronotum with sides strongly male widened distad, with a very small tooth before rounded to the short front angles, parallel basad to apex. Length 5.5-7 mm, width 3-4 mm. the rectangular hind angles, lateral sulci wide, Peru: Loreto, Estiron, Rio Ampiyacu, 13.XI- rather short. Front tibia of male gradually widened 9.XII.19Gl, B. Malkin, O'(FMNH). from base, with an angular expansion at apical Very similar to A. marginaf,us (Fabr.). third, hind tibia incurved at apex. Length G-7 mm, 112. Anidrytus humerosus Strohecker, n. sp. width 3.1-3.G mm. Fig. 113 A male from Rio de Janeiro (FSCA) conforms Holotype 0' and allotype Q, Peru: Loreto, Estiron, Rio closely to the Gerstaecker "type" but has entire Ampiyacu, 13.Xr-9.XII.1961, B. Malkin (FMNH). prothorax, meso- and metasternum and first ster- Paratypes: 30'0', 3 Q Q wit.h same data as holotype; nite quite black, legs and elytral epipleura rufous. 20'0', Loreto, 20 km from Ucayali on Rio Calleria, Colonia Calle ria, 10. IX-15.X. 1961, B. Malkin 115. Anidrytus pardalinis Strohecker (FMNH; FSCA);O', Peru (NHML). Bolivia: Coroico, Fig. 131 0' Q (FSCA). Anidrytns pardalinns Strohecker 1943:388, fig. 12f, g. Holotype 0', Costa Rica, F. Nevermann (USNM Oblong, subparallel. Venter and legs brown. 55888). Dorsum black with head, front angles of prono tum, elytral umbo amd apex reel; pubescence fine, gray, INSECTA MUNDI, Vol. 11, No.2, June, 1997 179

Oblong or somewhat oval, rather strongly con- 118. Anidrytus nigropiceus Gorham vex. General color deep rust-red, dorsum with Fig. 115 short, declivent, coppery pubescence. Pronotum with 2 large quadrate black pa tches on base. Elytra Anidrytus nigropiceus Gorham 1889: 126. Lectotype d' with base, most of suture and 2 round spots black, and 1 paralectotype d', Panama: Bugaba, Champion the anterior spot near mid-length, the posterior (NHML). spot smaller and on apical slope. Front tibia of rna Ie gradually widened in distal half, with a sharp tooth Broadly oblong, subparallel. Dark brown be- on inner edge. Length 8 mm, width 4 mm. low. Dorsum black. Antennomeres I-V and margins of pronotum and elytra reddish black. Lateral su lci 116. Anidrytus grandis Strohecker, n. sp. of pronotum very deep, almost reaching middle of Fig. 132 disc. Front tibia of male angulately widened beyond Holotype d', French Guiana: Cayenne, with Gorham's middle. Length 7 mm, width 4.6 mm. label A. marginatlLs (FSCA ex Janson). 119. Anidrytus fuliginosus (Guerin- Long-oval, rather strongly convex. Meso- and Meneville), new combination metasternum and first 2 sternites black; other E(pipOClLS) fuliginosus Guerin-Meneville 1857:266. Type parts of venter, including legs and epipleura ru- (sex ?), Colombia: Cartagena, Dejean ColI. fous. Dorsum black with sides of pronotum broadly, elytral base, side margin and apex narrowly red. Body blackish brown, antenna, legs, elytral Antennomeres III-V subequal, each hardly 2X as epipleura, sides of sterna and abdomen paler brown. long as wide, antennomeres VI-VIII similar to each Head, pronotum and elytra black, a little shining, other, slightly shorter than antennomere IV, an- punctured, a little pubescent. Length 6 mm, width nigricans) tennomere IX about as wiele at apex as long, its 3.66 mm. Very close to (A. in form. mediodistal angle very short. Pronotum relatively I have not recognized this insect among the small, 2X as wiele as long, sides parallel basad, material examined. It can hardly be the same as the Epipocus rounded to obtuse front angles, marginal bead specimens described by Gerstaecker as fuliginosus, rather wiele in front, evanescent at rectangular for which a new name has been pro- hind angles, lateral sulci long and wide, slightly posed (Strohecker 1977:323). convergent. Elytra 4X as long as pronotum, wid- 120. Anidrytus spadiceus (Guerin-Melleville) ened from base in anterior third, thence gradually narrowed caudad, densely andsubrugosely. Length E(pipocus) spadiceus Guerin-Meneville 1857:266. Type 8 mm, width 4.7 mm. (sex ?), Colombia, Dejean colI. Anidrytus spadiceus: Csiki 1901:35. 117. Anidrytus cardisoma Strohecker, n. sp. Fig. 114 Body black, mouthparts, legs and elytral epi- Holotype d', Bolivia: Coroico (FSCA). pleura yellowish brown. Pronotum and elytra very finely punctured, pubescent. Length 5.5mm, width Oval, moderately convex, elytra weakly flat- 3.25 mm. Very close to A. fuliginosus, possibly a tened. Sides of pronotum, elytral epipleura, legs variety. and distal sternites rust-red. Thoracic sterna and This brief diagnosis has not been sufficient for first sternite black. Dorsum generally black, head me to identify specimens as this species. and sides of pronotum reddish. Antenna long, antennomeres I-IV dark red, antennomeres V-XI black, Ackllow ledgments anntennomeres III-IV subequal, each more than I (Paul E. Skelley) thank R. H. Arnett, M. C. 2X as long as wiele, antennomeres V-VIII secularly Thomas, R. E. Woodruff, Florida State Collection of shorter but each longer than wide. Pronotum par- Arthropods; R. A. B. Leschen, Michigan State Uni- allel to rectangular hind angles, rounded to obtuse versity; and S. A. Slipinski, Polish Academy of front angles, lateral sulci, reaching middle of disc. Sciences, for their valuable editorial comments. Elytra 3X as long as pronotum, cordiform, widest in Florida Department of Agriculture and Consumer basal third, rather rapidly narrowed to apex. Front Services, Division of Plant Industry, Entomology tibia of male gradually but not greatly widened Contribution No. 840. distad, hind tibia weakly curved. Length 7 mm, width 4.3 mm. 180 Volume 11, No.2, June, 1997, INSECTA MUNDI

