Turk J Zool 2011; 35(3) 403-432 © TÜBİTAK Research Article doi:10.3906/zoo-0905-13

The studies made on Turkish Carabinae with checklist and bibliography (Coleoptera: Carabidae)

Sakine Serap AVGIN1,*, Pierfranco CAVAZZUTI2 1Osmaniye Korkut Ata University, Art and Science Faculty, Department of Biology, Osmaniye - TURKEY 2Via Chiesa 1, 12030 PAGNO (CN), ITALY

Received: 13.05.2009

Abstract: A synthesis of the studies carried out so far on Carabinae (Coleoptera: Carabidae) of Turkey is presented. Brief information about them is included here. Additionally, a species checklist and bibliography of Turkish Carabinae are given with important data. This study is intended to be a guide for future work on this subject.

Key words: Carabinae, Carabidae, Turkey

Türkiye Carabinae (Coleoptera: Carabidae) faunası üzerine yapılan tüm çalışmalar, fauna listesi ve biyografisi

Özet: Türkiye Carabinae (Coleoptera: Carabidae) faunası üzerine yapılan çalışmaların bir sentezi ve bu çalışmalar hakkında kısa bilgiler sunuldu. İlaveten, Türkiye Carabinae faunasının listesi ve biyografisi önemli bilgilerle birlikte verildi. Bu çalışma gelecek çalışmalar için bir rehber olacaktır.

Anahtar sözcükler: Carabinae, Carabidae, Türkiye

Introduction world (Deuve, 1991); or the nice Březina’s catalogue When studies on the species belonging to the (1999), who for the first time included all the type family Carabidae are examined, it is evident that most localities of species, subspecies, and synonyms; or a of these studies focus on the subfamily Carabinae, and synthesis of the genus Carabus in Europe (Turin et al., in particular on the genus Carabus. 2003); or an illustrated catalogue of the genus Carabus in the world (Deuve, 2004). For instance, we can cite some studies on the biology, morphology, taxonomy, and distribution of There are more studies on Carabinae subfamily of the species belonging to the subfamily Paussinae and Carabidae mainly for 2 reasons. The first reason is Carabinae of the Italian fauna (Casale et al., 1982); or that these species are of great importance for a catalogue and distribution of Carabus species in the biological control, and the second one is that most of

* E-mail: [email protected]

403 The studies made on Turkish Carabinae with checklist and bibliography (Coleoptera: Carabidae)

them are big in size, attractive in shape and colour, number is as many as 1200 today. In Turkey, the making them easier to examine. recorded number of Carabinae species is 87. When Most of the Carabinae species are predators. In the estimated new numbers of species belonging to Turkey, Calosoma sycophanta L. is used against Carabidae are taken into consideration in Turkey, Thaumatopea pityocampa (Denis & Schiffermüller) species belonging to Carabinae is 7 % of them. (Kanat, 2002). When the Carabidae family is examined in general Each study conducted on Carabinae is very in Turkey, it is seen that studies on Carabinae important for ecological and biogeographical aspects subfamily are more than the ones of the other and for biological control; therefore, any study carried subfamilies. out on this subject is a contribution to the knowledge The species belonging to the Carabinae subfamily regarding environment. have been classified in different ways by different Most of the studies conducted on Carabidae family researchers. in Turkey are about species belonging to the According to a recent classification by Löbl and Carabinae subfamily. The aim of this study is to Smetana (2003), the species and subspecies belonging provide general information about the studies carried to the Carabinae subfamily from Turkey are classified out on the Carabinae subfamily in Turkey, to in the genera Calosoma Weber, 1801, Carabus Linné, determine the investigations made on this subject, to 1758, and Cychrus Fabricius, 1794. present a checklist and bibliography of Turkish Among these genera, Carabus has the highest Carabinae, and to provide a source for the works to number of species and subspecies. There are many be conducted in the future. studies conducted on this genus in Turkey. Many of them have been conducted by entomologists from Materials and methods some European countries, such as Italy, France, All the studies on Turkish Carabinae, short notes, Austria, Germany, Hungary, Czech Republic, Great all the documents were compiled after a long and Britain, Russia, and Sweden. Some of these meticulous effort, and all the studies were reviewed entomologists are Silvano Battoni, Sandro Bruschi, and evaluated. Brief information about them is Achille Casale, Pierfranco Cavazzuti, Achille Costa, presented here. The first species that were described G. De Cristoforis, Stefano Dacatra, Alessandro Dalla from Turkey, the first records of some species from Torre, Piermauro Giachino, Giorgio Jan, Paolo Mazzi, Turkey, the review of some species, and all similar Gaetano Osculati, Beppe Osella, Maurizio Pavesi, Ivan studies, all the studies of the researchers who study on Rapuzzi, Mario Ruspoli, Arturo Schatzmayr, Enrico Turkish Carabinae, namely all the studies on Turkish Sturani (his father Mario described the larval Carabinae have been revised and this revision has morphology of some Anatolian species), Franco Tassi, been included in the “Remarks on the studies made Augusto Vigna Taglianti (Italy), Claude Auvray, on Turkish Carabinae” section of this paper. Gérard Charet, Philippe Basquin, Raymond Boulben, Additionally, “A Species Checklist and Bibliography Philippe Darge, Armand David, Charles Delagrange, of Turkish Carabinae” has been prepared meticulously Thierry Deuve, Guy Vacher de Lapouge, Bernard and it is submitted in the “A Checklist of Turkish Lassalle, Georges Ledoux, Jean-Philippe Legrand, Carabinae” and “A Bibliography of Turkish Carabinae” Patrice Machard, Daniel Prunier (France), Carl sections of this paper. Blumenthal, Eduard von Bodemeyer, Bodo von Bodemeyer, Carl Escherich, Henri Gadeau de Kerville, L. Ganglbauer, Friedman Göbel, Ernst Results Görgner, Walter Heinz, Adalbert Kindermann, Frank Remarks on the studies made on Turkish Kleinfeld, Max Korb, Armin Korell, Horst Korge, Carabinae Theobald J. Krueper, J. Lederer, Eberhard von In Turkey the number of species of Carabidae is Oertzen, Klaus Staven (Germany), Boleslav Březina, reported to be about 1100 (Casale and Vigna Martin Häckel, L.Hoberlandt, Josef Mařan, V.J. Taglianti, 1999). However, it is estimated that this Stanek, K. Taborsky (Czech Republic), Joahn Sahlberg

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(Finland), Stephan von Breuning, Rudolf Kenyeri, Lapouge (1914), and S. von Breuning (1928). Italian Karl Mandl, Arnold Penther, Harald Schweiger, Leo researcher Achille Costa carried out one important Weirather, Emerich Zederbauer (Austria), Igor research in the southwestern coast of the Anatolian Belousov, Maximilien de Chaudoir, G. Fischer vov Peninsula. Waldheim, C.G. Von Mannerheim, E. Ménétriés, V.I. Léon Fairmaire in 1884-1886 studied the Motschulski, Victor Shchurov, T. Tschitschrine, entomological materials collected in the Amanos Alexandr Zamotajlov (Russia). Mountains by abbot Armand David and Charles In addition, Avgın and Prunier (2007), Avgın and Delagrange, and published C. caramanus (1886). On Emre (2008), and Avgın and Prunier (2010) carried the same material S. von Breuning, in 1975, described out some studies on this subfamily in Turkey. C. scabrosus elbursensis (= elbursianus H. de Toulgoët,1986, new name). The first Carabus collected in Turkey was the bigger and one of the spectacular species of the world, In the last part of the 19th century E. Reitter and described as scabrosus Olivier, 1790. published the description of some new Carabus species: pirithous (1896), gordius, and omphreodes Subsequently, on the first part of the 1800, the (1898). Russian G. Fischer von Waldheim, Edouard Ménétriés, Victor I. Motschulski, and Maximilien de Viktor Apfelbeck, short time later (1904), in his Chaudoir explored the Caucasian region and its very important study on the “Käferfauna of the Balkan adjacent eastern Anatolians regions. In their peninsula” described C. graecus thessalonicensis and expedition, the following taxa were discovered: wiedemanni burgassiensis. Cychrus aeneus Fischer von Waldheim, 1824; The expedition carried out by Eduard von Callisthenes breviusculus orbiculatus (1839), Cychrus Bodemeyer on the Mount Bolkar, and by Austrian anatolicus Motschulski, 1865; Carabus gotschi, Arnold Penther and Emmerich Zederbauer on the puschkini kolenatii (1846), macrogonus (1847), victor Mount Erciyes, provided some important inconspicuus, bischoffi, nordmanni (1848), Carabidologic material. This material was studied by scabripennis, spinolae lamprus (1850), Carabus the top specialists of the time: L. Ganglbauer chalconatus chalcochlorus (1852), chevrolati thierki, published Carabus armeniacus korbianus, morawitzi anatolicus (1857), bonvouloiri (1863), microderus, (1887) and clathratus stygius (1890). E. Bodemeyer rumelicus (1867), acutangulus, Cychrus armeniacus described C. (Procerus) sommeri sterilis and (1879), Calosoma sicophanta severum, eversmanni bulgharmaadensis (1915). G. de Lapouge published C. Chaudoir (1850). torosus bodemeyeri (1914) and coriaceus cerisyi bodemeyeri (1923). At the same time, in 1832, Italian researchers Giuseppe De Cristoforis and Giorgio Jan visited Some naturalists in the beginning of 1900 carried Turkey and described: Carabus mariettii, torosus out entomological research. Finnish Joahn R. moestus, spinolae, chevrolati and saphyrinus De Sahlberg and his son, in 1904 visited the Central Cristoforis and Jan, 1837, while Gaetano Osculati in Anatolia, and published the result in his Catalogue of the expedition of the 1841-42 discovered Carabus the Coleoptera of the Eastern Mediterranean region maurus osculatii and cribratus orientalis Osculati, (Sahlberg, 1913). 1844. German Bodo von Bodemeyer published the Some of the most important naturalists of the result of his journey of 1911 in Turkey (Bodemeyer, Middle Europe visited Turkey; among others: 1927). French Henry Gadeau de Kerville, in 1912 Adalbert Kindermann (1838, 1850), Theobald J. went on an important journey in the Anatolian region Krueper (1875), Max Korbe (1886), Eberhard vov from İzmir to Ankara (Kerville, 1939). Oertzen (1887), and Carl Escherich (1896). The At that time, G. Vacher de Lapouge was a prolific materials they collected were studied by leading author of new taxa: Calosoma sicophanta prasinum scientists of the time, Maximilien de Chaudoir (1850), and Carabus punctatus vagepunctatus (1907), J. Waltl (1838), L. Ganglbauer (1887), Guy Vacher de scabriusculus bulgarus, wiedemanni bithynus, gotschi

