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A Checklist and Keys to North European Genera and Species of Dolichopodidae (Diptera)

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A Checklist and Keys to North European Genera and Species of Dolichopodidae (Diptera) All-Russian Institute of Plant Protection RAAS Справочный список и определитель родов и видов ISSN 1815-3682 хищных мух Dolichopodidae (Diptera) Северной Европы. ÂÅÑÒÍÈÊ Гричанов И.Я. Санкт-Петербург: ВИЗР РАСХН, 2006. ÇÀÙÈÒÛ ÐÀÑÒÅÍÈÉ 120c. (Приложение к журналу «Вестник защиты Приложение растений»). A checklist and keys to North European genera and species of Dolichopodidae (Diptera). Igor Ya. Grichanov. St.Petersburg: VIZR RAAS, 2006. 120p. (Plant Protection News, Supplement). Supplement Составлен справочный список и определитель 38 родов и 393 видов хищных мух Dolichopodidae (Diptera), известных в Северной Европе (Да- A checklist and keys to ния, Исландия, Латвия, Литва, Норвегия, Финляндия, Швеция, Эстония, Карелия, Республика Коми, Ненецкий автономный округ, Архангельская, North European genera Ленинградская, Мурманская, Новгородская, Псковская области). Для каж- дого вида даны оригинальные родовые комбинации, основные синонимы, глобальное распространение. В водном разделе приведены сведения о сис- and species of тематическом положении и морфологии имаго мух-зеленушек. Работа бу- дет полезна специалистам – энтомологам и экологам, интересующихся эн- Dolichopodidae (Diptera) томофагами, студентам и аспирантам учебных и научных учреждений. Рецензент: канд. биол. наук И.В. Шамшев Igor Ya. GRICHANOV Работа выполнялась в рамках международных проектов Минпромнауки Российской Федерации «Биорегуляция агроэкосистем» (2001-2003) и «Биотехнологии в защите растений» (2004-2005), была также поддержана фондами Swedish Institute (2001-2004) и Finnish Academy of Sciences (2003). Рекомендовано к печати редакционной коллегией Всероссийского научно-исследовательского института защиты растений РАСХН 15 июня 2006г. Copyright 2006© I.Ya. Grichanov ISSN 1815-3682 St.Petersburg 2006 2 Contents Introduction................................................................................................3 Systematic position of Dolichopodidae.......................................................3 Morphology of Dolichopodidae..................................................................5 Basic references..........................................................................................8 Checklist of North European Dolichopodidae (Diptera)..............................9 Key to North European genera of Dolichopodidae.....................................58 A checklist and keys to Keys to North European species of Dolichopodidae...................................67 Achalcus...............................................................................................67 North European genera and species of Argyra..................................................................................................68 Dolichopodidae (Diptera) Campsicnemus......................................................................................70 Chrysotimus..........................................................................................72 Chrysotus .............................................................................................72 Igor Ya. GRICHANOV Diaphorus.............................................................................................74 Dolichopus ...........................................................................................75 Hercostomus.........................................................................................90 Hydrophorus.........................................................................................95 Lamprochromus....................................................................................98 All-Russian Institute of Plant Protection, Medetera ..............................................................................................98 Podbelskogo 3, St.Petersburg-Pushkin, 196608, Russia. Micromorphus ....................................................................................102 Email: [email protected] Neurigona...........................................................................................103 Peodes................................................................................................103 Rhaphium ...........................................................................................104 Scellus ................................................................................................107 Abstract. Sciapus...............................................................................................107 Sybistroma..........................................................................................110 A check list and keys to North European genera and species of Sympycnus..........................................................................................110 Dolichopodidae are compiled. 393 species and 38 genera known in Syntormon ..........................................................................................112 northern part of Europe are included, belonging to the nine sub- Systenus..............................................................................................113 families. Introductionary notes concerning systematic position and Tachytrechus ......................................................................................113 morphology of Dolichopodidae and new species records for some Teuchophorus.....................................................................................115 regions of Russia are given. Thinophilus.........................................................................................115 Thrypticus...........................................................................................115 Key words: Diptera, Dolichopodidae, catalogue, keys, North Xanthochlorus ....................................................................................117 Europe. Acknowledgements.................................................................................118 References...............................................................................................118 3 4 Introduction – Empodium bristle-like or not discernible, at most at most 2 well-developed pads below claws........................................................................................4 The Dolichopodidae fauna of the world is very large, with approximately 4. Basal cells of wing long; vein CuA2 long, reaching wing margin near A1, or 7000 described species and 240 genera (Grichanov, 2003-2006). These mostly joining A1 close to wing margin (at less than a quarter of its length back predatory flies are distributed throughout the world including the tropics and from wing margin) ....................................................................................... high-latitude islands and territories. In North Europe adults and larvae of almost ................Asilidae, Syrphidae, Bombyliidae, Therevidae, Pipunculidae et al. all species of long-legged flies are predators inhabiting moist substrata. Species – Second basal cell short, or even confluent with discal cell; anal cell short and of only one genus (Thrypticus) are known to be phytophages living inside stems closed or absent; vein CuA2 absent, reduced, or joining A1 far from wing of cereal grasses. Most of the numerous species of the cosmopolitan genus margin........................................................................................................5 Medetera are associated with tree trunks, especially in boreal forests, where 5. Wing venation without any crossvein in median region of wing .................... their larvae are predacious mainly on bark-beetles (Coleoptera). .............................................................Lonchopteridae, Opetiidae, Phoridae North European Dolichopodidae have never been generalised in one book. – Venation with at least one crossvein in median region of wing......................6 Now 393 species and 38 genera are known in North Europe, belonging to the 6. Wing with both A1 and Sc reaching wing margin, and with posterior cubital following nine subfamilies: Achalcinae, Diaphorinae, Dolichopodinae, Hydro- cell (cup) acute at posterior apex; hind tarsus, at least in male, with one or phorinae, Medeterinae, Neurigoninae, Rhaphiinae, Sciapodinae and Sympycni- more basal segments expanded and flattened; arista (stylus) 3-segmented, nae. However, Achalcinae, Neurigoninae, Rhaphiinae and Sciapodinae each terminally situated.....................................................................Platypezidae contain only one genus known from North Europe, whereas definition of the – If A1 reaching wing margin, then either Sc incomplete or cup obtuse or other subfamilies is far from an ideal state. A number of new subfamilies and rounded at posterior apex, or cup very small; basal segments of hind tarsus tribes have been proposed in the second half of the XX century that means that rarely expanded or flattened........................................................................7 a revision of the family systematics on the global scale is needed. So, we do not 7. Vein R4+5 forked; if not, then either prosternum large (fused with episterna) think that a key to subfamilies is necessary to give in this work. and metapleura usually bristled, or costa running around the wing. Antennal scape bristled, at least with a few bristly hairs beneath; male hypopygium SYSTEMATIC POSITION OF DOLICHOPODIDAE symmetrical and unrotated............................................................Empididae – Vein R not forked; prosternum in the form of a small sclerite separated by The family Dolichopodidae
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