References Guerin-Meneville, F. E. 1857. Materiaux pour une Arrow, G. J. 1920. A contribution to the classification monographie des Coleopteres du groupe des Eumor- ofcoleopt.erous family Endomychidae. Transact.ions phides et plus specialement du genre Eumorphus. of the Entomological Society of London 1920:1-83,1 Archives ent.omologiques 1:237-280, 1 pI. p!. Kirsch, T. 1876. Beitrage zur Kenntnis der peruanis- Chapuis, F. 1876. In J. T. Lacordaire. Histoire naturel- chen Kaferfsuna, auf Dr. Abendroth's Sammlung Ie des Insectes, Genera des Coleopteres, Vo!' XII. basirt, Sechtes Stuck. Deutsche Entomoligische Paris, 374 pp. Zeitschrift XX:81-133. Chevrolat, L. A. A. 1844a. In F. E. Guerin-Meneville. Olivier, A. G. 1791-1792. In Encyclopedie Methodique. Iconographie du Regne Animal de Cuvier. Vol. 7. Dictionnaire des Insectes. Vol. VI. Paris. Paris, 576 pp., 104 pIs. Olivier, A. G. 1808. Entomologie ou histoire naturelle Chevrolat, L. A. A. 1844b. Dictionnaire universel des Insect.es. Pars VI. Paris, pp. 613-1104, 41 pIs. d'Histoire naturelle. Vo!' V. Perty, J. A. M. 1832. Delectus animalium articula- Csiki, E. 1901. Catalogus Endomychidarum. Ter- torum, quae in itinere per Brasiliam-collegerunt meszetrajzi Fuzetek XXIV: 1-53. Spix et Martius. Fasc. 2:61-124, 40 pIs. Erichson,W. F. 1847. Conspectus insectorum coleopter- Pic, M. 1931. Nouveautes diverses. Melanges Exotico- orum quae in Republica Peruana observata sunt. entomologiques 58. Moulins, 36 pp. Archiv flir Naturgeschichte XIII:67-185. Strohecker, H. F. 1943. Some fungus beetles of the Fabricius, J. C. 1801. Systema eleut.heratorum. Vol. I, family Endomychiclae in the U.S. National Muse- Kiliae, xxiv + 506 pp. um. Proceedings of the United States National Gerstaecker, A. 1858. Entomographien I, Monograph- Museum 93:381-392. ie del' Endomychiden. Leipzig, xii + 433 pp., 3 pIs. Strohecker, H. F. 1953. Genera Insectorum. Fasc. 210. Gorham, H. S. 1873. Enclomycici Recitati. London, 64 Endomychiclae. Bruxelles, 140 pp., 5 pIs. pp. 1 pI. Strohecker, H. F. 1957. New species of Endomychidae Gorham, H. S. 1874. Descriptions of new species of in the collections of the Hungarian National Muse- Encomycici. Transactions of the Entomological So- um and other institutions. Annales Historico-Natu- ciety of London 1874:437-446. rales Musei Nationalis Hungarici, Series Nova, Gorham, H. S. 1875. Descriptions of new species of 8:259-283. Endomycici. Transactions of the Entomological So- Strohecker, H. F. 1959. Neue Arten von Endomy- ciety of London 1875: 11-22. chiden im Berliner Museum. Mitteilungen aus dem Gorham, H. S. 1886. On new genera and species of Zoologischen Museum in Berlin 35:177-186. Endomychidae. Proceedings of t.he Zoological Soci- Strohecker, H. F. 1977. Revision of the genus Epipo etyofLondon 1886:154-163, 1 pI. cus. Transactions of the American Ent.omological Gorham, H. S. 1887-99. Biologia Central i-Americana. Society 103:303-325. Insecta, Coleoptera Vol. VII. London, xii + 276 pp., Strohecker, H. F. 1979. A new species of Anidrytus 13 pIs. from southern Brasil with notes on several other species. Revue Suisse Zoologie 86:715-717. INSECTA MUNDI, Vol. 11, No.2, June, 1997 181