405 The studies made on Turkish Carabinae with checklist and bibliography (Coleoptera: Carabidae)

unctus (1908), torosus escherichi, torosus bodemeyeri The most important contribution to the and mulsantianus akbesianus (1914), wiedemanni knowledge of the subgenus Oxycarabus Semenov, anceps and victor transeuns (1924). 1898, is the study carried out by the Austrian Karl V.J. Stanek in 1931 visited the Mount Ilgaz and Mandl dated back to 1971, in which he divided the discovered C. staneki, which was described as new subgenus into 4 species: saphyrinus Cristoforis & Jan, species by F. Štĕrba in the same year (1931). 1837, wagneri Küster, 1846, pirithous Reitter, 1896, and pseudopirithous Mandl, 1955. In 1975 he In 1932 Joseph Mařan published a study on completed a careful analysis of some species of Carabidae from the Ilgaz massif, from which he Sphodristocarabus of the group tokatensis Roeschke, described Cychrus armeniacus ilgazdaghensis among 1898, and macrogonus Chaudoir, 1847, but also of other new Carabidae species. blumenthaliensis Heinz & Korge, 1967, basilewskianus In 1947 the National Museum in Prague organized Breuning & Ruspoli, 1970, C. (Lamprostus) phaedimus a very important expedition of scientific research in Schaufuss, 1882, spinolae Cristoforis & Jan, 1837, and Turkey, which is considered as the first expedition of bonplandi Ménétriés, 1837. In a big study of the 1971 the modern time. The eminent entomologists taking he revised Sphodristocarabus belonging to the part were: L. Hoberlandt, K. Taborsky, J. Mařan, J. “macrogonus“ species group, giving a complete Dlabola, S. Jurecek, and others of the Museum of synthesis of the species: resplendens Breuning, 1934, Budapest (Hoberlandt and Taborsky, 1948; Mařan, macrogonus Chaudoir, 1847, simonianus Breuning & 1950). Ruspoli, 1970, scovitzi Faldermann, 1836a, and Other 3 important studies on the Turkish fauna pseudokolatensis Heinz & Korge, 1967. In that study, a followed the post-war period. At first, in 1947, Mařan great amount of data from historical bibliographic examined the distribution and the geographical sources and illustrations are indicated. In 1975, Mandl variation of the species C. (Procrustes) banoni Dejean, continued the revision of Sphodristocarabus analyzing 1829 (anatolicus Chaudoir, 1857 included) in the the “adamsi“ species group, distributed in the Aegean Islands and in the southern Anatolia. In the Caucasus, Transcaucasia, and North-East Turkey, and second study, Mařan, in 1949, made an important named many new taxa. In that work, the descriptions contribution to the knowledge of the geographical of all taxa are presented; moreover, many geographical distribution of the forms of C. (Procrustes) coriaceus indications are given, with a rich collection of Linné, 1758, present in Greece and Turkey, with the illustrations of aedeagus drawings, pronotum shapes, description of cerisyi bodemeyerianus. The and detailed distribution maps. A complete study of zoogeographical and phylogenetic conclusions of the Cychrus from Anatolia was edited in 1977. In the study are very interesting. taxonomic treatment proposed by Mandl, the genus In the last study, dated back to 1950, he showed the Cychrus is represented in Anatolia by 3 species: aeneus results obtained during the scientific expedition to Fischer von Waldheim, 1824, armeniacus Chaudoir, Turkey supported by the Museum of Natural Sciences 1879, and ruspolii Breuning, 1964. In 1980 he of Prague in 1947. The descriptions of C. published, with his typical meticulousness and (Lamprostus) torosus hoberlandti and C. (L.) hozari scrupulousness, the description of C. (Lamprostus) were presented in a larger and well detailed, critical hahni, in the context of a synthesis on the Lamprostus analysis of the species C. (L.) torosus Frivaldsky, 1835, of the western Anatolia. Accurate studies were and C. (Procrustes) chevrolati Cristoforis & Jan, 1837. conducted in 1986 attempting to define the identity Harald Schweiger (1962) identified some new of C. (Lamprostus) bonplandi Ménétriés, 1836, and of species of Carabus Linné, 1758, and also reported C. (L.) hozari Mařan, 1950. Finally, in 1987, Mandl some rare species from Turkey. This author (1964) performed a thorough study on the paleogeography, taxonomically examined 8 Carabus species from evolution, and the origin of some phyletic lineages of Turkey. In another study (1964), he examined the the most important Carabus subgenera in Turkey: species of Oxycarabus Semenov, 1898, and their Procrustes, Lamprostus, Chaetomelas, Oxycarabus, and zoogeographic relationships. Megodontus.

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The most important person who characterized the German Walter Heinz also started collecting carabidological research in Turkey in the post-war specimen in Turkey in the 1960s, and later on period was Austrian Stephan von Breuning, the published the descriptions of some new taxa. The author of the famous Monographie der Gattung most relevant ones were: C. (Megodontus) bonvouloiri Carabus (1932-1936). Since the 1960s, he published fortunatus, C. (Lamprostus) calley subnigrinus (1970), many contributions to the knowledge of the Turkish C. (Sphodristocarabus) scovitzi hedini, C. fauna. He described many new taxa, among which (Archicarabus) victor montanellus (1973), C. there were C. (Sphodristocarabus) heinzi and C. (Pachystus) ganymed (1975), C. (Procrustes) (Lamprostus) giresuni (1964). Breuning (1964) bernauerorum (1977), C. (Sphodristocarabus) hedini explained that the distribution of Procrustes chevrolati czipkai (1980). C. (Sphodristocarabus) enigmaticus was De Cristoforis & Jan, 1837 has close relationships with described together with its subspecies reiseri (1980), geological and climatic events in the area. In 1967 he and represents one of the most interesting species of published a noteworthy study on C. (Procrustes) the Turkish fauna. The collaboration with M. Pavesi chevrolati, and continued with systematic works, with (Heinz and Pavesi, 1994) originated an interesting the descriptions of many other Carabus and Cychrus study on Callisthenes from Turkey, with the species: C. (Tomocarabus) maljushencoensis, C. description of 2 new taxa: Callisthenes eversmanni (Megodontus) bonvouloiri kolatdaghensis, Cychrus pesarinii, n. ssp. and Callisthenes peksi, n. sp. ilgazdaghensis drannazdaghensis (1967). In 1972 he Since 1989 Heinz, together with Klaus Staven, has described C. (Sphodristocarabus) schuberti (a name published articles on the systematics of the Anatolian already assigned and replaced by Heinz, in 1975), and fauna. From their collaborations, a lot of excellent a long list of new taxa, many of which were published studies were derived, rich in biogeographic in the journal Entomops. Among them, there is one of annotations, containing the descriptions of many new the most beautiful gems from Turkey: C. taxa: C. (Lipaster) stjernvalli tenebrius (1989), C. (L.) (Titanocechenus) osellai (now included in Lipaster, gordius planostriatus (1995) and a well-argued study and downgraded to subspecies of gordius), for which on the subgenus Lipaster (1989). Papers on C. Ruspoli was the co-author. “Breuning and Ruspoli” (Sphodristocarabus) rotundicollis aequicostalis and C. were universally known in the carabidological (S.) tokatensis kirikensis (1995) followed. In 2000 they publications of that period: they carried out long published an important study on the distribution and scientific expeditions together in Anatolia. Sometimes nomenclature of Chaetomelas, a subgenus widely other people went with them, for instance Enrico spread in Asia Minor and in the eastern Sturani in 1970. Their collaboration allowed the Mediterranean region, and in the same year they discovery of many new taxa, among which there were wrote, in collaboration with B. Lassalle, a C. (Tomocarabus) rumelicus zaraensis (1970), C. (O.) development upon the identity of C. (Lipaster) cribratus haramburae (1974), C. (O.) cribratus gordius. schubertianus (1974), C. (Lipaster) stjernvalli latisi Armin Korell (1987, 1990) published some studies (1974), C. (S.) simonianus (1970), and others. on C. (Sphodristocarabus) tokatensis, and on the same Carl Blumenthal was one of the most assiduous co- species in Hatay (2001), from which he already operators in the studies of Breuning and Battoni. He described Procerus syriacus daphnensis (1985) and P. described, for instance, C.(Lamprostus) rabaroni syriacus limitaneus (1988). In 1998 he published a enricoi Battoni & Blumenthal, 1972, C. (L.) complete study on C. (Pachystus) trojanus, a very erenleriensis karadaghensis Battoni & Blumenthal, interesting species spread in the Aegean islands, and 1972, C. (Heterocarabus) mariettii sapancaensis in 1999, together with Ernst Görgner, he produced an Blumenthal & Breuning, 1966, C. (H.) mariettii interesting study on C. (Procrustes) impressus Klug, ormanensis Blumenthal & Breuning, 1966, C. 1832, of the province of Hatay, in which 2 new (Oxycarabus) pirithous sapancae Blumenthal & subspecies, halfeticus Korell and palmyrus Görgner, Breuning, 1966, and others. were described.

