Puget Sound Polychaetes
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Annelida, Phyllodocida)
A peer-reviewed open-access journal ZooKeys 488: 1–29Guide (2015) and keys for the identification of Syllidae( Annelida, Phyllodocida)... 1 doi: 10.3897/zookeys.488.9061 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research Guide and keys for the identification of Syllidae (Annelida, Phyllodocida) from the British Isles (reported and expected species) Guillermo San Martín1, Tim M. Worsfold2 1 Departamento de Biología (Zoología), Laboratorio de Biología Marina e Invertebrados, Facultad de Ciencias, Universidad Autónoma de Madrid, Canto Blanco, 28049 Madrid, Spain 2 APEM Limited, Diamond Centre, Unit 7, Works Road, Letchworth Garden City, Hertfordshire SG6 1LW, UK Corresponding author: Guillermo San Martín ([email protected]) Academic editor: Chris Glasby | Received 3 December 2014 | Accepted 1 February 2015 | Published 19 March 2015 http://zoobank.org/E9FCFEEA-7C9C-44BF-AB4A-CEBECCBC2C17 Citation: San Martín G, Worsfold TM (2015) Guide and keys for the identification of Syllidae (Annelida, Phyllodocida) from the British Isles (reported and expected species). ZooKeys 488: 1–29. doi: 10.3897/zookeys.488.9061 Abstract In November 2012, a workshop was carried out on the taxonomy and systematics of the family Syllidae (Annelida: Phyllodocida) at the Dove Marine Laboratory, Cullercoats, Tynemouth, UK for the National Marine Biological Analytical Quality Control (NMBAQC) Scheme. Illustrated keys for subfamilies, genera and species found in British and Irish waters were provided for participants from the major national agencies and consultancies involved in benthic sample processing. After the workshop, we prepared updates to these keys, to include some additional species provided by participants, and some species reported from nearby areas. -
Phylogeny and Biogeography of Branchipolynoe
Phylogeny and Biogeography of Branchipolynoe (Polynoidae, Phyllodocida, Aciculata, Annelida), with Descriptions of Five New Species from Methane Seeps and Hydrothermal Vents Johanna Lindgren, Avery Hatch, Stéphane Hourdez, Charlotte Seid, Greg Rouse To cite this version: Johanna Lindgren, Avery Hatch, Stéphane Hourdez, Charlotte Seid, Greg Rouse. Phylogeny and Biogeography of Branchipolynoe (Polynoidae, Phyllodocida, Aciculata, Annelida), with Descriptions of Five New Species from Methane Seeps and Hydrothermal Vents. Diversity, MDPI, 2019, 11 (9), pp.153. 10.3390/d11090153. hal-02313505 HAL Id: hal-02313505 https://hal.sorbonne-universite.fr/hal-02313505 Submitted on 11 Oct 2019 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. diversity Article Phylogeny and Biogeography of Branchipolynoe (Polynoidae, Phyllodocida, Aciculata, Annelida), with Descriptions of Five New Species from Methane Seeps and Hydrothermal Vents Johanna Lindgren 1, Avery S. Hatch 1, Stephané Hourdez 2, Charlotte A. Seid 1 and Greg W. Rouse 1,* 1 Scripps Institution of Oceanography, -
RECON: Reef Effect Structures in the North Sea, Islands Or Connections?