5(51) 1(38) 2(39) 3(15) 4(51)

6(40) 7(40)

14 (22) 15 16 11 (35) (26) (23) 13(37)

Figs. 1-16. Elytral pattern. 1) Epopterus fasciatus fasciatus -lectotype, 2) E. fasciatus al!gustatus - holotype,3) E. zebra - holotype, 4) E. similis - holotype, 5) E. similis - Ecuador, 6) E. variegatus - Peru:Callanga, 7) E. variegatus - Amazon, 8) E. retiCILlatus - holotype, 9) E. cyclicus -holotype, 10) E. cordatus -holotype, 11) E. deltoideus -holotype, 12) E. decempul!ctatus - Trinidad, 13) E. decempul!ctatus - Colombia: Cucuta, 14) E. ? vicil!uS - Peru: Callanga, 15) E. obliquus - holotype, 16) E. l!otatus - holotype. 182 Volume 11, No.2, June, 1997, INSECTA MUNDI

30(10)

~31(12)

Figs. 17-32. Elytral pattern. 17) Epopterus histrio - topotype, 18) E. rubiginosus - holotype, 19) E. parvus -para type, 20) E. quaesitus -holotype, 21) E. [allax -holotype, 22) E. pictus sensu Gerstaecker -Brazil, 23) E. biocuZatus -holotype, 24) E. ocellatus sensu Gerstaecker - Brazil, 25) E. dilectus -holotype, 26) E. vemicatus -holotype, 27) E. quechuanus -holotype, 28) E. picticollis -holotype, 29) E. signaticollis - Panama, 30) E. sigllaticollis - Colombia, 31) E. testudinarius - Colombia, 32) E. gracilis - holotype. INSECTA MUNDI, Vol. 11, No.2, June, 1997 183

36(7) 33 34(2) 35(5) 37(8} (1 )

48 (20)

55 54 (19) (45) 53(37)

Figs. 33-57. Aedeagus, dextro-ventral aspect. 33) Epopterus ocellatus sensu Gerstaecker -Brazil: Corupa, 34) E. allllulatus- Guyana, 35) E. pm·titus partl:[us - Honduras, 36) E. partitus hirschi - holotype, 37) E. tigrillus - holotype, 38) E. gemilllls - holC' type, 39) E. sigtlctticollis - Colombia: Villavicencio, 40) E. vacuus - holotype, 41) E. Illldulatus - Argentina: Tucuman, 42) E. pietus sensu Gerstaecker - Brazil, 43) E. vClriegatus - holotype, 44) E. eomptus - topotype (syntype?), 45) E. testuditlctrius -lectotype, 46) E. fasciatus fasciatus - lectotype, 47) E. egallus -lectot)1le, 48) E. cUCllllillus - holotype, 49) E. dileetus - holotype, 50) E. dives - holotype, 51) E. clara - Brazil: Para, 52) E. histrio - Brazil: Amapa, 53) E. decempu.lletatus - Colombia: Puerto Berrio, 54) E. ephippiger - holotype, 55) E. bioculatus holotype, 56) E. lIigrocillctus - holot)1le, 57) E. trallsversus - holotype. 184 Volume 11, No.2, June, 1997, INSECTA MUNDI