407 The studies made on Turkish Carabinae with checklist and bibliography (Coleoptera: Carabidae)

Achille Casale (1973) described C. which the diagnosis of C. (Sphodristocarabus) (Sphodristocarabus) resplendens cavazzutianus, and coruhnehriensis, a new species, and C. (S.) enigmaticus discussed some groups of Sphodristocarabus species, exiliatus (1990) are given. C. (Sphodristocarabus) giving photographs and drawings of their genitalia. giachinoi, C. (S.) karasudominus, and C. (S.) Pierfranco Cavazzuti, together with Casale, visited resplendens diogenei were published in 1991. C. Anatolia several times between 1973 and 1991. They (Tomocarabus) acutangulus mackaensis (1991) was described a new subspecies of Carabus from Turkey in described by Cavazzuti, together with Korell; with 1976: C. (Sphodristocarabus) resplendens lilianae, an him he also described C. (Tribax) merdeniki (1992), a interesting taxon with lively variable colors. In 1984 species with completely unknown habits, which only Cavazzuti published the descriptions of C. recently (2008) has been found again by Ivan Rapuzzi. (Sphodristocarabus) simonae, C. (S.) tokatensis In 1992 he published also a work with the description kosensis, and C. (Tribax) curlettii, a macrocephalous of 2 new species: C. (Pachystus) mazzii from South species rated among the most rare and mysterious Anatolia, and C. (Sphodristocarabus) pavesii, an from Anatolia. Its distribution is point form, localized interesting species coming, probably, from the in the area of Sarıkamış. C. (Sphodristocarabus) mountains located at the east of İzmir, but never georgia, another very rare and enigmatic species, found again. In some following studies he gave again coming from the surroundings of Tunceli, was descriptions of new Carabus from Turkey, among described in 1984. In the following years this author which C. (Lamprostus) pseudorobustus pseudocolossus, continued the analysis of the Turkish fauna publishing C. (L.) nordmanni exsultans (1993), and C. the description of many new taxa, as C. (Ischnocarabus) rubizzaniorum (1994). Moreover, in (Sphodristocarabus) macrogonus azaleae, C. (S.) collaboration with Rapuzzi (1998), he described C. armeniacus sarikamisensis and C. (Megodontus) (Pachystus) pisidicus samai, a new subspecies from bonvouloiri casalei, all in the same year (1986) in south Turkey, an unexpected discovery in a region which Procerus scabrosus ispiratus and P. s. callipygius very far away from the type locality of the species. In were described, as well. Procerus scabrosus 1999 he described C. (Lamprostus) spinolae leonidas, munzurensis (1987), a rare endemic from the Munzur a new unrecognized subspecies of C. spinolae that, massif, was described in collaboration with Lassalle. because of its isolated geographical localization In the same year (1987) he published a study on the among the southern populations of C. (L.) torosus, Lamprostus from the northern forests in which some raises many questions on the dynamics of the original new subspecies were described; among them the most population and on the systematic position of the 2 important is C. nordmanni triumphans with a very variable chromatic range, coming from inviolate species. There followed the description of Cychrus forests at low altitude of the Pontic Alps, near Tonya, anatolicus ziyathi (2003), a unique taxon of Cychrus from where also the earthworm Eophyla cavazzutii anatolicus present in northeastern Anatolia, isolated Omodeo, 1988 comes, one of the biggest Lumbricidae among numerous populations of C. aeneus. In 2004, (Oligochaeta) of the Mediterranean basin, which is in cooperation with Myska, he produced a study on almost surely a common prey of Lamprostus. some new subspecies of Procerus, from the Azerbaijan and the southern Armenia, and one from the In 1989, Cavazzuti presented a monographic paper Kahramanmaraş zone, in southern Turkey: C. (P. ) on Procerus Dejean, 1821, treated as a distinct genus. scabrosus kahramanicus. In this work many notes on systematics, biology, habitat, ecology, and distributions were given on all A more detailed study on Turkish Carabinae was species. This monograph treated all Procerus species prepared by Cavazzuti (2006). The distribution in from Anatolia and included descriptions of new Turkey, short definitions, and biologies of the species subspecies from this region. belonging to this subfamily were reported in this Later, Cavazzuti continued his investigations of the paper. Turkish fauna with a work on C. (Sphodristocarabus) In addition to these studies, some other studies species from the medium Valley of the Çoruh River, in were also published.

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Vigna Taglianti and Bruschi (1976) defined localized in the southeastern part of Anatolia, with Carabus (Sphodristocarabus) keltepensis as a new peculiar characteristics and, together with species for Turkey (in the mountains near Sapanca Relictocarabus from the high Moroccan Atlas, lake). They also discussed the systematic relations representing the remains of a relictual population of between the species of Sphodristocarabus Géhin, and the Mediterranean Basin. There followed the prepared a new identification key. description of C. (Apoplesius) legrandi (1991), Casale and Vigna Taglianti (1999) discussed the expanding to the Kars region, the range of the origin and biogeography of carabids of the Anatolian subgenus. In the following studies he described: C. fauna, presented a short history of its discovery, and (Sphodristocarabus) scovitzi sarigoelensis and C. gave a checklist of Anatolian Carabidae, with species (Lamprostus) spinolae ladikensis; he did the latter and subspecies of Carabinae included. together with Charet. In 1993 he proposed the division of C. (Cytilocarabus) cribratus in 3 different species: Silvano Battoni published 2 reports on his trips to cribratus Quensel, 1806, porrectangulus Géhin, 1885, the Balkans and Anatolia (1973, 1978), and described and staneki Sterba, 1931, based on important some taxa: C. (Procrustes) coriaceus mehmeti, C. differences in the morphology of the aedeagus. Then, (Sphodristocarabus) resplendens zanderi, and C. he described C. (Cytilocarabus) mahouxi ciloensis (Lamprostus) rabaroni enricoi (1973) in collaboration with Blumenthal, while he described by himself the (1994), C. (Heterocarabus) mariettii hroni (1997) and taxon C. (Oreocarabus) staneki yaraligozi (1982). C. (Sphodristocarabus) biroi iorbi (1998) with Charet. In 2001 he published an interesting systematic study Ivan Rapuzzi described (1990) C. (Ischnocarabus) on the Lamprostus of the robustus-nordmanni-torosus tenuitarsis prasinosomatus, a nice subspecies from the species group, with a great revision of its nomenclature, surroundings of Çankırı. Later on, he published some and later (2002) published a study on the Carabini and studies on Carabus from the province of Hatay, in Cychrini of the Caucasic and Ponto-Caucasic regions, which he described C. (Lamprostus) hemprichi in which he described some new subspecies of Cychrus propheta (1995), C. (Procrustes) impressus aeneus of the eastern Pontic Alps. Later, Deuve gave simulacrum, and C. (Chaetomelas) morawitzi some systematic studies with the descriptions of C. discipulus (1997). In 2000, together with Frank (Ischnocarabus) tenuitarsis balgoezensis (2004) and C. Kleinfeld he described C. (Tomocarabus) scabripennis (Sphodristocarabus) biroi leonardoi (2005) from the martaiellus, from Abant lake, and together with the surroundings of Sinop. More recently (2009), Deuve same author he published in 2004 a very complete and treated the distribution of C. (Minocarabus) maurus in detailed study on the Carabus and Procerus from the Anatolia and Cyprus, and described C. (M.) maurus Near East, including Hatay, in which they gave all the smyrnensis from İzmir. distribution localities, the maps of all taxa in those regions and a rich iconographic documentation. Bernard Lassalle discovered one of the most interesting and peculiar species from Turkey, Georges Ledoux published in 1972 the description described as C. (Tribax) deuvei (1986), that Deuve in of C. (Cytilocarabus) mahouxi followed by the 1990 attributed to the new subgenus Apoplesius, description of C. (Sphodristocarabus) resplendens created by himself. Some years later he published the rizensis, 1975. descriptions of some new taxa of Carabus: C. Thierry Deuve produced a great number of studies (Sphodristocarabus) rotundicollis aabadensis and C. on Carabids, some of which concern the study of the (Heterocarabus) mariettii mouthiezi (1990). In 1991 Turkish fauna. His first publication, together with he collaborated with Machard, and described many Simard, in 1977 was the description of C. new taxa, almost all coming from the eastern Pontic (Trachycarabus) ponticola. He then described C. Alps and belonging to the subgenera Tribax, (Tomocarabus) simardianus (1990), from specimens Megodontus and Sphodristocarabus, but also C. found in the Roeschke collection, about which there (Lipaster) osellai bontempsi, C. (L.) osellai lasoeli, and are still some doubts. In the same year he published an Callisthenes adehinotus. Later, he gave the descriptions excellent paper on the new subgenus Apoplesius, of C. (Oreocarabus) cribratus echarouxi and C. (O.)

