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1758 to the End of the Second Millennium CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM. II. TACHYPORINE GROUP LEE H. HERMAN BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY NUMBER 265 NEW YORK : 2001 Recent issues of the Bulletin may be purchased from the Museum. Lists of back issues of the Bulletin, Novitates, and Anthropological Papers published during the last five years are available at World Wide Web site http://nimidi.amnh.org. Or address mail orders to: American Museum of Natural History Library, Central Park West at 79th St., New York, N.Y. 10024. TEL: (212) 769- 5545. FAX: (212) 769-5009. E-MAIL: [email protected] CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM. II. TACHYPORINE GROUP LEE H. HERMAN Curator, Division of Invertebrate Zoology American Museum of Natural History BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Number 265, pages 651–1066 Issued July 18, 2001 (Parts I–VII) Copyright © American Museum of Natural History 2001 ISSN 0003-0090 CATALOG OF THE STAPHYLINIDAE, PARTS I–VII PART I, pages 1–650 Introduction Brief History of Taxonomic Studies of the Staphylinidae Research History • Biographical Sketches Overview of Classification Omaliine Group Dasycerinae • Empelinae • Glypholomatinae • Micropeplinae • Microsilphinae • Neophoninae • Omaliinae • Proteininae • Protopselaphinae PART II, pages 651–1066 Tachyporine Group Habrocerinae • Olisthaerinae • Phloeocharinae • Tachyporinae • Trichophyinae PART III, pages 1067–1806 Oxyteline Group Apateticinae • Osoriinae • Oxytelinae • Piestinae • Trigonurinae PART IV, pages 1807–2440 Staphylinine Group Euaesthetinae • Leptotyphlinae • Megalopsidiinae • Oxyporinae • Pseudopsinae • Solieriinae • Steninae PART V, pages 2441–3020 Staphylinine Group, continued Staphylininae Diochini • Maorothiini • Othiini • Platyprosopini • Staphylinini (Amblyopinina • Hyptiomina • Anisolinina • Philonthina) PART VI, pages 3021–3840 Staphylinine Group, continued Staphylininae, continued Staphylinini (Quediina • Staphylinina • Tanygnathinina • Xanthopygina • Incertae Sedis) Xantholinini Fossils: Incertae Sedis Protactinae†: Incertae Sedis PART VII, pages 3841–0000 Bibliography Index iii CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM ABSTRACT This catalog (published in seven parts, all released on the same day) is based on only the published lit- erature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylin- inae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or descrip- tion are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or syn- onyms are listed for each valid name. Annotated subsequent references are presented for all names. Dis- tributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Alesˇ Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies. Address correspondence to: Lee H. Herman, Curator, Division of Invertebrates, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA ([email protected]). Please send comments, corrections, or updates to either the postal or e-mail address. Reprints of new publications would be helpful and appreciated. iv CONTENTS PART II TACHYPORINE GROUP Tachyporine Group . 651 Habrocerinae . 651 Olisthaerinae . 659 Phloeocharinae . 663 Tachyporinae . 675 Cordobanini . 676 Deropini . 676 Megarthropsini . 678 Mycetoporini . 679 Symmixini . 802 Tachyporini . 803 Vatesini . 1055 Tachyporinae: Tribe Incertae Sedis . 1059 Trichophyinae . 1061 PARTS I, III–VII Introduction, History, and Biographical Sketches . Part I Omaliine Group . Part I Oxyteline Group . Part III Staphylinine Group . Parts IV–VI Staphylinidae: Subfamily Incertae Sedis . Part VI Protactinae† . Part VI Bibliography . Part VII Index . Part VII v HABROCERINAE [2 genera; 21 species and 1 species incertae sedis; Neotropical, Nearctic, Palaearctic, Oriental, and Australian (introduction) Regions] Habrocerinae Mulsant and Rey, 1877: 66 (cited as Habrocériens; genera included: Habro- cerus). TYPE GENUS: Habrocerus (see Newton and Thayer, 1992: 56). — Eichelbaum, 1909: 190 (generic references; number of species in genus; general distribution of genus). — Bernhauer and Schubert, 1916: 441 (world catalog; genera). — Porta, 1926: 131 (characters; species of Italy). — Böving and