English Holly Ilex Aquifolium L
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Toward a Resolution of Campanulid Phylogeny, with Special Reference to the Placement of Dipsacales
TAXON 57 (1) • February 2008: 53–65 Winkworth & al. • Campanulid phylogeny MOLECULAR PHYLOGENETICS Toward a resolution of Campanulid phylogeny, with special reference to the placement of Dipsacales Richard C. Winkworth1,2, Johannes Lundberg3 & Michael J. Donoghue4 1 Departamento de Botânica, Instituto de Biociências, Universidade de São Paulo, Caixa Postal 11461–CEP 05422-970, São Paulo, SP, Brazil. [email protected] (author for correspondence) 2 Current address: School of Biology, Chemistry, and Environmental Sciences, University of the South Pacific, Private Bag, Laucala Campus, Suva, Fiji 3 Department of Phanerogamic Botany, The Swedish Museum of Natural History, Box 50007, 104 05 Stockholm, Sweden 4 Department of Ecology & Evolutionary Biology and Peabody Museum of Natural History, Yale University, P.O. Box 208106, New Haven, Connecticut 06520-8106, U.S.A. Broad-scale phylogenetic analyses of the angiosperms and of the Asteridae have failed to confidently resolve relationships among the major lineages of the campanulid Asteridae (i.e., the euasterid II of APG II, 2003). To address this problem we assembled presently available sequences for a core set of 50 taxa, representing the diver- sity of the four largest lineages (Apiales, Aquifoliales, Asterales, Dipsacales) as well as the smaller “unplaced” groups (e.g., Bruniaceae, Paracryphiaceae, Columelliaceae). We constructed four data matrices for phylogenetic analysis: a chloroplast coding matrix (atpB, matK, ndhF, rbcL), a chloroplast non-coding matrix (rps16 intron, trnT-F region, trnV-atpE IGS), a combined chloroplast dataset (all seven chloroplast regions), and a combined genome matrix (seven chloroplast regions plus 18S and 26S rDNA). Bayesian analyses of these datasets using mixed substitution models produced often well-resolved and supported trees. -
Department of Planning and Zoning
Department of Planning and Zoning Subject: Howard County Landscape Manual Updates: Recommended Street Tree List (Appendix B) and Recommended Plant List (Appendix C) - Effective July 1, 2010 To: DLD Review Staff Homebuilders Committee From: Kent Sheubrooks, Acting Chief Division of Land Development Date: July 1, 2010 Purpose: The purpose of this policy memorandum is to update the Recommended Plant Lists presently contained in the Landscape Manual. The plant lists were created for the first edition of the Manual in 1993 before information was available about invasive qualities of certain recommended plants contained in those lists (Norway Maple, Bradford Pear, etc.). Additionally, diseases and pests have made some other plants undesirable (Ash, Austrian Pine, etc.). The Howard County General Plan 2000 and subsequent environmental and community planning publications such as the Route 1 and Route 40 Manuals and the Green Neighborhood Design Guidelines have promoted the desirability of using native plants in landscape plantings. Therefore, this policy seeks to update the Recommended Plant Lists by identifying invasive plant species and disease or pest ridden plants for their removal and prohibition from further planting in Howard County and to add other available native plants which have desirable characteristics for street tree or general landscape use for inclusion on the Recommended Plant Lists. Please note that a comprehensive review of the street tree and landscape tree lists were conducted for the purpose of this update, however, only -
Hollies for the Landscape