409 The studies made on Turkish Carabinae with checklist and bibliography (Coleoptera: Carabidae)

cribratus bilmenekus (1996), C. (Chaetomelas) 1. Help to make an identification key for the morawitzi tainei and C. (Oxycarabus) saphyrinus Carabinae found in Turkey, yenicensis (1998). In 2004 he reported the findings of 2. Provide more detailed studies on the ecology of the type locality of C. (Lipaster) gordius, providing a Carabinae species, map of the distribution of Lipaster in Turkey and the description of C. (L.) stjernvalli ilgarensis. 3. Investigate Carabinae species for their functions as predator or prey. Patrice Machard published in 1977 a short synthesis on Lamprostus from Turkey, where he A Checklist of Turkish Carabinae described: C. (L.) rabaroni drouxi and C. (L.) calley The checklist of the Carabinae recorded from martinae. In 1979 he published a taxonomic revision Turkey is given below. of the Turkish taxa of the subgenera Oxycarabus, A total of 365 species and subspecies are included, Lipaster, Archicarabus and Tribax, and later on he distributed in 3 genera and 25 subgenera as follows: described some new taxa belonging to the subgenera Tribe CARABINI Latreille, 1802 Lamprostus, Procrustes, Sphodristocarabus and Tribax. In cooperation with Auvray (1990) he reported the GENUS CARABUS Linné, 1758: 413 presence in Turkey, in the region of Kars, of C. Div. SPINULATI Ishikawa, 1973: 46 (Procrustes) clypeatus abchasicus, a species new for the Subdiv. LEPIDOSPINULATI Imura, Kim, Su & Turkish fauna, previously known only in the Osawa 1998: 20 Caucasus. In 1994 he published a study on some new Carabus from north eastern Anatolia: C. (Lamprostus) Subgen. LIMNOCARABUS Géhin, 1876: 15 nordmanni subrobustus, C. (Archicarabus) gotschi clatratus Linné, 1761: 218 pseudorugosus, and C. (A.) victor subminimus (1995). ssp stygius Ganglbauer, 1890: 117 Claude Auvray published a pair of studies with Div. DIGITULATI Deuve, 1991a: 25 Machard (1990, 1994), already cited, and in 2005 a revision of the subgenus Tomocarabus in Turkey. Subgen. CARABUS Linné, 1758: 413 Belousov and Zamotajlov published the granulatus Linné, 1758: 413 astonishing discovery of the subgenus Cechenochilus ssp aetolicus Schaum, 1857: 122 in Turkey in 1999, with the description of the new Div. LIPASTROMORPHI Deuve, 1991a: 41 species lazorum. The subgenus, spread in the Caucasian region, was reported for the first time in Subgen. TRACHYCARABUS Géhin, 1876: 33 Turkey, on the Kaçkar massif, where many previous scabriusculus Olivier, 1795: 47 investigations were already carried out by several ssp bulgarus Lapouge 1908: 37 entomologists. This important discovery allows the possibility of finding other taxa of the same subgenus = interruptus Born 1902: 156 on the higher massifs of the eastern Pontic Alps. bosphoranus Fischer von Waldheim, 1823: 34 Finally, concerning the representatives of the genus = laceratus Csiki 1927: 273 Calosoma (in the widest sense) in Turkey, at least the ssp tscherkessicus Heinz & Korge, 1964: 236 classic monograph by Jeannel (1940), and the revision by Obydov (2002) of the subgenus Callisthenes = sibiriculus Mandl, 1955: 246 (treated as distinct genus), have to be cited. Subgen. MIMOCARABUS Gehin 1876: 36 In addition to all these studies, many short papers maurus Adams 1817: 281 and notes on Turkish Carabinae were issued by ssp maurus Adams, 1817: 281 several entomologists in different journals. = discoides Reitter, 1880: 457 Many more important studies should be = gemellus Fischer von Waldheim, 1844: 14 conducted on this subfamily in addition to the ones already made in Turkey. These studies should: ssp hochhuthi Chaudoir, 1846: 103

410 S. S. AVGIN, P. CAVAZZUTI

= maurulus Mandl, 1955: 247 wiedemanni Ménétries, 1836: 149 ssp alagoensis Kraatz, 1897a: 336 ssp burgassiensis Apfelbeck, 1904: 39 ssp osculatii Osculati, 1844: 72 = vaitoiani Strasser, 1900: 148 ssp geminatulus Reitter, 1896: 147 ssp wiedemanni Ménétries, 1836: 149 ssp paphius Redtenbacher, 1843: 981 ssp bythinus Lapouge, 1908: 19 ssp smyrnensis Deuve, 2009: 216 = cekirgensis Schweiger, 1964b: 160 pumilio Kuster, 1846: (Cah. n. 23) = nilufer Schweiger, 1964b: 160 ssp pumilio Kuster, 1846: (Cah. n. 23) = goekdaghensis Mandl, 1955: 252 = araratensis Kraatz, 1896: 336 ssp besucheti Schweiger, 1964b: 161 Subgen. LIPASTER Motschoulsky, 1866: 296 ssp akcakocaensis Blumenthal & Breuning, 1967: 56 = Lamprocarabus Thomson, 1875: 639 ssp winklerianus Breuning, 1932: 119 = Titanocechenus Breuning & Ruspoli, 1970: 215 ssp mengenicola Schweiger, 1969: 157 gordius Reitter 1898: 360 ssp mussardianus Breuning, 1964b: 213 ssp gordius Reitter, 1898: 360 = zindangensis Lassalle, 2003: 437 ssp bontempsi Lassalle, 1992b: 310 ssp paphlagoniensis Breuning, 1954: 134 ssp osellai Breuning & Ruspoli, 1970: 87 = patruelis Schweiger, 1964a: 144 = lasoeli Lassalle, 1992b: 311 ssp anceps Lapouge, 1924: 181 = pseudolangneri Machard, 1979: 290 victor Fischer von Waldheim, 1836: 350 ssp planostriatus Heinz & Staven, 1995: 15 = falsarius Reitter, 1896: 142 stjernvalli Mannerheim, 1830: 55 = motschoulskyi Kolenati, 1845: 31 ssp stjernvalli Mannerheim, 1830: 55 = nanus Kraatz, 1897b: 379 = boschniaki Faldermann, 1836a: 24 = dissolutus Csiki, 1927: 282 = raddeanus Schaufuss, 1882a: 526 = obtritus Lapouge, 1924: 181 = metzleri Mandl, 1955: 268 ssp transeuns Lapouge, 1924: 181 ssp posofensis Auvray, 1994: 137 = akkusanus Breuning, 1964b: 213 = purpureoviolaceus Machard, 1979: 290 = pseudotransenus Machard, 1979: 291 = ilgarensis Lassalle, 2004: 158 = inconspicuus Chaudoir, 1848b: 449 ssp latisi Breuning & Ruspoli, 1974: 238 ssp loebli Schweiger, 1969: 159 = langneri Breuning & Ruspoli, 1976: 222 = torulensis Lassalle, 2003: 434 ssp tenebrius Heinz & Staven, 1989: 49 ssp subminimus Machard, 1995: 274 Div. ARCHICARABOMORPHI Deuve, 1991a: 59 ssp cordithorax Blumenthal & Breuning, 1967: 56 Subgen. ARCHICARABUS Seidlitz, 1887: 6 = hortulanus Darge, 1994: 15 = Deuterocarabus Reitter, 1896: 141 ssp borckanus Cavazzuti, 2003: 10 montivagus Palliardi ,1825: 31 ssp montanellus Heinz & Korge, 1967: 91 ssp bulgaricus Csiki, 1927: 280 gotschi Chaudoir, 1846: 77 = ponticus Apfelbeck, 1904: 38 ssp inonunus Cavazzuti, 2003: 13

411 The studies made on Turkish Carabinae with checklist and bibliography (Coleoptera: Carabidae)

ssp ilgazicus Breuning & Ruspoli, 1970: 85 Div. METACARABI Bengtsson, 1927: 83 ssp regismontium Schweiger, 1964a: 143 Subdiv. MULTISTRIATI Reitter, 1896: 55 = subnivalis Schweiger ,1964a: 144 Latipalpi Reitter, 1896: 55 ssp congruus Motschoulsky, 1865: 288 Subgen. TOMOCARABUS Reitter, 1896: 135 = turcicus Motschoulsky, 1865: 288 decolor Fischer von Waldheim, 1823: 72 = ottomanus Reitter, 1896: 142 ssp decolor Fischer von Waldheim, 1823: 72 = koyulicus Lassalle, 2003: 438 = biseriatus Chaudoir ,1846: 87 = kochisarensis Lassalle, 2003: 440 convexus Fabricius, 1775: 238 ssp diversus Schweiger, 1969: 158 ssp gracilior Géhin, 1885: 49 ssp pseudogotschi Breuning, 1926: 75 ssp salax Motschoulsky, 1865: 286 ssp nicolae Auvray, 1995: 274 ssp immodicus Cavazzuti, 2006: 64 ssp rhinopterus Hampe, 1852: 309 ssp naumanni Breuning, 1971: 33 ssp sultandaghensis Breuning, 1966: 24 ssp horioni Korge, 1968: 37 ssp acutangulus Chaudoir, 1879: 109 ssp goeksunensis Breuning, 1966: 24 ssp mackaensis Cavazzuti & Korell, 1991: 99 ssp unctus Lapouge, 1908: 21 ssp erzeliki Schweiger, 1962a: 120 = bulghardaghensis Breuning, 1926: 78 = abantayaginiensis Schweiger, 1962a: 121 = demelti Heinz, 1977b: 12 scabripennis Chaudoir, 1850b: 156 = eregliensis Breuning, 1974b: 167 ssp scabripennis Chaudoir, 1850b: 156 ssp alayliensis Cavazzuti, 2006: 53 ssp ponticola Deuve & Simard, 1977: 1 ssp caramanus Fairmaire, 1886: xxi = reticulatus Hampe, 1852: 308 = mollis Lapouge, 1924: 183 ssp reductus Roeschke, 1896: 345 = depressipennis Mandl, 1955: 262 = nocturnus Schweiger, 1969: 160 ssp pseudorugosus Machard, 1994: 141 ssp subreductus Breuning, 1977: 48 Subgen. ISCHNOCARABUS Kraatz, 1877b: 78 ssp martaiellus Rapuzzi & Kleinfeld, 2000: 359 cychropalpus Peyron, 1858: 356 ssp kocaelicus Cavazzuti, 2006: 71 = cychropalpulus Mandl, 1955: 268 ssp microderus Chaudoir, 1867: 259 tenuitarsis Kraatz, 1877b: 78 = finikensis Lassalle, 1993: 304 ssp tenuitarsis Kraatz, 1877b: 78 ssp gautieri Géhin, 1885: 91 = cottarellii Breuning & Ruspoli, 1970: 96 = ledereri Gautier, 1868: 322 ssp balgoezensis Deuve & Charet, 2004: 36 simardianus Deuve, 1990a: 14 ssp ankarensis Breuning, 1961: 113 rumelicus Chaudoir, 1867: 259 = cankiriensis Breuning , 1962: 37 ssp rumelicus Chaudoir, 1867: 259 ssp prasinosomatus Rapuzzi, 1991: 36 = guelekensis Breuning, 1964b: 214 ssp auvrayi Charet, 1992b: 21 ssp zaraensis Breuning & Ruspoli, 1970: 85 ssp rubizzaniorum Cavazzuti,1994: 579 ssp cappadociae Breuning, 1932: 132

