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Please Scroll Down for Article This article was downloaded by: [Deakin University] On: 11 May 2009 Access details: Access Details: [subscription number 907852123] Publisher Taylor & Francis Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK Alcheringa: An Australasian Journal of Palaeontology Publication details, including instructions for authors and subscription information: http://www.informaworld.com/smpp/title~content=t770322720 New Lopingian (Late Permian) rugosochonetid species from Sichuan, South China Monica J. Campi a; Guang R. Shi a a School of Ecology and Environment, Deakin University, Burwood, Australia Online Publication Date: 01 January 2005 To cite this Article Campi, Monica J. and Shi, Guang R.(2005)'New Lopingian (Late Permian) rugosochonetid species from Sichuan, South China',Alcheringa: An Australasian Journal of Palaeontology,29:2,275 — 285 To link to this Article: DOI: 10.1080/03115510508619306 URL: http://dx.doi.org/10.1080/03115510508619306 PLEASE SCROLL DOWN FOR ARTICLE Full terms and conditions of use: http://www.informaworld.com/terms-and-conditions-of-access.pdf This article may be used for research, teaching and private study purposes. Any substantial or systematic reproduction, re-distribution, re-selling, loan or sub-licensing, systematic supply or distribution in any form to anyone is expressly forbidden. The publisher does not give any warranty express or implied or make any representation that the contents will be complete or accurate or up to date. The accuracy of any instructions, formulae and drug doses should be independently verified with primary sources. The publisher shall not be liable for any loss, actions, claims, proceedings, demand or costs or damages whatsoever or howsoever caused arising directly or indirectly in connection with or arising out of the use of this material. New Lopingian (Late Permian) rugosochonetid species from Sichuan, South China MONICA J. CAMPI and GUANG R. SHI CAMP|, M.J. & SHI, G.R., 30.9.2005. New Lopingian (Late Permian) rugosochonetid species from Sichuan, South China. Alcheringa 29, 275-285. ISSN 0311 5518. Two rugosochonetid species, Neochonetes (Huangichonetes) geniculatus sp. nov. and Neochonetes (Zhongyingia) linshuiensis sp. nov., are described from the Lopingian (Late Permian) of the Chuanmu section, Sichuan, South China. Ecological changes from the diverse upper Changhsingian brachiopod palaeocommunity to the depauperate post- extinction brachiopod community are briefly discussed. Monica J. Campi [[email protected]] & Guang R. Shi [[email protected]], School of Ecology and Environment, Deakin University, Melbourne Campus, 221 Burwood Highway, Burwood 3125, Victoria, Australia; received 13.6.2004, revised 19.11.2004. Key words: brachiopods, chonetids, ecology, Lopingian, Permian-Triassic boundary. TWO NEW RUGOSOCHONETID species from The Feixianguan Formation at the Chuanmu the Changhsing and lowermost Feixianguan section is a thick sequence (>1 000 m) of fairly Formations at the Chuanmu section, Sichuan, monotonous thin-bedded marl and micrite. South China are described in this paper. The Brachiopods were only found at the very base of Chuanmu section is located in the Huaying this formation at the Chuanmu section. Early Mountains (Huayingshan) in eastern Sichuan, Triassic sequences are fairly uniform across South about 125 km northeast of Chongqing City (Fig. China (Peng et al. 2001), and were deposited Downloaded By: [Deakin University] At: 04:24 11 May 2009 1). Permian and Triassic sequences are well during a transgressional event and subsequent exposed along a road cutting at this section, highstand following a regression that occurred which is fairly continuous from the Cisuralian towards the end of the Permian (Chert et al. 1998). Liangshan Formation to the Late Triassic. This They are characterised initially by thin bedded work forms part of a larger study on the Permian mudstone and shale, followed by fairly thinly Brachiopoda from the Chuanmu section by MJC. bedded marly to argillaceous micritic facies, and are generally well laminated with little evidence Lithology and biostratigraphy of bioturbation. The new species were collected from the The Changxing Formation is the youngest upper half of the Changxing Formation and the Permian formation in South China, and is lowermost 20 cm of the overlying Feixianguan conformably overlain by Triassic marine sediments at the Chuanmu section. In this section Formation. The abundant and highly diverse and the surrounding area in Sichuan, the brachiopod fauna of the upper half of the Changxing Formation is dominated by cherty Changxing Formation yielded more than 70 limestone with subordinate argillaceous identifiable species (304 individuals) associated limestone and marl, representing a shallow with the two new species. The most common carbonate platform environment (Fig. 2). species was Haydenella kiangsiensis (Kayser, 0311/5518/2005/02275-11 $3.00 © AAP 1883), followed by Spinomarginifera lopingensis 276 MONICA J. CAMPI and G R. SHI ALCHERINGA R ~Carbonifereua ~~---- LiangshanFormation ~~ okouFormation ~~ LongtanFormation mines ~ -_--~_~ Fig. 1A-B. A, Location map of the Chuanmu section in Sichuan. 