RECON: Reef effect structures in the North Sea, islands or connections? Summary Report Authors: Coolen, J.W.P. & R.G. Jak (eds.). Wageningen University & Research Report C074/17A RECON: Reef effect structures in the North Sea, islands or connections? Summary Report Revised Author(s): Coolen, J.W.P. & R.G. Jak (eds.). With contributions from J.W.P. Coolen, B.E. van der Weide, J. Cuperus, P. Luttikhuizen, M. Schutter, M. Dorenbosch, F. Driessen, W. Lengkeek, M. Blomberg, G. van Moorsel, M.A. Faasse, O.G. Bos, I.M. Dias, M. Spierings, S.G. Glorius, L.E. Becking, T. Schol, R. Crooijmans, A.R. Boon, H. van Pelt, F. Kleissen, D. Gerla, R.G. Jak, S. Degraer, H.J. Lindeboom Publication date: January 2018 Wageningen Marine Research Den Helder, January 2018 Wageningen Marine Research report C074/17A Coolen, J.W.P. & R.G. Jak (eds.) 2017. RECON: Reef effect structures in the North Sea, islands or connections? Summary Report Wageningen, Wageningen Marine Research, Wageningen Marine Research report C074/17A. 33 pp. Client: INSITE joint industry project Attn.: Richard Heard 6th Floor East, Portland House, Bressenden Place London SW1E 5BH, United Kingdom This report can be downloaded for free from https://doi.org/10.18174/424244 Wageningen Marine Research provides no printed copies of reports Wageningen Marine Research is ISO 9001:2008 certified. Photo cover: Udo van Dongen. © 2017 Wageningen Marine Research Wageningen UR Wageningen Marine Research The Management of Wageningen Marine Research is not responsible for resulting institute of Stichting Wageningen damage, as well as for damage resulting from the application of results or Research is registered in the Dutch research obtained by Wageningen Marine Research, its clients or any claims traderecord nr. -
Zootaxa, Loandalia (Polychaeta: Pilargidae)
Zootaxa 1119: 59–68 (2006) ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ ZOOTAXA 1119 Copyright © 2006 Magnolia Press ISSN 1175-5334 (online edition) New species of Loandalia (Polychaeta: Pilargidae) from Queensland, Australia SHONA MARKS1 & SCOTT HOCKNULL2 1 S. A. Marks. CSIRO Marine Research, PO Box 120, Cleveland QLD 4163. [email protected]. 2 S. A. Hocknull. Queensland Museum, 122 Gerler Rd Hendra QLD 4711. [email protected] Abstract Two new species of Loandalia are described from Queensland, Australia. Loandalia fredrayorum sp. nov. is described from Moreton Bay, south eastern Queensland and is distinguished from all other species of Loandalia by the presence of singular palpostyles; uniramous parapodia at chaetiger 1; an emergent notopodial spine at chaetiger 9; neurochaetae numbering 20–24; ventral cirri begin on chaetiger 7 and the pygidium with two lateral papillae-like anal cirri. Loandalia gladstonensis sp. nov. is described from Gladstone Harbour, central eastern Queensland and is distinguished from all other species of Loandalia by the presence of bifid palpostyles; chaetiger 1 uniramous with remaining chaetigers biramous; an emergent notopodial spine from chaetiger 7–8; ventral cirri present from chaetiger 5 and neurochaetae numbering 5–6. Key words: Loandalia fredrayorum sp. nov., Loandalia gladstonensis sp.nov., Pilargidae, Queensland, Australia, new species, systematics. Introduction Saint-Joseph (1899) established the Pilargidae for the new species Pilargis verrucosa Saint-Joseph. Prior to this, pilargids had been placed in several different families including the Syllidae, Hesionidae and Polynoidae (Licher & Westheide 1994). Recent cladistic analyses of the Phyllodocida firmly recognise Pilargidae as a distinct clade (Glasby 1993; Pleijel & Dahlgren 1998). -
Neanthes Limnicola Class: Polychaeta, Errantia
Phylum: Annelida Neanthes limnicola Class: Polychaeta, Errantia Order: Phyllodocida, Nereidiformia A mussel worm Family: Nereididae, Nereidinae Taxonomy: Depending on the author, Ne- wider than long, with a longitudinal depression anthes is currently considered a separate or (Fig. 2b). subspecies to the genus Nereis (Hilbig Trunk: Very thick segments that are 1997). Nereis sensu stricto differs from the wider than they are long, gently tapers to pos- genus Neanthes because the latter genus terior (Fig. 1). includes species with spinigerous notosetae Posterior: Pygidium bears two, styli- only. Furthermore, N. limnicola has most form ventrolateral anal cirri that are as long as recently been included in the genus (or sub- last seven segments (Fig. 1) (Hartman 1938). genus) Hediste due to the neuropodial setal Parapodia: The first two setigers are unira- morphology (Sato 