412 S. S. AVGIN, P. CAVAZZUTI

ssp kurdistanicus Eidam, 1931: 78 ssp porrectangulus Gehin, 1885: 5 = kurdistanicola Deuve, 1990a: 15 ssp ispiriacus Deuve, 1993a: 7 ssp syriensis Breuning, 1943: 14 ssp farineli Machard, 1975a: 52 Oreocaraboides Deuve, 1997b: 3 ssp tazieffi Breuning & Ruspoli, 1974: 89 Subgen. PACHYSTUS Motschoulsky, 1865: 294 = arpaalaniensis Machard, 1985: 27 = Melancarabus, Thomson, 1875: 640 staneki Sterba, 1931: 118 irmasanus Heinz & Korge, 1967: 92 ssp staneki Sterba, 1931: 118 ssp irmasanus Heinz & Korge, 1967: 92 = kreissli Schweiger, 1964a: 145 = felix Heinz & Korge, 1967: 92 ssp graziolii Cavazzuti, 1987: 502 ssp mazzii Cavazzuti, 1992: 115 ssp yaraligozi Battoni, 1982: 64 morio Mannerheim, 1830: 58 mahouxi Ledoux, 1972: 87 ssp morio Mannerheim, 1830: 58 ssp mahouxi Ledoux, 1972: 87 = olivieri Castelnau, 1835: 89 ssp ciloensis Deuve, 1994a: 17 = carceli Gory, 1833: 211 Subgen. PACHYCARABUS Géhin, 1876: 31 = acuminatus Ménétries, 1838: 10 = Caucasocarabus Lapouge, 1930: 241 = aethiops Cristoforis & Jan, 1837: 181 roseri Faldermann, 1836a: 22 ssp thessalonicensis Apfelbeck, 1904: 24 ssp roseri Faldermann, 1836a: 22 = cavernicola Kraatz, 1880: 157 = sphodrinus Fischer von Waldheim, 1844: 20 ssp muglanus Cavazzuti, 2006: 86 ssp ciftekopruleriensis Machard, 1975b: 142 ssp ganymed Heinz ,1975a: 145 ssp soganliensis Heinz & Korge,1964: 230 pisidicus Peyron, 1855: 669 Div. ARCİFERA Imura, 1996: 7 ssp pisidicus Peyron, 1855: 669 Subgen. HETEROCARABUS Morawitz, 1886: 49 ssp samai Cavazzuti & Rapuzzi, 1998: 398 akensis Haury, 1889: 106 tamsi Ménétries, 1832: 109 ssp akensis Haury, 1889: 106 Subgen. CYTILOCARABUS Reitter, 1896: 193 ssp akensiculus Cavazzuti, 2006: 108 cribratus Quensel, 1806: 171 ssp sapancaensis Blumenthal & Breuning, 1967: 56 ssp cribratus Quensel, 1806: 171 mariettii Cristoforis & Jan, 1837: 5 ssp orientalis Osculati, 1844: 72 ssp mariettii Cristoforis & Jan, 1837: 5 ssp schubertianus Breuning & Ruspoli, 1974: 229 ssp charetianus Auvray, 1992: 20 ssp negreianus Deuve, 1993b: 17 ssp ormanensis Blumenthal & Breuning, 1967, 57 ssp pulumuricus Cavazzuti, 1994: 580 ssp ativsengueni Schweiger, 1962a: 122 ssp haramburae Breuning & Ruspoli, 1974: 230 = pseudomarietti Schweiger, 1964b: 161 ssp bilmenekus Lassalle, 1996: 51 = petrovitzi Mandl, 1964: 2 ssp maironei Cavazzuti, 1994: 581 ssp necopinatus Basquin & Darge, 1987a: 406 ssp echarouxi Lassalle, 1996: 51 bischoffi Chaudoir, 1848b: 450 porrectangulus Gehin, 1885: 5 ssp bischoffi Chaudoir, 1848b: 450

413 The studies made on Turkish Carabinae with checklist and bibliography (Coleoptera: Carabidae)

ssp balboi Cavazzuti, 2005: 114 ssp simonianus Breuning & Ruspoli, 1970: 91 ssp mouthiezi Lassalle, 1991: 324 = pseudosimonianus Mandl, 1975a: 70 ssp ornatus Heinz, 1976: 66 ssp koselerensis Lassalle & Machard, 1991: 66 ssp muchei Breuning, 1961: 115 ssp azaleae Cavazzuti, 1986a: 3 ssp stephaniruspolii Breuning, 1966: 24 ssp giresunensis Deuve, 1992a: 55 = inclarus Darge, 1994: 15 ssp lemeslei Machard, 1995: 273 ssp terahci Charet, 1994: 55 ssp rizensis Ledoux, 1975: 28 ssp delmastroi Cavazzuti, 2003: 13 = michaelae Darge, 1994: 14 ssp inexpectatus Basquin & Darge, 1987a: 406 ssp pseudokolatensis Heinz & Korge, 1967: 97 ssp hroni Deuve, 1997a: 214 giachinoi Cavazzuti, 1991b: 192 Div. NEOCARABI Bengtsson, 1927: 83 biroi Csiki, 1927: 75 Subgen. SPHODRISTOCARABUS Géhin, 1885: 27 = biroi Csiki, 1927: 75 = Sphodristus Thomson, 1875: 640 = kindermanni Chaudoir, 1850b: 155 pavesii Cavazzuti, 1992: 118 ssp iorbi Deuve & Charet, 1999: 174. macrogonus Chaudoir, 1847: 8 ssp leonardoi Deuve, 2005: 267 ssp macrogonus Chaudoir, 1847: 8 heinzi Breuning,1964b: 215 = theophilei Deyrolle, 1869: 6 ssp heinzi Breuning, 1964b: 215 ssp keltepensis Vigna Taglianti & Bruschi, 1976: 209 ssp mateuellus Deuve, 2003: 264 = kolatensis Lapouge, 1925: 162 = matteui Breuning & Ruspoli, 1970: 93 ssp schweigerinae Schweiger, 1962a: 124 gilnickii Deyrolle, 1869: 6 = pseudoschweigerinae Schweiger, 1964a: 140 ssp hasandaghensis Darge & Basquin, 1990: 5 = goettingi Schweiger, 1964a: 139 ssp gilnickii Deyrolle, 1869: 6 ssp drannazdaghensis Breuning, 1964b: 214 = acutus Lapouge, 1904: 16 = clientulus Schweiger, 1969: 162 = giulielli Thomson, 1875: 686 = mollioni Breuning, 1976: 229 = viridifossulatipennis Breuning, 1964b: 214 ssp diogenei Cavazzuti, 1991b: 198 = brignolii Breuning & Ruspoli, 1970: 94 ssp resplendens Breuning, 1932: 135 = ovoides Machard, 1994: 142 = zanderi Battoni & Blumenthal, 1972: 52 = pseudoelongatus Machard, 1994: 142 = eiselti Mandl, 1975a: 67 = yakalamakus Lassalle, 1992b: 309 = Cavazzutianus Casale, 1973: 133 ssp lavezorum Darge, 1994: 14 = chambruni Breuning & Ruspoli, 1970: 92 ssp brunneicupreus Deuve, 1993b: 20 ssp akkusensis Breuning, 1964a: 106 coruhnehriensis Cavazzuti, 1991a: 106 = daumani Breuning, 1976: 232 enigmaticus Heinz, 1980: 80 = guergentepeensis Blumenthal, 1975: 59 ssp enigmaticus Heinz, 1980: 80 = ingloriosus Breuning & Ruspoli ,1970: 93 ssp reiseri Heinz, 1980: 82 ssp lilianae Cavazzuti & Casale,1976: 181 ssp citamgineus Lassalle & Charet, 1992a: 106

414 S. S. AVGIN, P. CAVAZZUTI

ssp bechteli Charet, 1996: 42 = basquini Darge, 1990b: 32 ssp exiliatus Cavazzuti, 1991a: 108 ssp machardianus Machard, 1986: 208 scovitzi Faldermann, 1836a: 20 ssp kurdaorum Lassalle, 1992b: 308 ssp scovitzi Faldermann, 1836a: 20 georgia Cavazzuti, 1984a: 496 ssp otigoensis Deuve, 1993b: 23 = hardei Heinz, 1990a: 162 ssp czipkai Heinz, 1980: 79 Subgen. CECHENOCHILUS Motschoulsky, 1850: 74 ssp bouilloni Breuning & Ruspoli, 1976: 238 = Cechenus Fischer von Waldheim, 1822: 110 ssp hedini Heinz, 1973: 26 lazorum Belousov & Zamotajlov, 1999: 67 ssp sarigoelensis Deuve & Charet, 1992b: 184 Subeng. APOPLESIUS Deuve, 1990b: 25 armeniacus Mannerheim, 1830: 59 deuvei Lassalle, 1986: 36 ssp armeniacus Mannerheim, 1830: 59 ssp deuvei Lassalle, 1986: 36 = subincatenatus Kraatz, 1878: 108 ssp hajdaji Hackel, 2007: 5 = mutabilis Géhin, 1885: 29 pseudodepressus Machard, 1988: 33 = fulminans Kraatz, 1878: 108 legrandi Deuve, 1991b: 20 = incatenatus Mannerheim, 1830: 60 omphreodes Reitter, 1897: 337 ssp korbianus Ganglbauer, 1887b: 136 Subgen. TRIBAX Fischer von Waldheim, 1817: 463 ssp pseudoarmeniacus Breuning, 1932: 136 = Microtribax Gottwald, 1982: 211 = arsiyandagensis Machard, 1985: 30 = Platychrus Kolenati, 1845: 25 = robichei Auvray, 1994: 144 =PlectesFischer von Waldheim, 1823: 52 ssp sarikamisensis Cavazzuti, 1986a: 4 puschkini Adams, 1817: 292 ssp rotundithorax Deuve, 2003: 265 ssp kolenatii Chaudoir, 1846: 97 = rotundicollis Mandl, 1955: 261 ssp kintrishensis Gottwald, 1980: 42 ssp basilewskianus Breuning & Ruspoli, 1970: 90 ssp ishikawaianus Breuning & Ruspoli, 1970: 94 ssp aabadensis Lassalle, 1990b: 324 = kabacaensis Machard, 1991: 21 ssp blumenthaliensis Heinz & Korge, 1967: 98 ssp zyzzyus Lassalle, 1992b: 306 ssp alagoensoides Mandl, 1955: 261 ssp pseudoponticus Lassalle & Machard, 1991: 59 tokatensis Roeschke, 1898: 162 = ispirensis Lassalle & Machard, 1991: 60 ssp tokatensis Roeschke, 1898: 162 = uskutdaghensis Lassalle & Machard, 1991: 60 = almushensis Mandl, 1975a: 62 ssp strobinoicus Breuning & Ruspoli, 1976: 224 ssp kosensis Cavazzuti, 1984b: 135 ssp zeytinlikensis Charet, 1996: 41 ssp kirikensis Staven, 1995a: 5 ssp ayrdensis Blumenthal & Breuning, 1966: 57 ssp simonae Cavazzuti, 1984b: 132 ssp ponticus Deyrolle, 1869: 6 ssp karasudominus Cavazzzuti, 1991b: 195 ssp praenuntius Heinz & Korge, 1967: 100 kurdicus Heinz, 1975b: 196 ssp mescitliensis Staven, 1998: 408 ssp kurdicus Heinz, 1975b: 196 merdeniki Cavazzuti & Korell, 1992: 61 = schuberti Breuning, 1972: 143 = koropeckyi Deuve & Macek, 1992: 1