HOLLIES FOR THE LANDSCAPE Ilex crenata ‘Beehive’ Japanese Holly A Rutgers NJAES introduction - This is a northern hardy, Japanese Holly with a compact globe habit and shiny green foliage. Ilex ‘Winter Bounty’ A Rutgers NJAES introduction – ‘Winter Bounty' (I. ciliospinosa x. I. latifolia) is a unique holly with bold, shiny, long, nearly smooth leaves, rounded evergreen foliage. Heavy berry display, pollinate with 'Blue Prince', reportedly deer resistant. Ilex crenata ‘Jersey Pinnacle’ Japanese Holly A Rutgers NJAES introduction – this holly has a dense upright tight growing form with glossy dark green leaves. Ilex rugosa × I. (integra×pernyi) Rutgers ‘Spartan’ Holly U.S. Plant Patent #20,804 A Rutgers NJAES introduction – ‘Spartan’ is a new and distinct variety of evergreen holly distinguished in that it combines from three species (Ilex rugosa, Ilex integra, and Ilex pernyi), and has desirable landscape and production traits which distinguish it from all other forms of Ilex. In a landscape setting, the growth habit can be described as dense, self-compacting, and conical. Spartan has a unique combination of characteristics: nearly conical shape with minimal pruning; moderately vigorous, dense and self-compacting; easy to maintain at desired height; leaves are consumer friendly (no spines); abundant and well-distributed fruit; and low susceptibility to disease and insect pests. The plants can be used as foundation plants, hedge plants, and as single specimen plants since they are easy to maintain at a desired height. Ilex x ‘Rutzan’ ‘Red Beauty’ Holly U.S. Plant Patent No. 14,750 A Rutgers NJAES introduction - Red Beauty® is a new and distinct variety of evergreen holly distinguished in that it combines from three species (Ilex aquifolium, Ilex rugosa, and Ilex pernyi), and has desirable landscape and production traits which distinguish it from all other forms of Ilex. -
State of New York City's Plants 2018
STATE OF NEW YORK CITY’S PLANTS 2018 Daniel Atha & Brian Boom © 2018 The New York Botanical Garden All rights reserved ISBN 978-0-89327-955-4 Center for Conservation Strategy The New York Botanical Garden 2900 Southern Boulevard Bronx, NY 10458 All photos NYBG staff Citation: Atha, D. and B. Boom. 2018. State of New York City’s Plants 2018. Center for Conservation Strategy. The New York Botanical Garden, Bronx, NY. 132 pp. STATE OF NEW YORK CITY’S PLANTS 2018 4 EXECUTIVE SUMMARY 6 INTRODUCTION 10 DOCUMENTING THE CITY’S PLANTS 10 The Flora of New York City 11 Rare Species 14 Focus on Specific Area 16 Botanical Spectacle: Summer Snow 18 CITIZEN SCIENCE 20 THREATS TO THE CITY’S PLANTS 24 NEW YORK STATE PROHIBITED AND REGULATED INVASIVE SPECIES FOUND IN NEW YORK CITY 26 LOOKING AHEAD 27 CONTRIBUTORS AND ACKNOWLEGMENTS 30 LITERATURE CITED 31 APPENDIX Checklist of the Spontaneous Vascular Plants of New York City 32 Ferns and Fern Allies 35 Gymnosperms 36 Nymphaeales and Magnoliids 37 Monocots 67 Dicots 3 EXECUTIVE SUMMARY This report, State of New York City’s Plants 2018, is the first rankings of rare, threatened, endangered, and extinct species of what is envisioned by the Center for Conservation Strategy known from New York City, and based on this compilation of The New York Botanical Garden as annual updates thirteen percent of the City’s flora is imperiled or extinct in New summarizing the status of the spontaneous plant species of the York City. five boroughs of New York City. This year’s report deals with the City’s vascular plants (ferns and fern allies, gymnosperms, We have begun the process of assessing conservation status and flowering plants), but in the future it is planned to phase in at the local level for all species. -
Eryngium Yuccifolium A. Michaux Rattlesnake Master (Eryngium Synchaetum)
A. Michaux Eryngium yuccifolium Rattlesnake Master (Eryngium synchaetum) Other Common Names: Button Eryngo, Button Snakeroot. Family: Apiaceae (Umbelliferae). Cold Hardiness: With proper provenances, this species grows in USDA hardiness zones 4 to 9. Foliage: Alternate, simple, yucca-like sword-shapted blue-green foliage clasps stout stems; basal leaves to 30 long, but leaves on flower stalks much shorter; strap-like ½ to 1½ wide, margins toothed on terminal portions becoming spiny at the base; the specific epithet refers to the yucca-like foliage. Flower: Tiny individual fragrant flowers in ¾diameter ball-like clusters in open flattened clusters atop tall flower stalks in late spring to summer; clusters subtended by holly or thistle-leaf like bracts; individual flowers are numerous and tightly packed; greenish white to white flowers have five-petals and two filiform styles. Fruit: Seed heads eventually turn brown and are retained on the plant into winter until stems die back. Stem / Bark: Stems — stout, stiffly erect, somewhat