1999; Bakken and Wilson mous. All other parapodia are biramous 2005; Tusuji and Sato 2012). However, re- (Nereididae, Blake and Ruff 2007) where both production is markedly different in N. limni- notopodia and neuropodia have acicular lobes cola than other Hediste species (Sato 1999). and each lobe bears 1–3 additional, medial Thus, synonyms of Neanthes limnicola in- and triangular lobes (above and below), called clude Nereis limnicola (which was synony- ligules (Blake and Ruff 2007) (Figs. 1, 5). The mized with Neanthes lighti in 1959 (Smith)), notopodial ligule is always smaller than the Nereis (Neanthes) limnicola, Nereis neuropodial one. The parapodial lobes are (Hediste) limnicola and Hediste limnicola. conical and not leaf-like or globular as in the The predominating name in current local in- family Phyllodocidae. (A parapodium should tertidal guides (e.g. -
OREGON ESTUARINE INVERTEBRATES an Illustrated Guide to the Common and Important Invertebrate Animals
OREGON ESTUARINE INVERTEBRATES An Illustrated Guide to the Common and Important Invertebrate Animals By Paul Rudy, Jr. Lynn Hay Rudy Oregon Institute of Marine Biology University of Oregon Charleston, Oregon 97420 Contract No. 79-111 Project Officer Jay F. Watson U.S. Fish and Wildlife Service 500 N.E. Multnomah Street Portland, Oregon 97232 Performed for National Coastal Ecosystems Team Office of Biological Services Fish and Wildlife Service U.S. Department of Interior Washington, D.C. 20240 Table of Contents Introduction CNIDARIA Hydrozoa Aequorea aequorea ................................................................ 6 Obelia longissima .................................................................. 8 Polyorchis penicillatus 10 Tubularia crocea ................................................................. 12 Anthozoa Anthopleura artemisia ................................. 14 Anthopleura elegantissima .................................................. 16 Haliplanella luciae .................................................................. 18 Nematostella vectensis ......................................................... 20 Metridium senile .................................................................... 22 NEMERTEA Amphiporus imparispinosus ................................................ 24 Carinoma mutabilis ................................................................ 26 Cerebratulus californiensis .................................................. 28 Lineus ruber ......................................................................... -
Polychaetes of the Western Black Sea Meiobenthos
J. Black Sea/Mediterranean Environment Vol. 15: 109- 121 (2009) Meiobenthic bristle worms (Polychaeta) of the western Black Sea shelf Ludmila V. Vorobyova* and Olena S. Bondarenko *Odessa Branch, A.O. Kovalevsky Institute of Biology of the Southern Seas (OB IBSS), National Academy of Sciences of Ukraine. 37 Pushkinskaya Street, 65125 Odessa. Abstract The species composition, frequency and quantitative distribution of polychaetes (larvae, adult forms) which in size belong to meiobenthos have been studied. Thirty four species of polychaetes, six of which belong to eumeiobenthos have been discovered. In species composition the Bulgarian shelf (33 species) differs significantly from the Romanian (22 species) and Ukrainian (21 species). The mean density of assemblages is higher on the Bulgarian shelf. Maximum indices of abundance occur at a 15 m depth. Keywords: Black Sea, western shelf, meiobenthos, polychaetes, eumeiobenthos, pseudomeiobenthos. Introduction The western Black Sea shelf (WBS) is of special interest for ecological research. Very diverse environmental conditions from the Dnieper-Bug area to Burgas Bay related to the level of impact of river runoff of the Dnieper, Dniester and Danube are observed. The dynamics of hydrological and hydrochemical parameters determine the marked variability of characteristics of sea bottom invertebrates. The present day state of meiobenthic polychaetes was analyzed. Small (up to 2-5 mm) meiobenthic invertebrates of different water bodies have been described by Mare 1942, Hulings and Gray 1971. The term meiobenthos in scientific literature has been introduced much later than the terms benthos and plankton. The former was proposed by Peterson in 1911 *Corresponding author: [email protected] 109 and the latter by Hensen in 1887. -
Identification Guide to the Planktonic Polychaete Larvae Around the Island of Helgoland (German Bight)