415 The studies made on Turkish Carabinae with checklist and bibliography (Coleoptera: Carabidae)

curlettii Cavazzuti, 1984b: 138 ssp vexator Schweiger, 1969: 164 Subgen. LAMPROSTUS Motschoulsky, 1865: 297 ssp salignus Schweiger, 1969: 165 = Chaetogaster Lapouge, 1930: 197 ssp ladikensis Deuve & Charet, 1992: 125 = Chaetoprostus Reitter, 1896: 66 ssp lamprus Chaudoir, 1850b: 154 = Pseudoprocrustes Morawitz, 1886: 8 ssp dominici Thomé, 1987: 23 torosus Frivaldsky, 1835: 252 ssp rigouti Basquin & Darge, 1986a: 21 ssp torosus Frivaldsky, 1835: 252 ssp nordmanni Chaudoir, 1848b: 446 ssp escherichi Lapouge,1914: 35 = enricoi Battoni & Blumenthal, 1972: 52 ssp xanthicus Darge, 1992: 155 = productus Hampe, 1852: 309 ssp cankirianus Blumenthal & Breuning, 1966: 57 ssp rabaroni Breuning, 1964a: 107 = basquinianus Darge, 1992: 156 ssp sturanicus Breuning & Ruspoli, 1979: 281 = crudus Darge, 1992: 157 ssp giresuni Breuning, 1964a: 107 = reynieri Charet, 1992b: 21 ssp paululus Heinz & Korge, 1964: 233 ssp bodemeyeri Lapouge, 1914: 35 erenleriensis Schweiger, 1962a: 127 ssp jani Géhin, 1885: 8 ssp ahmetustaensis Cavazzuti, 1987: 492 = moestus Cristoforis & Jan, 1837: 1 ssp bithynicus Heinz & Korge, 1967: 102 = mundus Lapouge, 1914: 38 ssp erenleriensis Schweiger, 1962a: 127 = hoberlandti Mařan, 1952: 5 = kaynasliensis Machard, 1975b: 142 = vignatagliantii Breuning & Ruspolii, 1977: 68 ssp karadagensis Battoni & Blumenthal, 1972: 51 ssp gajaci Breuning & Ruspoli, 1979: 281 ssp fortepunctatus Battoni & Blumenthal, 1973: 105 = hahni Mandl, 1980: 152 = rotundicollis Blumenthal & Breuning, 1966: 57 ssp procrustoides Géhin, 1885: 7 ssp pseudotorosus Schweiger, 1969: 163 spinolae Cristoforis & Jan, 1837: 5 ssp hamus Basquin & Legrand, 1991: 68 ssp leonidas Cavazzuti, 1999: 406 robustus Deyrolle, 1869: 6 ssp pseudescherichi Breuning, 1962: 37 ssp robustus Deyrolle, 1869: 6 ssp spinolae Cristoforis & Jan, 1837: 5 = perfectus Schaufuss, 1882a: 517 = bonplandi Ménétries, 1836: 149 = callipeplus Schaufuss, 1882a: 517 = obesus Lapouge, 1909: 117 = semirugosus Schaufuss, 1882a: 517 = phaedimus Schaufuss, 1882a: 520 = sexpunctatus Schaufuss,1882a: 517 ssp mercatii Breuning & Ruspoli, 1970: 99 = trapesunticus Eichler, 1924: 61 ssp rouziei Breuning & Ruspoli, 1970: 97 = triumphator Schaufuss, 1882a: 517 ssp pseudoducalis Schweiger, 1964a: 146 = colossus Bleuse, 1896: 19 ssp usurpator Cavazzuti, 1987: 495 ssp pseudorobustus Heinz & Blumenthal, 1977: 219 ssp sinopensis Breuning, 1961: 113 = extremus Schaufuss, 1882a: 517 ssp coeruleoviolaceus Breuning & Ruspoli, 1970: 99 ssp yavuzkemalensis Deuve & Remond, 2001: 37 ssp lohsei Heinz & Blumenthal, 1978: 14 ssp egribelensis Deuve & Remond, 2001: 38

416 S. S. AVGIN, P. CAVAZZUTI

ssp drouxi Machard, 1977: 174 ssp aleppensis Lapouge, 1914: 43 ssp pseudocolossus Cavazzuti, 1993: 135 ssp bernhauerorum Heinz, 1977a: 35 ssp triumphans Cavazzuti, 1987: 498 punctatus Castelnau, 1834: 89 ssp exsultans Cavazzuti, 1993: 137 ssp punctatus Castelnau,1834: 89 ssp ducalis Schaufuss, 1882a: 514 ssp vagepunctatus Lapouge, 1907a: 148 = singanensis Breuning, 1932: 160 Subgen. PROCRUSTES Bonelli, 1809: 39 = subrobustus Machard, 1994: 140 = Macrogenus Motschoulsky, 1845: 346 ssp kaptanpasa Cavazzuti, 2001: 396 = Procrusticus White, 1845: 111 ssp athleta Semenov, 1897: 193 = Procrustocarabus Géhin, 1885: 4 chalconatus Mannerheim, 1830: 57 = Pseudoprocrustes Morawitz,1886: 8 ssp chalconatus Mannerheim, 1830: 57 = Sphodristus Motschoulsky, 1866: 295 = luxuriosus Motschoulsky, 1844: 10 payafa White, 1845: 111 = hozari Mařan, 1952: 6 ssp payafa White, 1845: 111 = hisarensis Darge, 1990d: 285 ssp pseudoacuticollis Catloš & Moravec, 1999: 110 = koyunensis Darge, 1990a: 57 ssp acuticollis Motschoulsky, 1858: 187 = maguini Auvray, 1994: 138 ssp drouxianus Machard, 1986: 206 ssp chalcochlorus Chaudoir, 1852: 97 clypeatus Adams, 1817: 299 = tschimenensis Breuning, 1964b: 215 ssp kurnakovi Kryzhanovskij, 1968: 174 ssp subnigritulus Heinz, 1990b: 349 = abchasicus Motschoulsky, 1850: 88 calleyi Fischer von Waldheim, 1824: 96 = fischeri Faldermann, 1836a: 14 ssp machardiellus Deuve ,1991a: 138 = rectoimpressus Motschoulsky, 1850: 88 = kurdistanicus Machard, 1986: 205 chevrolati Cristoforis & Jan, 1837: 2 ssp chaudoiriellus Deuve, 1995: 74 ssp internatus Heinz & Korge, 1964: 235 ssp martinae Machard, 1977: 175 = cifcandaghensis Blumenthal & Breuning, 1966: 59 = catakensis Machard, 1985: 28 = salignicola Schweiger, 1969: 166 ssp subnigrinus Heinz, 1970: 372 ssp ilgazdagensis Schweiger, 1964a: 147 ssp claudeauvrayi Cavazzuti, 2001: 395 = ilgazdaghensis Heinz & Korge, 1964: 234 propheta Rapuzzi, 1995: 473 = golkoyensis Schweiger, 1964b: 162 mulsantianus Morawitz, 1886: 7 = dadayanus Schweiger, 1964a: 142 ssp mulsantianus Morawitz, 1886: 7 = esmeraldae Breuning & Ruspoli, 1977: 69 = asperatus Mulsant & Wachanru, 1853: 124 = portei Breuning & Ruspoli, 1977: 70 = kotschyi Ganglbauer, 1887a: 2 = rodriguezi Breuning & Ruspoli, 1977: 69 = adspersarius Lapouge, 1907a: 148 ssp chevrolati Cristoforis & Jan, 1837: 2 ssp nurdagensis Blumenthal, 1967: 122 = assimilis Cristoforis & Jan, 1837: 3 ssp antakyae Korell, 1985: 141 = persimilis Csiki, 1927: 39 ssp akbesianus Lapouge, 1914: 45 ssp thierki Chaudoir, 1857a: 81