1: mountains; 2: rivers, 3: provincial boundaries. B, Distribution of formations at the Chuanmu section. 1: gravel road; 2: river. (Kayser, 1883), Prelissorhynchia pseudoutah 20 cm (Bed 178a) of the overlying Feixianguan (Huang, 1933), Neochonetes (Zhongyingia) Formation (Fig. 2). linshuiensis sp. nov., Spinomarginifera alpha The brachiopod fauna in the Feixianguan Huang, 1932, Acosarina minuta (Abich, 1878), Formation at the Chuanmu section is found only Spinomarginifera chengyaoyenensis Huang, at the very base of this formation in one bed only 1932, Crurithyris speciosa Wang, 1955, (Bed 178a, CC- 11). It consists entirely of Permian Haydenella nasuta Zeng, 1993, Linshuichonetes holdover taxa which did not immediately succumb flatus (Shen & Archbold, 2002), Cathaysia to the end-Permian extinction. The brachiopod orbicularis Xu & Grant, 1994, Neochonetes fauna from Bed 178a (CC-11) contains eight (Huangichonetes) cursothornia (Xu & Grant, species along with an unidentifiable specimen Downloaded By: [Deakin University] At: 04:24 11 May 2009 1994), Neochonetes (Huangichonetes) sub- belonging to the Rugosochonetidae. The species strophomenoides (Huang, 1932), Spirigerella? are Cathaysia orbicularis, Spinomarginifera guizhouensis (Liao, 1980b), Huatangia sulcatifera lopingensis, Crurithyris sp., Linshuichonetes (Liao & Meng, 1986) and Orthotetina frechi flatus, Neochonetes (Huangichonetes) sub- (Huang, 1933), which represented just over 60% strophomenoides, Acosarina minuta, Neo- of the collections; the other 60 species make up chonetes (Huangichonetes) geniculatus sp. nov. the remainder of the species diversity. and Spirigerella? sp., in order of decreasing Neochonetes (Zhongyingia) tinshuiensis sp. abundance. nov. is a relatively common species in the upper As conodont work is yet to be fully part of the Changxing Formation at this section, completed on this section, the Permian-Triassic with twelve specimens collected from seven boundary was primarily located through the use horizons in the upper 110 m of the Changxing of lithostratigraphy, biostratigraphy and Formation (Fig. 2). The second new species, eventostratigraphy (Peng & Tong 1999, Peng et Neochonetes (Huangichonetes) geniculatus sp. al. 2001) mainly through correlation of the clay nov. appears to be much rarer, with only two beds immediately above and below the boundary specimens collected from two horizons, one bed. From correlation with the PTBSS (Permian- within 20 cm (Bed 174) of the top of the Triassic Boundary Stratigraphic Set) of Peng et Changxing Formation and the other in the basal al. (2001) the Permian-Triassic boundary has ALCHERINGA NEW PERMIAN BRACHIOPODS FROM SOUTH CHINA 277 In U n n I I 139 181 I U I I U I I $ 0 nil n ! n u n I I U I I~1 180 ~ -- 181 138 iI~ I~ ' ~.,~. ~-i]: ¢ 178b ~:i,.i.~i.~l:.~ ,:-'~ ;"~ ;"-'~i:i _.., A U- [ 17~" JO~ ~ I'll'II] ....IIl"'.i] 21II :In :II ~ i t37 174 i~_ 17z 173 175 136 172 ,° (~ 174 171 135 170 134 169 d) _ 168 133 167 165 164 132 163 162 131 161 3 rn general 160 ~ scale 159 o 158 130 0C 157 o¢ 1. lJ~ l in =. E E ~55 L~. 154 2_ 2. ~ 129 153 152 3 gN ¢.~ 151 0 128 150 149 4. 148 147 146 127 Downloaded By: [Deakin University] At: 04:24 11 May 2009 145 144 143 142 126 7. 141 125 8. 124 140 123 9. 10. F'~ Fig. 2. Stratigraphic column of the Changhsing (top 110 m only) and Feixianguan Formations at the Chuanmu section. 1, limestone; 2, argillaceous limestone; 3, cherty limestone; 4, cherty bioclastie limestone; 5, mudstone; 6, calcareous mudstone; 7, claystone; 8, Neochonetes (Zhongyingia) linshuiensis sp. nov.; 9, Neoehonetes (Huangichonetes) geniculatus sp. nov.; 10, other brachiopods. The arrow marked 'A' at top right indicates the Permian-Triassic biostratigraphic boundary, which occurs between Beds 178a and b, about 20 cm above the top of the Changxing Formation. The arrow marked 'B' indicates the lithostratigraphic boundary between the Changxing and Feixianguan Formations. 278 MONICA J. CAMPI and G. R. SHI ALCHERINGA been placed by us between Beds 178a and 178b, China (pers. obs.). The S-H-NPalaeocommunity which appear to be the equivalents of Beds 27a/ was associated with an abundant invertebrate fauna b and 27c/d of Meishan D section respectively dominated by crinoids, with bivalves, corals, (Peng et al. 2001,
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