swollen at the nodes; glabrous, green to bluish green; Buds — small; green to blue-green; Bark — not applicable; basal leaves and floral stalks from semi-woody base. Habit: Erect, 3 to 4 (6) tall, sparsely branched herbaceous perennials from a woody base, with the vegetative tissues sort of reminiscent of a cross between an Iris and a Yucca; over time a cluster of foliage forms at the base; the plant's texture is attractively coarse. Cultural Requirements: Sunny sites with moist well drained soils are required; drainage is particularly important as plants are grown in mesic locations, less so in more arid regions; overly fertile soils result in lodging and plants benefit from being surrounded by shorter plants that can lend support to the tall flower stalks; transplant from containers or seed in place as taproots hinder successful transplant; prickly leaves may hinder maintenance activities around the plants. -
Ilex Crenata and Its Varieties
ARNOLDIA A continuation of the BULLETIN OF POPULAR INFORMATION - of the Arnold Arboretum, Harvard University VOLUME 20 SEPTEMBER 23, 1960 NUMBER7 ILEX CRENATA AND ITS VARIETIES . Japanese holly was introduced into cultivation in the United States in THE1864 and has proved popular in all types of ornamental plantings. Its small, evergreen leaves and the dense, twiggy character of its branching give it quali- ties that make it useful as a specimen, a background planting or a hedge, since it withstands shearing very well indeed. American nurserymen have grown it from cuttings for a long time and have found that when seed is planted, numerous variations arise. In fact, there are so many now that if one studies the nursery catalogues carefully, one will find some 40 names purporting to be different vari- eties of this useful species and one nurseryman is known to have as many as 200 selected seedlings. In order to assist those interested in making a selection among these plants, this issue of Arnoldia is devoted to a cursory study of the varieties. Some are so new that definite information concerning their performance ~s not available, and will not be for several years. Because many people are interested m the lower- growing types of shrubs, especially those which require no special care or spraying to combat insects and diseases, this is a group well worth noting. Forty years ago there were only the species, the variety microphylla, and possi- bly one or two others available in the trade. Now there are forty names being listed, the most recent being one named ‘Glossy’ offered in an advertisement in the August 1, 1960, issue of the American A-urseryman. -
Ilex Aquifolium
Ilex aquifolium Ilex aquifolium (holly, common holly, English holly, European holly, or occasionally Christmas holly), is a species of holly native to western and southern Europe, northwest Africa, and southwest Asia. It is regarded as the type species of the genus Ilex, which by association is also called "holly". It is an evergreen tree or shrub found, for example, in shady areas of forests of oak and in beech hedges. In the British Isles it is one of very few native evergreen trees. It has a great capacity to adapt to different conditions and is a pioneer species that repopulates the margins of forests or clearcuts. I. aquifolium can exceed 10 m in height, but is often found at much smaller heights, typically 2–3 m (6.6–9.8 ft) tall and broad, with a straight trunk and pyramidal crown, branching from the base. It grows slowly and does not usually fully mature due to cutting or fire. It can live 500 years, but usually does not reach 100. Ilex aquifolium is the species of holly long associated with Christmas, and previously the Roman festival of Saturnalia. Its glossy green prickly leaves and bright red berries (produced only by the female plant) are represented in wreaths, garlands and cards wherever Christmas is celebrated. It is a subject of music and folklore, especially in the British tradition. It is also a popular ornamental shrub or hedge, with numerous cultivars in a range of colours. Description Ilex aquifolium grows to 10–25 m (33–82 ft) tall with a woody stem as wide as 40–80 cm (16– 31 in), rarely 100 cm (39 in) or more, in diameter. -