HELGOL.~NDER MEERESUNTERSUCHUNGEN Helgol/inder Meeresunters. 48, 1-58 (1994) Identification guide to the planktonic polychaete larvae around the island of Helgoland (German Bight) S. Plate* & E. Husemann* * Biologische Anstalt Helgoland (Meeresstation); D-27483 Helgoland, Federal Republic of Germany ABSTRACT: The purpose of this work is to provide the means of identifying the planktonic larvae of the polychaete species appearing in the plankton around the island of Helgoland (North Sea). During a three-year survey in this area, the larvae of 54 species out of 24 families belonging to the orders Orbiniida, Spionida, Capitelhda, Phyllodocida, Oweniida, Terebelhda, Sabelhda and the former Archiannelida have been recorded. Illustrated keys to the families, genera and species are presented. To facilitate the identification, additional descriptions and information about the seasonal appearance of the species are given. INTRODUCTION More than 13 000 species of polychaetous annelids take part in the marine benthos communities worldwide. Their distribution, species composition and population density are monitored within various benthos surveys. For the North Sea, especially the German Bight and the Wadden Sea, much information about the benthic polychaete fauna is available (Caspers, 1950; Stripp, 1969; DSrjes, 1977; Rachor & Gerlach, 1978; Gillandt, 1979; Salzwedel et al., 1985; Rachor, 1990; Bosselmann, 1991; Kr6ncke, 1991). In contrast, the holoplanktonic polychaete species and the meroplanktonic polychaete larvae, which are only part of the plankton during a more or less expanded phase of their ontogenesis, have never received much attention. Meroplanktonic polychaete larvae are seldomly recorded during studies monitoring the North Sea plankton (Smidt, 1951; Giere, 1968; Fransz, 1981; Bosselmann, 1989; Belgrano et al., 1990). -
Polychaete Worms Definitions and Keys to the Orders, Families and Genera
THE POLYCHAETE WORMS DEFINITIONS AND KEYS TO THE ORDERS, FAMILIES AND GENERA THE POLYCHAETE WORMS Definitions and Keys to the Orders, Families and Genera By Kristian Fauchald NATURAL HISTORY MUSEUM OF LOS ANGELES COUNTY In Conjunction With THE ALLAN HANCOCK FOUNDATION UNIVERSITY OF SOUTHERN CALIFORNIA Science Series 28 February 3, 1977 TABLE OF CONTENTS PREFACE vii ACKNOWLEDGMENTS ix INTRODUCTION 1 CHARACTERS USED TO DEFINE HIGHER TAXA 2 CLASSIFICATION OF POLYCHAETES 7 ORDERS OF POLYCHAETES 9 KEY TO FAMILIES 9 ORDER ORBINIIDA 14 ORDER CTENODRILIDA 19 ORDER PSAMMODRILIDA 20 ORDER COSSURIDA 21 ORDER SPIONIDA 21 ORDER CAPITELLIDA 31 ORDER OPHELIIDA 41 ORDER PHYLLODOCIDA 45 ORDER AMPHINOMIDA 100 ORDER SPINTHERIDA 103 ORDER EUNICIDA 104 ORDER STERNASPIDA 114 ORDER OWENIIDA 114 ORDER FLABELLIGERIDA 115 ORDER FAUVELIOPSIDA 117 ORDER TEREBELLIDA 118 ORDER SABELLIDA 135 FIVE "ARCHIANNELIDAN" FAMILIES 152 GLOSSARY 156 LITERATURE CITED 161 INDEX 180 Preface THE STUDY of polychaetes used to be a leisurely I apologize to my fellow polychaete workers for occupation, practised calmly and slowly, and introducing a complex superstructure in a group which the presence of these worms hardly ever pene- so far has been remarkably innocent of such frills. A trated the consciousness of any but the small group great number of very sound partial schemes have been of invertebrate zoologists and phylogenetlcists inter- suggested from time to time. These have been only ested in annulated creatures. This is hardly the case partially considered. The discussion is complex enough any longer. without the inclusion of speculations as to how each Studies of marine benthos have demonstrated that author would have completed his or her scheme, pro- these animals may be wholly dominant both in num- vided that he or she had had the evidence and inclina- bers of species and in numbers of specimens. -
Eteone Lighti Order: Phyllodocida, Phyllodociformia
Phylum: Annelida Class: Polychaeta, Errantia Eteone lighti Order: Phyllodocida, Phyllodociformia A paddleworm Family: Phyllodocidae, Eteoninae Taxonomy: The genus Eteone was revised tentacular cirri (Fig. 2). into three genera (Eteone, Hypereteone, Parapodia: Uniramous with neuropodia only. Mysta) by Wilson (1988) based on anal cirri All but first segment with a flat triangular dor- morphology and the presence and location sal cirrus, about as wide as long (Fig. 3), of proboscis papillae. Thus, E. lighti is these become longer and narrower posterior- sometimes referred to as Hypereteone lighti. ly. The ventral cirrus has a broad base taper- While the presence of three major groups ing to a blunt tip and is shorter than the