64(60) 65 (57) 67(25)

70 (18)

76 (16) 77 (24)

Figs. 58-78. Aedeagus, dextro-ventral aspect. 58) Epopterus cordatus - holotype, 59) E. lIigrocillctus - Colombia: Leticia, 60) E. /Zavolilleatus - holotype, 61) E. deltoideus - holotype, 62) E. cyclicus - holotype, 63) E. similis - holotype, 64) E. mimeticus • holotype, 65) E. /Zavollotatus - holotype, 66) E. reticulatus - holotype, 67) E. longus - holotype, 68) E. batesi - holotype, 69) E. zebra - holotype, 70) E. lIigerrimus - holotype, 71) E. lIotatus - holotype, 72) E. obliquus - holotype, 73) E. parvus - holotype, 74) E. gracilis - holotype, 75) E. quec!luallus - holotype, 76) E. picticollis - holotype, 77) E. COIl!USUS - holotype, 78) E. crypticus - holotype. INSECTA MUNDI, Vol. 11, No.2, June, 1997 185

79 (30)

Figs. 79-87. Aedeagus, dextro-vental aspect. 79) Epopterus? vemicatl1s - Brazil: Rio de Janeiro, 80) E. atrivelltris - holotype, 81) E. loretellsis - holotype, 82) E. c(llierialllls - holotype, 83) E. (lIIOIII(l/I1S- holotype, 84) E. p(lntherilll1s - Panama, 85) E. aravacus - holotype, 86) E. decelllpnllctatus -lectotype, 87) E. sl1/JII/(/cn/atlls - holotype. 186 Volume 11, No.2, June, 1997, INSECTA MUNDI

91 (106)

93 94 95 96 (66) (67) (101) (102)

100 (100)

105 (96)

Figs. 88-lOG. Aedeagus, dextro-ventral aspect. 88) Al!idrytus major - holotype, 89) A. juvel!cus ? - holotype of A. al!gllstuius, 90) A. tril!itatis - holotype, 91) A. guatelllaiae - holotype, 92) A. bipul!ctatus -holotype, 93) A. piagiatus -holotype, 94) A. lIimbiferus - holotype, 95) A. iiquefcfCtus, 9G) A. mexicallus - holotype, 97) A. cOlltmctus - Costa Rica: Turrialba, 98) A. ovatuius -lectotype, 99) A. lIigriwlls - Rio de Janeiro, 100) A. hirschi - holotype, 101) A. amCJ.Zollicus - Brazil: Fonte Boa, 102) A. cllpreatus - holotype, 103) A. epliippium- French Guiana, det. Gorham, 104) A. iugubris - no locality, det. Gorham, 105) A. batesi - holotype, 106) A. circumcillctus - holotype. INSECTA MUNDI, Vol. 11, No.2, June, 1997 187

110 (111)

113 (112) 116 (8S)

119 121 (84) (77)

122 123 126 127 (88) (81 ) (76) (80)

Figs. 107-127. Aedeagus, dextro-ventral aspect. 107) Anidrytus bechyneorum - holotype, 108) A. latus - Gerstaecker "type", 109) A. arrowi - holotype, 110) A. IwmeraUs - lectotype, Ill) A. humilis -lectotype, 112) A. decoratlls - holotype, 113) A. humeroslls - holotype, 114) A. cardisoma - holotype, 115) A. nigropiceus -lectotype, 116) A. pilosus -lectotype, 117) A. parallelus - holotype, 118) A. stellllS - holotype, 119) A. helllolus - Venezuela, comp. lectotype, 120) A. concolor - Peru, comp. holotype, 121) A. gibbosus - holotype, 122) A. atratillus - Brazil, comp. holotype, 123) A. [IlSCUS-lectotype, 124) A. compCtctus - holotype, 125) A. glaber - Peru, comp. holotype, 126) A. cOllsobrillus - holotype, 127) A. tenllipes - holotype. 188 Volume 11, No.2, June, 1997, INSECTA MUNDI

128 (74) \ \ .. ' 129 (73)

Figs. 128-1~H.Aedeagus, dextro-ventral aspect. 128) Anidrytus ullicolor - Peru, compo holotype, 129) A. poreus - holotype, 130) A. gerstaeclwri -lectotype, 1:31) A. pardo/inns - holotype, 1:32) A. grandis - holotype, 1:3:3)A. parki - holotype, 1:34) A. variegatns - Rio de Janeiro.