Meadows Farms Nurseries Nellie R. Stevens Holly
Nellie R. Stevens Holly Ilex 'Nellie R. Stevens' Height: 20 feet Spread: 10 feet Sunlight: Hardiness Zone: 5b Description: This splendid ornamental evergreen keeps its pyramidal shape with little to no pruning; it has glossy, toothed foliage and loads of brilliant red fruit that attracts birds; unusual drought and heat tolerance; a vigorous grower Ornamental Features Nellie R. Stevens Holly fruit Photo courtesy of NetPS Plant Finder Nellie R. Stevens Holly is primarily grown for its highly ornamental fruit. It features an abundance of magnificent red berries in late fall. It has attractive dark green foliage. The spiny oval leaves are highly ornamental and remain dark green throughout the winter. The flowers are not ornamentally significant. Landscape Attributes Nellie R. Stevens Holly is a dense multi-stemmed evergreen tree with a distinctive and refined pyramidal form. Its average texture blends into the landscape, but can be balanced by one or two finer or coarser trees or shrubs for an effective composition. This is a relatively low maintenance tree, and is best Nellie R. Stevens Holly pruned in late winter once the threat of extreme cold has Photo courtesy of NetPS Plant Finder passed. It is a good choice for attracting birds and bees to your yard. It has no significant negative characteristics. Nellie R. Stevens Holly is recommended for the following landscape applications; - Mass Planting - Hedges/Screening - Naturalizing And Woodland Gardens Planting & Growing Nellie R. Stevens Holly will grow to be about 20 feet tall at maturity, with a spread of 10 feet. It has a low canopy with a typical clearance of 2 feet from the ground, and is suitable for planting under power lines. -
Variety Height Characteristics Shiny Dark Green Leaves with Creamy White Margins
Variety Height Characteristics Shiny dark green leaves with creamy white margins. Argenteo- Bright red berries in winter. Ilex aquifolium 15-20’ marginatus Requires a male pollinator for berries to set. Use Gold Coast to pollinate. Very dense, compact growth habit. Dark green leaves are edged in bright Ilex aquifolium Gold Coast 4-6’ yellow. Male form with no berries, but will pollinate other hollies. Ilex aquifolium Hedgehog 15-20’ Great for containers or Ilex aquifolium Myrtifolia 6-8’ smaller gardens. Very slow growing. This extremely vigorous growing holly is great for Nellie R. Ilex aquifolium 12-18’ hedging. Female species- Stevens requires male to produce berries. An evergreen shrub with abundant red berries for Ilex cornuta Dazzler 8-10' winter! Spines on the margins of the foliage. Great for Christmas greenery. Valued for its delicate, narrow, twisted leaves and heavy crop of bright red fruit Ilex cornuta Needlepoint 8-10’ that lasts through winter. Performs well as a lush screen, border, accent or foundation plant. The hardies of all hollies. Bright red berries stand out Ilex meserveae Blue Girl 6-7’ against blue-green foliage. Makes a great hedge. Great as a background Ilex meserveae Blue Prince 12-15’ plant or hedge. Good pollinator for female hollies. Variety Height Characteristics Compact, pyramidal form. Dark green leaves with Ilex meserveae Castle Spire 8-10’ bright red berries. Use Castle Wall to pollinate. Great compact, pyramidal form. Great in place of Yew Ilex meserveae Castle Wall 5-8’ hedging. Use to pollinate Castle Spire. Dark green foliage and dark Ilex meserveae Dragon Lady 10-15’ purple stems. -
Holly - Ilex Sp
Holly - Ilex sp. General Information: Available in both evergreen and deciduous species, holly is grown for its toothed glossy green leaves and its showy berries, which are red in most popular varieties, but can also be a showy yellow. Holly can range from under one foot to over 80, and is found in both temperate and tropical regions. Both male and female plants are needed for fruiting. Family: Aquifoliaceae Lighting: Can tolerate both sun and shade, although semi-shade is preferable in midsummer. Increased light tends to produce dense foliage. Temperature: In general, evergreen varieties are hardy to zone 7, deciduous varieties to zone 5. Most varieties will require some frost protection, and all varieties should be