417 The studies made on Turkish Carabinae with checklist and bibliography (Coleoptera: Carabidae)

= wiedemanni Cristoforis & Jan, 1837: 3 Subgen. OXYCARABUS Semenov, 1898a: 507 = surejae Csiki, 1927: 39 pirithous Reitter, 1896: 66 ssp korbi Breuning, 1928: 1 ssp pirithous Reitter, 1896: 66 = chauveloti Breuning & Ruspoli, 1977: 70 ssp pseudopirithous Mandl, 1955: 269 ssp vanensis Machard, 1985: 30 ssp kenyeryi Schweiger, 1962a: 124 coriaceus Linné, 1758: 413 ssp sapancae Blumenthal & Breuning, 1967: 58 ssp kindermanni Waltl, 1838: 452 ssp yenicensis Lassalle, 1998: 91 = olympicus Géhin, 1885: 3 ssp saphyrinus Cristoforis & Jan, 1837: 4 = brussensis Ganglbauer, 1888: 395 ssp notabilis Roeschke, 1898: 163 = caraboides Ganglbauer, 1888: 394 ssp akdagicola Mandl, 1973: 80 = pseudofurax Mařan, 1952: 6 ssp pseudosaphyrinus Schweiger, 1965: 121 = mehmeti Battoni & Blumenthal, 1972: 51 = kainashensis Schweiger, 1965: 124 ssp cerisyi Dejean, 1826: 30 = boluensis Schweiger, 1965: 127 = melancholicus Gistel, 1857: 37 = ilgazdagicus Blumenthal & Breuning, 1967: 57 = minor Gautier, 1866b: 288 = mengenensis Blumenthal & Breuning, 1967: 58 = bodemeyeri Lapouge, 1923: 158 = korgei Heinz & Korge, 1964: 232 = bodemeyerianus Mařan, 1952: 6 ssp rabaronianus Breuning, 1964b: 216 ssp mopsucrenae Peyron, 1858: 357 ssp besanconi Breuning & Ruspoli, 1970: 99 ssp ressli Mandl, 1961: 30 ssp ruspolii Breuning, 1964b: 216 = montisgavurensis Mandl, 1975a: 78 ssp juengeri Breuning & Ruspoli, 1970: 101 anatolicus Chaudoir, 1857a: 81 ssp saphyrinus Cristoforis & Jan, 1837: 4 ssp anatolicus Chaudoir, 1857a: 81 = wagneri Kuster, 1846: (cah. 6, n. 12) = cypricus Roeschke, 1915: 294 = artusi Schweiger, 1965: 128 ssp lycicus Breuning, 1932: 163 = pseudowagneri Mandl, 1973: 78 = gueneycensis Catloš & Moravec, 1999: 112 Subgen. MEGODONTUS Solier, 1848: 58 ssp resslianus Mandl, 1963: 48 = Aulacocarabus Géhin, 1876: 24 impressus Klug, 1834: 71 = Nabicarabus Kwon & Lee, 1984: 102 ssp impressus Klug, 1834: 71 = Pachycranion Solier, 1848: 58 ssp simulacrum Rapuzzi, 1997: 564 =ProteocarabusGéhin, 1876: 5 = halfeticus Korell, 1999: 242 = Pseudocoptolabrus Reitter, 1896: 95 Subgen. CHAETOMELAS Thomson, 1875: 635 septemcarinatus Motschoulsky, 1840: 181 morawitzi Ganglbauer, 1887a: 2 = trigintiduosulcatus Lapouge, 1898: 60 ssp morawitzi Ganglbauer, 1887a: 2 bonvouloiri Chaudoir, 1863b: 447 = tainei Lassalle, 1998: 91 ssp gracilicollis Breuning, 1964a: 107 ssp montesamanus Mandl, 1967: 385 ssp bonvouloiri Chaudoir, 1863b: 447 ssp discipulus Rapuzzi, 1997: 565 = clandestinus Schaufuss, 1882b: 558

418 S. S. AVGIN, P. CAVAZZUTI

= nobilis Schaufuss, 1882b: 558 ssp munzurensis Cavazzuti & Lassalle, 1987: 71 = foveicollis Schaufuss, 1882b: 558 ssp montisabanti Schweiger, 1962a: 126 = pastillatus Lapouge, 1902: 310 caucasicus Adams, 1817: 282 = kolatdaghensis Breuning, 1967: 24 ssp armenius Zaitzev, 1930: 261 ssp kaschkarensis Blumenthal & Breuning, 1967: 59 ssp audouini Brullé, 1835: 116 = soganlidaghensis Lassalle & Machard, 1991: 64 ssp colchicus Motschoulsky, 1844: 7 ssp fortunatus Heinz, 1970: 370 = shawsheticus Zaitzev, 1930: 261 ssp casalei Cavazzuti, 1986a: 6 ssp ispiratus Cavazzuti, 1986b: 146 Subgen. PROCERUS Dejean, 1826: 22 bulgharmaadensis Bodemeyer E.,1915b: 571 scabrosus Olivier, 1790: 17 ssp toulgoetianus Deuve, 2002b: 281 ssp scabrosus Olivier, 1795: 17 = transversalis Csiki, 1927: 37 = olivieri Dejean, 1826: 24 = laticollis Kraatz, 1876: 127 = aegyptiacus Motschoulsky, 1844: 8 ssp bulgharmaadensis Bodemeyer E., 1915b: 571 = bosphoranus Motschoulsky, 1844: 4 ssp akbesiensis Toulgoët, 1986b: 499 = cribratus Motschoulsky, 1850: 87 = akbesianus Breuning, 1975b: 132 ssp propinquus Csiki, 1927: 36 ssp weidneri Lassalle, 1990a: 2 = dejeani Géhin, 1885: 1 syriacus Kollar 1843: 980 = angusticollis Haller, 1885: 200 ssp muruvvetae Avgın & Prunier, 2007: 83 = giganteus E. Bodemeyer, 1915a: 439 ssp syriacus Kollar, 1843: 980 ssp mentor Blumenthal & Breuning, 1967: 58 = daphnensis Korell, 1985: 140 ssp schuberti Breuning, 1968: 51 ssp limitaneus Korell, 1988b: 176 ssp culminicola Cavazzuti, 1989: 109 Gen. CALOSOMA Weber, 1801: 20 ssp kahramanicus Cavazzuti & Myska, 2004: 165 Subgen. CALOSOMA Weber, 1801: 20 sommeri Mannerheim, 1844: 868 sycophanta Linné, 1758: 414 ssp sommeri Mannerheim, 1844: 868 ssp sycophanta Linné, 1758: 414 = breviusculus Kraatz, 1876: 133 = nigrocyaneum Letzner, 1850: 95 = uludagensis Schweiger ,1964a: 140 =marginatumLetzner, 1850: 95 = basoglui Schweiger, 1964b: 163 = azureum Letzner, 1850: 95 ssp sterilis Bodemeyer E., 1915a: 439 = purpureoaureum Letzner, 1850: 95 ssp dardanellicus Kraatz Koschlau, 1886: 421 = cupreum Letzner, 1850: 95 ssp estegeicus Cavazzuti, 1989: 79 = smaragdinum Rossi, 1882: 204 ssp bureschianus Breuning, 1928b: 109 ssp prasinum Lapouge, 1907b: 124 ssp callipygius Cavazzuti, 1986b: 149 ssp severum Chaudoir, 1850c: 422 ssp cerkes Basquin & Darge, 1986b: 20 = rapax Motschoulsky, 1865: 309 ssp amasicus Csiki, 1927: 36 inquisitor Linné, 1758: 414 = modestus Kraatz, 1876: 133 ssp cupreum Dejean, 1826: 196

419 The studies made on Turkish Carabinae with checklist and bibliography (Coleoptera: Carabidae)

= clathratum Kolenati, 1845: 33 = mengenensis Breuning, 1975a: 48 = viridescens Reitter, 1896: 46 ssp ziyathi Cavazzuti, 2003: 14 Subgen. CAMPALITA Motschoulsky, 1865: 304 aeneus Fischer von Waldheim , 1824: 43 maderae Fabricius, 1775: 237 ssp aeneus Fischer von Waldheim, 1824: 43 auropunctatum Herbst, 1782: 131 = granulatus Motschulsky, 1850: 91 olivieri Dejean, 1831: 559 = signatus Faldermann, 1836a: 13 denticolle Gebler, 1833: 274 ssp ayderensis Deuve, 2002a: 122 Subgen. CALLISTHENES Fischer von Waldheim, ssp baiburtensis Breuning, 1975a: 48 1821: pl.7 ssp hopaensis Deuve, 2002a: 122 breviusculus Mannerheim, 1830: 61 ssp trabzonensis Deuve, 2002a: 120 = orbiculatus Motschulsky, 1839: 88 = araraticus Chaudoir, 1846: 104 Discussion = tamerlanum Zaitzev, 1918: 261 Many researchers from different countries of the eversmanni Chaudoir, 1850b: 157 world came to Turkey and collected samples of ssp eversmanni Chaudoir, 1850b: 157 Carabinae for many years up to the present and they conducted many research studies, such as the ssp persicus Géhin, 1885: 68 description of the new species that belong to this ssp adehinotus Lassalle, 1992b: 305 subfamily, their variety, distribution, and review. ssp pesarinii Heinz & Pavesi, 1994: 5 Furthermore, some researchers from Turkey have made some important investigations on this peksi Heinz & Pavesi, 1994: 6 subfamily. Combining all these studies and submitting Tribe CYCHRINI Laporte, 1834 them in one single source will surely be very essential Gen. CYCHRUS Fabricius, 1794: 440 for further studies to be performed in the future. All the studies on Carabinae from Turkey are submitted anatolicus Motschulsky, 1865: 310 in this publication from the first described species ssp anatolicus Motschulsky, 1865: 310 Carabus scabrosus Olivier, 1790 to the last described = medodjiensis Breuning, 1969: 36 subspecies Carabus (mimocarabus) maurus smyrnensis Deuve, 2009. ssp armeniacus Chaudoir, 1879: 109 Because the checklist and bibliography of this ssp occidentalis Mandl, 1977: 83 subfamily provide a basis for other studies, = armeniacus Roeschke, 1907: 253 submission of these in one single study is of great ssp drannazdaghensis Breuning, 1967: 24 importance for further studies. For example, the studies of Casale et al. (1982), Deuve (1991), Brezina ssp frivaldszkyi Roeschke, 1907: 250 (1999), Turin et al. (2003), Deuve (2004) are very ssp ilgazdaghensis Mařan, 1932: 30 significant studies that we always utilize regarding = angustemarginatus Mandl, 1977: 89 Carabinae. Like these papers, this study will certainly act as a basis for future studies in Turkey. = ruspolii Breuning, 1964b: 217 ssp korgei Breuning, 1964b: 217 Acknowledgement ssp ponticus Schweiger, 1962a: 119 The authors thank an anonymous reviewer who = breuningi Schweiger, 1964a: 141 highly improved the original version of this = akcakocae Breuning, 1969: 36 manuscript.