neu- are apparent, splitting Eteone into these ropodial lobe (Fig. 3). Note: parapodium genera has not been recognized by most should be examined in side view to check for authors and E. lighti is the name most com- flatness, inflatedness, etc. monly seen (Pleijel 1991; Blake 1992; Blake Setae (chaetae): Setae are compound 1997; Blake and Ruff 2007). (Phyllodocidae, Blake 1975b) and consist of long, fine, colorless spinigers (Hartman 1968) Description (Figs. 4a,b). Size: Individuals to 30 mm in length and 1– Eyes/Eyespots: Two eyespots are present 1.5 mm in width (Hartman 1968). A 25-mm on posterior third of the prostomium (Fig. 2a). long Coos Bay specimen weighed 0.17 g Anterior Appendages: Prostomium bears (wet weight, Baker et al. 1970). two pairs of short, conical antennae and ap- Color: Pale or white, deep yellow dorsal pendages on the first segment include two transverse stripes (Hartman 1968) and dor- pairs of short and slender tentacular cirri sal cirri with deep yellow tips. -
An Annotated Checklist of the Marine Macroinvertebrates of Alaska David T
NOAA Professional Paper NMFS 19 An annotated checklist of the marine macroinvertebrates of Alaska David T. Drumm • Katherine P. Maslenikov Robert Van Syoc • James W. Orr • Robert R. Lauth Duane E. Stevenson • Theodore W. Pietsch November 2016 U.S. Department of Commerce NOAA Professional Penny Pritzker Secretary of Commerce National Oceanic Papers NMFS and Atmospheric Administration Kathryn D. Sullivan Scientific Editor* Administrator Richard Langton National Marine National Marine Fisheries Service Fisheries Service Northeast Fisheries Science Center Maine Field Station Eileen Sobeck 17 Godfrey Drive, Suite 1 Assistant Administrator Orono, Maine 04473 for Fisheries Associate Editor Kathryn Dennis National Marine Fisheries Service Office of Science and Technology Economics and Social Analysis Division 1845 Wasp Blvd., Bldg. 178 Honolulu, Hawaii 96818 Managing Editor Shelley Arenas National Marine Fisheries Service Scientific Publications Office 7600 Sand Point Way NE Seattle, Washington 98115 Editorial Committee Ann C. Matarese National Marine Fisheries Service James W. Orr National Marine Fisheries Service The NOAA Professional Paper NMFS (ISSN 1931-4590) series is pub- lished by the Scientific Publications Of- *Bruce Mundy (PIFSC) was Scientific Editor during the fice, National Marine Fisheries Service, scientific editing and preparation of this report. NOAA, 7600 Sand Point Way NE, Seattle, WA 98115. The Secretary of Commerce has The NOAA Professional Paper NMFS series carries peer-reviewed, lengthy original determined that the publication of research reports, taxonomic keys, species synopses, flora and fauna studies, and data- this series is necessary in the transac- intensive reports on investigations in fishery science, engineering, and economics. tion of the public business required by law of this Department. -
THE SUBLITTORAL FAUNA of TWO SANDY BAYS on the ISLE of CUMBRAE, FIRTH of CLYDE by R
.'1. Mar. bioI. Ass. U.K. (1955) 34, 161--180 161 Printed in Great Britain THE SUBLITTORAL FAUNA OF TWO SANDY BAYS ON THE ISLE OF CUMBRAE, FIRTH OF CLYDE By R. B. Clark Department of Zoology, Glasgow University and A. Milne Department of Agriculture, King's College, Newcastle on Tyne (Text-figs. I and 2) INTRODUCTION There is a considerable literature on the ecologyof intertidal animals and a growing one on the sublittoral fauna. Largely because of the difficulty of taking samples in very shallow water, litde attention has been paid to the continuation of the intertidal zonation below the low-water mark. Although incomplete in some respects, the results of the present survey are published, partly to help bridge the gap between studies of sublittoral and intertidal faunas, pardy because it is unlikelythat this surveywill everbe completed and partly because the intertidal fauna of one of the bays is particularly well known. The work was begun in 1938by one of the authors (A.M.) but was discontinued at the outbreak of the late war. Since 1949 further collections have been made and the identity of most of the species taken in the earlier sampling has been checked. The collectionsof animals and a full account of the results have been deposited in the Marine Laboratory at Millport. METHODS None of the larger and more reliable bottom samplers can be operated from the small boat that must be used in shallow water. All samples in the quanti- tative survey were taken by the Robertson mud bucket, that is, a cylindrical bucket about 15 in.