Catalog of the Staphylinidae( Insecta: Coleoptera). 1758 to the End of The
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CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM. I. INTRODUCTION, HISTORY, BIOGRAPHICAL SKETCHES, AND OMALIINE GROUP LEE H. HERMAN BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY NUMBER 265 NEW YORK : 2001 Recent issues of the Bulletin may be purchased from the Museum. Lists of back issues of the Bulletin, Novitates, and Anthropological Papers published during the last five years are available at World Wide Web site http://nimidi.amnh.org. Or address mail orders to: American Museum of Natural History Library, Central Park West at 79th St., New York, N.Y. 10024. TEL: (212) 769- 5545. FAX: (212) 769-5009. E-MAIL: [email protected] CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM. I. INTRODUCTION, HISTORY, BIOGRAPHICAL SKETCHES, AND OMALIINE GROUP LEE H. HERMAN Curator, Division of Invertebrate Zoology American Museum of Natural History BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Number 265, pages 1–650 Issued July 18, 2001 (Parts I–VII) Copyright © American Museum of Natural History 2001 ISSN 0003-0090 ... I Vnnametf'friassic fFossi{ .. Top: Staphylinus hirtus Linne. [he first species of Staphylinidae described !illustration after Panzer, 1796, Faunae insectorum Germanicae). Bottom: Unnamed Triassic fossil, the oldest known species of Staphy linidae (from photOgraph of original fossil). CATALOG OF THE STAPHYLINIDAE, PARTS I–VII PART I, pages 1–650 Introduction Brief History of Taxonomic Studies of the Staphylinidae Research History • Biographical Sketches Overview of Classification Omaliine Group Dasycerinae • Empelinae • Glypholomatinae • Micropeplinae • Microsilphinae • Neophoninae • Omaliinae • Proteininae • Protopselaphinae PART II, pages 651–1066 Tachyporine Group Habrocerinae • Olisthaerinae • Phloeocharinae • Tachyporinae • Trichophyinae PART III, pages 1067–1806 Oxyteline Group Apateticinae • Osoriinae • Oxytelinae • Piestinae • Trigonurinae PART IV, pages 1807–2440 Staphylinine Group Euaesthetinae • Leptotyphlinae • Megalopsidiinae • Oxyporinae • Pseudopsinae • Solieriinae • Steninae PART V, pages 2441–3020 Staphylinine Group, continued Staphylininae Diochini • Maorothiini • Othiini • Platyprosopini • Staphylinini (Amblyopinina • Hyptiomina • Anisolinina • Philonthina) PART VI, pages 3021–3840 Staphylinine Group, continued Staphylininae, continued Staphylinini (Quediina • Staphylinina • Tanygnathinina • Xanthopygina • Incertae Sedis) Xantholinini Staphylinidae: Subfamily Incertae Sedis Protactinae† PART VII, pages 3841–0000 Bibliography Index iii CATALOG OF THE STAPHYLINIDAE (INSECTA: COLEOPTERA). 1758 TO THE END OF THE SECOND MILLENNIUM ABSTRACT This catalog (published in seven parts, all released on the same day) is based on only the published lit- erature for the Staphylinidae. Of the 32 subfamilies, the following 28 are included herein: Apateticinae, Dasycerinae, Empelinae, Euaesthetinae, Glypholomatinae, Habrocerinae, Leptotyphlinae, Megalopsidiinae, Micropeplinae, Microsilphinae, Neophoninae, Olisthaerinae, Omaliinae, Osoriinae, Oxyporinae, Oxytelinae, Phloeocharinae, Piestinae, Protactinae†, Proteininae, Protopselaphinae, Pseudopsinae, Solieriinae, Staphylin- inae, Steninae, Tachyporinae, Trichophyinae, and Trigonurinae. The Aleocharinae, Paederinae, Pselphinae, and Scaphidiinae are excluded from this edition of the catalog. References to the original citation or descrip- tion are given for available family-group, genus-group, and species-group names of both extant and extinct forms. The type genus is cited for each family-group name, the type species for each genus-group name, and the type locality for each species-group name. Where appropriate, all subgenera, subspecies, or syn- onyms are listed for each valid name. Annotated subsequent references are presented for all names. Dis- tributional summaries are given for each valid taxon. Full bibliographic citations are in Part VII. A short historical review, coauthored with Alesˇ Smetana, follows the Introduction (Part I), with the main focus on biographical sketches that include many photographs. The goal of this catalog is to summarize the current state of knowledge of the family and to stimulate worldwide monographic studies. Address correspondence to: Lee H. Herman, Curator, Division of Invertebrates, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024-5192, USA ([email protected]). Please send comments, corrections, or updates to either the postal or e-mail address. Reprints of new publications would be helpful and appreciated. iv CONTENTS PART I INTRODUCTION, HISTORY, BIOGRAPHICAL SKETCHES, AND OMALIINE GROUP Introduction . .1 Acknowledgments . 