420 S. S. AVGIN, P. CAVAZZUTI

References Březina, B. 1999. World catalogue of the genus Carabus L. Sofia, Kanat, M. 2002. Çam Keseböceği (Thaumetopoea pityocampa Moscow: Pensoft Publ., 170 pp. Schiff.)’ne karşı biyolojik mücadelede Calosoma sycophanta L.’nın kullanımı. In: Proceedings of Pine Processionary Moth Casale, A., Sturani, M. and Vigna Taglianti, A. 1982. Fauna D’Italia, Symposium (Ed. M. Kanat), Kahramanmaraş- Turkey pp. 93- Coleoptera Carabidae. I Introduzione, Paussinae, Carabinae. 101. Calderini, Bologna, Italy, 499 pp. Turin, H., Penev, L. and Casale, A. 2003. The Genus Carabus in Deuve, T. 1991. La Nomenclature Taxonomique Du Genre Carabus. Europe. Pensoft Ed., 511 pp. Bull. Soc. Sciences Nat. Venette-France. 197 pp. Deuve, T. 2004. Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae). Pensoft Publishers. 440 pp.

A Bibliography of Turkish Carabinae

Adams, M.F. 1817. Descriptio Kahramanmaraş and surrounding towns, Battoni, S. and Blumenthal, C. 1972. insectorum novorum Imperii Russici, in South Central Turkey. Entomol. News. Expédition entomologique en Grèce et inprimis Caucasi et Siberiae. Mémoires de 119 (2): 151-170. Turquie. Entomops. 26: 51-58. la Société Impériale des Naturalistes de Avgın, S.S. and Prunier, D. 2010. Battoni, S. and Blumenthal, C. 1973. Moscou. 5: 278-314. Contribution to the taxonomy of the Beitrag zur Verbreitung und Systematik Apfelbeck, V. 1904. Die Käferfauna Genus Cychrus Fabricius, 1794 der Gattung Carabus in Nordanatolien der Balkanhalbininsel, mit (Coleoptera: Carabidae) from Turkey and und Griechenland (Col.). Mitt. Münch. Berücksichtigung Klein-Asien und der their relations with Cychrus species from Entom. Ges. 63: 93-111. Insel Kreta. I. Caraboidea. Friedlander, Balkans to Caucasus. Türk. Entomol. Beheim, D. and Breuning, S. Von. Berlin, ix+ 422 pp. Derg. 34(3): 307-322. 1943. Neubeschreibungen von Caraboidea u. Revisionen an den v. Auvray, C., Charet, G. and Machard, Basquin, P. and Darge, Ph. 1986a. Breuning’schen Monographien von P. 1991. Faune carabologique de Turquie. Contribution à la faune de Turquie: une Carabus, Calosoma und Ceroglossus Description de quatre races nouvelles. nouvelle sous-espèce de Lamprostus Bull. Sciences Nat. 74: 20-22. (Kol.). Mitteilungen d. München Ent. Ges. rabaroni Breuning (Coleoptera, 33 H. 25 pp. Auvray, C. and Machard, P. 1990. Un Carabidae). Bull. Sciences Nat. 49: 21-22. Carabe nouveau pour la faune de Turquie Belousov, I. and Zamotajlov, A. 1999. Basquin, P. and Darge, Ph. 1986b. The first record of the subgenus (Col.. Carabidae). L’Entomologiste. 46 (1): Deuxième contribution à la faune de Cechenochilus Motschulsky, 1850 for 23-25. Turquie, avec description d’un nouveau Turkey (Coleoptera, Carabidae genus Auvray, C. and Machard, P. 1994. Procerus (Coleoptera, Carabidae). Bull. Carabus L.). Advances in Carabidology. Faune carabologique de Turquie. Sciences Nat. 52: 20-21. 67-74. Description de sept races nouvelles. Basquin, P. and Darge, Ph. 1987a. Bengtsson, S. 1927. Die Larven der Remarques sur les sous-genres Deux nouveaux Heterocarabus d’Anatolie nordischen arten von Carabus Lin. Eine Lamprostus Motschulsky et Lipaster (Col. Carabidae). Nouv. Revue Ent. 4 (4): morphologische studie. Kungliga Motschulsky (Coleoptera: Carabidae). 406. Fysiografiska Sällskapets i Lund L’Entomologiste. 50 (2): 137-145. Förhandlingar (N.F.). 39 (2): 1-89. Basquin, P. and Darge, Ph. 1987b. Auvray, C. and Machard, P. 1995. Bleuse, L. 1896. Trois variétés Faune carabologique de Turquie. Note sur la synonymie de deux Procerus (Coleoptera, Carabidae). Bull. Sciences nouvelles de Coléoptères du Genre Description de quatre races nouvelles. Carabus L. Miscellanea Entomologica. IV Nat. 56: 21. Résultats d’élevages (Coleoptera: (2):18-19. Carabidae). L’Entomologiste. 51 (6): 273- Basquin, P. and Legrand, J.P. 1991. Blumenthal, C. 1967. Carabus 278. Diagnose d’un Carabus nouveau (Procrustes) mulsantianus Mor.n. Avgın, S.S. and Prunier, D. 2007. d’Anatolie. (Col. Carabidae). Bull. Soc. nurdagensis nov. Entomol. Blätter. 68 (2): Description d’une nouvelle sous-espèce Entom. de Fr. 96:68. 121-122. de Procerus de Turquie méridionale. Le Battoni, S. 1982. Nouvelle sous-espèce Blumenthal, C. 1975. Beschreibung Coléoptériste. 10 (2): 83-86. de Carabus staneki Sterba (Col. einer neuen Sphodristocarabus rasse aus Avgın, S.S. and Emre, İ. 2008. Some Carabidae). Bull. Soc. Ent. Mulhouse. 63- Anatolien (Carabidae, Col.). Kol. Carabinae (Coleoptera: Carabidae) from 64. Rundsch. 52: 59-60.

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Blumenthal, C. 1976. Beitrag zur Breuning, S. 1928b. Beitrag zur Breuning, S. 1971. Beschreibung einer verbreitung und systematik der Gattung carabenfauna von Bulgarien (Carabidae. neuen rasse der gattung Carabus L. Carabus in Griechenland und der Turkei Col.) mit Berücksichtigung den (Coleopt. Carabidae). Kol. Rundsch. 49: 3. (Col. Carabidae). Nouv. Rev. Ent. 6 (2): angrenzenden Gebiete. Mitteilungen des Breuning, S. 1972. Carabus 113-120. Naturwissenschaftlichen Institutes Sofia. (Sphodristocarabus) schuberti n.sp. 1: 108-117. Blumenthal, C. and Breuning, S. 1967. Zeitsch. Arbeit. Ost. Ent. 22: 143. Beitrag zur kenntnis der Carabus Fauna Breuning, S. V. 1932. Monographie Breuning, S. 1974a. Quelques Anatoliens (Coleoptera, Carabidae). Boll. der Gattung Carabus (1932-1936), Best. remarques concernant l’article intitulé Ass. Romana Ent. 21: 56-59. Tab. Europ. Coleopt., 104-110, Reitter, “Beitrag zur kenntnis der fauna des Iran Troppau. 41: 1610 pp. Blumenthal, C. and Deuve, T. 1986. und Anatoliens”, par Walter Heinz Carabus (Chaetomelas) praestigiator Breuning, S. 1950. Einige neue Arten (Entomol. Blätter, 69: 14-36, 1973) (Col. pinguis Lapouge, n. comb. Désignation und Rassen der Gattungen Carabus und Carabidae, Carabinae). Nouv. Rev. Ent. 4 d’un néotype (Col. Carabidae). Nouv. Cychrus aus Ostasien. Ent. Arb. Mus. G. (2): 115-117. Revue Ent. 3 (4): 438 pp. Frey. 1: 198-201. Breuning, S. 1974b. Eine neue Blumenthal, C. and Heinz, W. 1982. Breuning, S. 1954. Weiterer Beitrag Carabus Art aus Anatolien. Mitt. Zool. Bemerkungen zu einer Arbeit von K. zur kenntnis asiatischen Carabini. Ent. Mus. Berlin, L: 167. Mandl mit dem Titel “Eine neue Art der Arb. Munich. 5: 134-135. Breuning, S. 1975a. Description de Untergattung Lamprostus Motschulsky Breuning, S. 1961. Neuer Beitrag zur Cychrus nouveaux d’Anatolie (Col. der Gattung Carabus L. und kritische kenntnis der Gattung Carabus L. Entom. Carabidae). Bull. Soc. Ent. Mulhouse. Bemerkungen zu einer Publikation über Abh. Ber. Mus. Tierk. Dresden. 26 (13): 1975:48. diese Untergattung aus dem Jahre 1978 113-115. Breuning, S. 1975b. Description de (Col. Carabidae)”. Entomologica Breuning, S. 1962. Neue und nouvelles sous-espèces du genre Carabus Basiliensa. 7: 217-225. interessante Formen der Gattungen L. (s.l.) (Coléoptères Carabidae Bodemeyer, B. 1927. Über meine Carabus L. und Dorcadion Muls. aus Carabinae). Nouv. Rev. d’Ent. 5: 129-134. entomologischen Reisen. I. Kleinasien. Anatolien. Reichenbachia. I (6): 37-40. Breuning, S. and Ruspoli, M. 1970. Stuttgart, 85 pp. Breuning, S. 1964a. Beschreibung Description de 15 espèces et sous-espèces Bodemeyer, E. 1900. Quer durch einiger neuer Formen der Gattung nouvelles de Carabidae (Col.) d’Anatolie Klein-Asien in der BulgharDağh. Eine Carabus L. Ent. 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