sheltered from strong or cold winds. Watering: Needs a fair amount of water, especially before fruit production. Holly can be badly damaged by draught. Reduce watering in winter. Likes misting, unless it is in full sun. Feeding: Every two weeks during growth, using half strength liquid plant food, or bonsai food. Pruning and wiring: Cut back new shoots to the one or two nodes closest to the trunk. Branches can be very brittle, so shaping is best done by pruning rather than wiring. If wiring must be done, it is best to wire in spring- summer, taking care to protect the bark. Leaf pruning to reduce leaf size is possible. Suitable for all sizes and styles, although the evergreen varieties do not take as well to broom style. Ilex asprella has a tendency towards horzontal growth which must be compensated for; Ilex vomitoria, on the other hand, has a strong inclination to grow upwards. -
Foster Holly Production Ilex X Attenuata ‘Fosteri’ (Foster Holly) by Mark Halcomb UT Extension Area Nursery Specialist (Revised 5-28-2012)
Foster Holly Production Ilex x attenuata ‘Fosteri’ (Foster Holly) by Mark Halcomb UT Extension Area Nursery Specialist (Revised 5-28-2012) The secret to growing quality Foster Hollies is two shearings per year, a wide base and full bottom foliage. Full lower foliage is attained with a sufficiently wide spacing to allow sunlight penetration to the bottom foliage and weed control around the plant to prevent weeds from shading out the bottom foliage. Propagation Ilex x attenuata 'Fosteri' resulted from a cross of Ilex cassine x I. opaca. Seed propagation of Foster will yield 50 percent I. opaca. Cutting propagation is preferred. Cuttings are stuck Aug. to Nov. and can be potted in 6-10 weeks generally. A plant no less than 12 to 18” tall is transplanted in the field. Liner Selection Consider planting 1 gallon liners, at least for the first crop, to get a good start. During the first year, also consider buying rooted cuttings and plant them into a liner row or a transplant bed for 1-2 years with irrigation or pot them into #2 containers. Let them grow a larger root system before being spaced out to dig. This frees up some land and helps insure survival, taking a larger plant to the field. One of every 25 or 4 percent should be Foster # 4 males interplanted to increase fruit set if potential customers will be brought to the field. Consider growing a small number of the male #4's for those retailers and landscapers that realize and promote greater fruit set. Also consider growing a few Ilex x attenuata 'Blazer', 'Big John'; Ilex opaca 'Carolina #2' and 'Greenleaf'. -
Rutgers Gardens Holly Collection Accession List
Area Plant Species names, hybrid formulas, and cultivar names Common name Accession # Area #1 (American Holly Collection) 1 1 Ilex opaca 'Judge Brown' × Ilex opaca 'David' Hybrid American Holly 0200-R Ilex opaca 'Old Heavy Berry' × Ilex opaca 'Silica King' 1 2 *REMOVED Hybrid American Holly 0201-R 1 3 Ilex opaca 'Boyce Thompson Xanthocarpa' Yellow-fruited American Holly 0092-1945*A 1 4 Ilex opaca 'Jersey Delight' American Holly 0202-R Ilex opaca 'Old Heavy Berry' × Ilex opaca 1 5 'Point Pleasant #4' Hybrid American Holly 0203-R 1 6 Ilex R-5, T-47 × Ilex opaca 'Point Pleasant #5' Hybrid Holly 0204-R 1 7 Ilex 'Nellie R. Stevens' Hybrid Holly 0175-1960 1 8 Ilex opaca 'Hedgeholly' American Holly 0121-1949*A 1 9 Ilex opaca 'Portia Orton' Hybrid American Holly 0205-R 1 10 Ilex opaca Cardinal™ × Ilex opaca R-38, T-18 Hybrid American Holly 0206-R 1 11 Ilex opaca 'Clark's Valley #2' American Holly 0075-1950 1 12 Ilex opaca 'Mae' American Holly 0007-1954 1 13 Ilex opaca 'Farage' *REMOVED American Holly 0028-1946*G 1 14 Ilex opaca 'Manig' *REMOVED American Holly 0032-1946*I 1 15 Ilex aquifolium 'Ex. Herb Kale' English Holly 0207-R 1 16 Ilex opaca 'Jersey Knight' American Holly 0050-1965*A 1 17 Ilex opaca 'Griscom' American Holly 0033-1947*F 1 18 Ilex opaca 'Andorra' American Holly 0021-1950 1 19 Ilex opaca 'Reeve West' American Holly 0045-1947 1 20 Ilex opaca 'Clark' American Holly 0026-1946 1 21 Ilex opaca 'Judge Brown' × Ilex R-36, T-15 Hybrid Holly 0208-R 1 22 Ilex opaca 'Allen' American Holly 0031-1947 1 23 Ilex opaca 'Wheeler #4' American Holly