Craighead, 1930: 30 (larval characters). — Scheerpeltz, 1933: 1469 (world catalog supplement). — Scheerpeltz, 1940: 51 (key to Palaearctic genera). — Bernet Kempers, 1941: 34, 36 (larval characters). — Blackwelder, 1944: 1450 (checklist of species from Mexico, Central and South America, and the West Indies). — V. Hansen, 1952: 241 (characters; Danish species). — Hatch, 1957: 48, 149 (characters). — Arnett, 1963: 280 (characters; list of North American genera). — Székessy, 1963: 63 (characters). — Lohse, 1964: 220 (characters; central European species). — Moore, 1964a: 89 (characters; notes). — Kasule, 1966: 271 (included in Tachyporinae in larval key). — Palm, 1966: 3 (characters; species of Sweden). — Pototskaia, 1967: 17 (larval characters). — Scheerpeltz, 1968a: 97 (catalog of Austrian species). — Moore and Legner, 1974h: 475 (catalog of the taxonomy, biology, and ecology of develop- mental stages of species of America north of Mexico). — Moore and Legner, 1975: 304 (catalog; North America north of Mexico). — Pope, 1977: 32 (list of British species). — Topp, 1978: 307 (larval characters). — Welch, 1978: 61 (egg characters). — Muona, 1979: 20 (list of Scandinavian species). — Burakowski, Mroczkowski, and Stefan´ska, 1980: 155 (catalog; Poland). — Watt and McColl, 1982: 19, 20 (characters; notes on New Zealand taxa). — Newton, 1985: 200 (notes on austral distribution). — Newton and Thayer, 1992: 56 (summary of current classification). — Tóth, 1993: 1 (characters). — Assing and Wunderle, 1995: 308, 353 (revision of and key to genera and species; characters). — Lawrence and Newton, 1995: 826 (discussion). — Sparacio, 1995: 166 (characters; species of Sicily). — Klimaszewski, Newton, and Thayer, 1996: 148 (review of subfamily for New Zealand; notes; list of New Zealand genera; number of New Zealand species). — M. Hansen, 1996: 38, 111 (listed in Tachyporine group of subfamilies; monophyletic group; list of Danish species). — Downie and Arnett, 1996: 483 (characters; key to species of Northeastern North America). — M. Hansen, 1997: 123, 175, 178, 179 (autapomorphic features; key adult and larval characters; notes). — Newton, Thayer, Ashe, and Chandler, 2000: 358 (characters; notes). — Klimaszewski, 2000: 44 (characters; notes on classification, natural history, and distribution in Canada). Habroceridae Mulsant and Rey, 1877. — Coiffait and Saiz, 1968: 464 (Chilean species). — Outerelo and Gamarra, 1985: 14 (genera of Iberian Peninsula). 651 652 HABROCERINAE Bulletin AMNH Habrocerini Mulsant and Rey, 1877. — Blatchley, 1910: 457 (characters). — Arnett, 1963: 280 (characters; list of North American genera). — Kasule, 1968: 124 (larval characters; notes). Habrocerina Mulsant and Rey, 1877. — Jakobson, 1908: 447 (characters). — Johansen, 1914: 293 (characters). Habrocerus [15 species (plus 1 incertae sedis); Neotropical, Nearctic, Palaearctic, Oriental, and Australian (introduction) Regions] Habrocerus Erichson, 1839a: 400 (species included: capillaricornis). TYPE SPECIES: Tachy- porus capillaricornis Gravenhorst, fixed by monotypy. — Heer, 1839: 292 (characters). — Erichson, 1839b: 242 (characters). — Stephens, 1839: 352 (characters). — Redtenbacher, 1849: 682 (characters). — Schaum, 1852: 24 (list of species; Europe). — Lacordaire, 1854: 56 (characters; notes). — Wollaston, 1854: 568 (characters; notes). — Fairmaire and Laboulbène, 1856: 481 (characters). — Jacquelin du Val, 1856: 26 (characters). — Kraatz, 1857d: 391 (characters). — Redtenbacher, 1857: 173 (characters; Austrian species). — Thomson, 1858: 36 (characters). — G. Waterhouse, 1858: 20 (catalog of British species). — Thomson, 1859: 45 (characters; type species: capillaricornis). — Schaum, 1859: 24 (catalog; European species). — Thomson, 1861: 146 (characters). — Fauvel, 1866a: 331 [= 1866b: 82] (characters). — Wencker and Silbermann, 1866: 25 (list of species; collecting notes; France). — Gemminger and Harold, 1868: 553 (catalog). — Jacquelin du Val, 1868: 64 (catalog; European species). — Pandellé, 1869: 275, 310 (characters; key to species). — Sharp, 1871d: 10 (list of British species). — Redtenbacher, 1874: 184 (characters). — Fauvel, 1875: 590 [= 1876: 72] (characters). — Fauvel, 1875a: XXXVII
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