14 Brief History of Taxonomic Studies of the Staphylinidae Including Biographical Sketches of the Investigators, by Alesˇ Smetana and Lee H. Herman . 17 Research History . 17 Biographical Sketches . 38 Overview of Classification . 161 Staphylinidae . 167 Omaliine Group . 175 Dasycerinae . 175 Empelinae . 179 Glypholomatinae . 181 Micropeplinae . 183 Microsilphinae . 205 Neophoninae . 207 Omaliinae . 209 Anthophagini . 210 Aphaenostemmini . 377 Corneolabiini . 378 Coryphiini . 381 Boreaphilina . 381 Coryphiina . 389 Eusphalerini . 409 Hadrognathini . 467 Omaliini . 468 Proteininae . 599 Anepiini . 600 Austrorhysini . 600 Nesoneini . 601 Proteinini . 602 Silphotelini . 648 Protopselaphinae . 649 PARTS II–VII Tachyporine Group . Part II Oxyteline Group . Part III Staphylinine Group . Parts IV–VI v vi CONTENTS Bulletin AMNH Staphylinidae: Subfamily Incertae Sedis . Part VI Protactinae† . Part VI Bibliography . Part VII Index . Part VII INTRODUCTION atalogs are among the most indispensable tools used by systematists and other biologists; Ctheir scarcity is a major impediment to progress in the study of a group. Without catalogs it is laborious to determine what groups are described, what species are included in each group, and what has been published about them. Minimally, catalogs present a list of species and gen- era. Maximally, they tabulate the names of the taxa with their author and date and place of pub- lication; their subsequent use; their synonyms; notation of homonyms; the current classification; the type species for each genus-group name; the type locality for each species-group name; the original, subsequent, and current genus-species combination; the geographical distribution, habi- tat, host, natural history; and so on for each species, in short, every published mention of a taxon. Ideally, catalogs permit us to keep track of available names, valid names, unavailable names, and the composition of taxonomic groups. Catalogs state the spelling, validity, and availability of names and their dates of publication; verification of these data, particularly the last, are not as simple as might first appear. They permit discovery of homonyms and help prevent publication of others. They organize the scattered original descriptions and subsequent literature. Catalogs summarize the historical development and studies of a group. They outline the currently recog- nized classification and the state of knowledge of a taxon, bringing together in one place all that we know about it. Catalogs provide a common language through which we can communicate, and without which we would be immersed in a bedlam of names and literature. My intention is to provide a guide to the taxa and literature of the Staphylinidae and to make both the catalog as practical as possible and the literature accessible. This project was undertaken because a new catalog was desperately needed. The Bernhauer- Schubert-Scheerpeltz catalogs and supplements were long out of date and no one else volunteered to produce a new one. It had become necessary to review over 60 years of the Zoological Record to begin a new research project, to gather information about a taxon and its composition, or to know which names were valid and which were synonyms. The most recent catalog for the Staphylinidae of the world was published in six parts between 1910 and 1926 (Bernhauer and Schubert, 1910, 1911, 1912, 1914, 1916; Bernhauer and Scheer- peltz, 1926) with major supplements by Scheerpeltz (1933, 1934). All subsequent catalogs or checklists have been restricted to specified geographical regions. Some examples of such lists and catalogs include those for Mexico, Central America, the West Indies, and South America (Blackwelder, 1944), central Europe (Horion, 1963, 1965, 1967; Lucht, 1987), Austria (Scheer- peltz, 1968a), the former Soviet Union (Tikhomirova, 1973a), Taiwan (Shibata, 1973c, 1986a), America north of Mexico (Moore and Legner, 1975), Japan (Shibata, 1976, 1977, 1983, 1984, 1985a), Britain (Pope, 1977), Poland (Burakowski et al., 1979; Burakowski et al., 1980), Fenno- scandia and Denmark (Muona, 1979), Korea (Yuh et al., 1985), Fennoscandia, Denmark, and the Baltic region (Silfverberg, 1992), the former Czechoslovakia (various authors, in Jelínek, 1993, Check-list of Czechoslovak Insects IV), Denmark (Hansen, 1996, along with subsequent multi- authored supplements), Latvia (Telnov et al., 1997), Italy (in Ciceroni, Puthz, and Zanetti, 1995), Ireland (Anderson et al., 1997), and northeastern Asia (Ryabukhin, 1999), and so on. The pre- ceding list is not complete; it includes only some of the multitude of such lists, particularly for Europe, many of which are cited in the catalog and many of which are not. These checklists pro- vide lists of taxa, citations to their original descriptions, and (in some cases) their synonyms for