What Natural Factors Limit the Growth of Ecosystems?
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Rhythms of the Brain
Rhythms of the Brain György Buzsáki OXFORD UNIVERSITY PRESS Rhythms of the Brain This page intentionally left blank Rhythms of the Brain György Buzsáki 1 2006 3 Oxford University Press, Inc., publishes works that further Oxford University’s objective of excellence in research, scholarship, and education. Oxford New York Auckland Cape Town Dar es Salaam Hong Kong Karachi Kuala Lumpur Madrid Melbourne Mexico City Nairobi New Delhi Shanghai Taipei Toronto With offices in Argentina Austria Brazil Chile Czech Republic France Greece Guatemala Hungary Italy Japan Poland Portugal Singapore South Korea Switzerland Thailand Turkey Ukraine Vietnam Copyright © 2006 by Oxford University Press, Inc. Published by Oxford University Press, Inc. 198 Madison Avenue, New York, New York 10016 www.oup.com Oxford is a registered trademark of Oxford University Press All rights reserved. No part of this publication may be reproduced, stored in a retrieval system, or transmitted, in any form or by any means, electronic, mechanical, photocopying, recording, or otherwise, without the prior permission of Oxford University Press. Library of Congress Cataloging-in-Publication Data Buzsáki, G. Rhythms of the brain / György Buzsáki. p. cm. Includes bibliographical references and index. ISBN-13 978-0-19-530106-9 ISBN 0-19-530106-4 1. Brain—Physiology. 2. Oscillations. 3. Biological rhythms. [DNLM: 1. Brain—physiology. 2. Cortical Synchronization. 3. Periodicity. WL 300 B992r 2006] I. Title. QP376.B88 2006 612.8'2—dc22 2006003082 987654321 Printed in the United States of America on acid-free paper To my loved ones. This page intentionally left blank Prelude If the brain were simple enough for us to understand it, we would be too sim- ple to understand it. -
Effective Population Size and Genetic Conservation Criteria for Bull Trout
North American Journal of Fisheries Management 21:756±764, 2001 q Copyright by the American Fisheries Society 2001 Effective Population Size and Genetic Conservation Criteria for Bull Trout B. E. RIEMAN* U.S. Department of Agriculture Forest Service, Rocky Mountain Research Station, 316 East Myrtle, Boise, Idaho 83702, USA F. W. A LLENDORF Division of Biological Sciences, University of Montana, Missoula, Montana 59812, USA Abstract.ÐEffective population size (Ne) is an important concept in the management of threatened species like bull trout Salvelinus con¯uentus. General guidelines suggest that effective population sizes of 50 or 500 are essential to minimize inbreeding effects or maintain adaptive genetic variation, respectively. Although Ne strongly depends on census population size, it also depends on demographic and life history characteristics that complicate any estimates. This is an especially dif®cult problem for species like bull trout, which have overlapping generations; biologists may monitor annual population number but lack more detailed information on demographic population structure or life history. We used a generalized, age-structured simulation model to relate Ne to adult numbers under a range of life histories and other conditions characteristic of bull trout populations. Effective population size varied strongly with the effects of the demographic and environmental variation included in our simulations. Our most realistic estimates of Ne were between about 0.5 and 1.0 times the mean number of adults spawning annually. We conclude that cautious long-term management goals for bull trout populations should include an average of at least 1,000 adults spawning each year. Where local populations are too small, managers should seek to conserve a collection of interconnected populations that is at least large enough in total to meet this minimum. -
Routledge Handbook of Ecological and Environmental Restoration the Principles of Restoration Ecology at Population Scales
This article was downloaded by: 10.3.98.104 On: 26 Sep 2021 Access details: subscription number Publisher: Routledge Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: 5 Howick Place, London SW1P 1WG, UK Routledge Handbook of Ecological and Environmental Restoration Stuart K. Allison, Stephen D. Murphy The Principles of Restoration Ecology at Population Scales Publication details https://www.routledgehandbooks.com/doi/10.4324/9781315685977.ch3 Stephen D. Murphy, Michael J. McTavish, Heather A. Cray Published online on: 23 May 2017 How to cite :- Stephen D. Murphy, Michael J. McTavish, Heather A. Cray. 23 May 2017, The Principles of Restoration Ecology at Population Scales from: Routledge Handbook of Ecological and Environmental Restoration Routledge Accessed on: 26 Sep 2021 https://www.routledgehandbooks.com/doi/10.4324/9781315685977.ch3 PLEASE SCROLL DOWN FOR DOCUMENT Full terms and conditions of use: https://www.routledgehandbooks.com/legal-notices/terms This Document PDF may be used for research, teaching and private study purposes. Any substantial or systematic reproductions, re-distribution, re-selling, loan or sub-licensing, systematic supply or distribution in any form to anyone is expressly forbidden. The publisher does not give any warranty express or implied or make any representation that the contents will be complete or accurate or up to date. The publisher shall not be liable for an loss, actions, claims, proceedings, demand or costs or damages whatsoever or howsoever caused arising directly or indirectly in connection with or arising out of the use of this material. 3 THE PRINCIPLES OF RESTORATION ECOLOGY AT POPULATION SCALES Stephen D. -
A Limiting Factor
Published on Plants in Action (http://plantsinaction.science.uq.edu.au/edition1) Home > Printer-friendly > Printer-friendly Chapter 15 - Water: a limiting factor [1] A superb stand of flooded gums, (Eucalyptus grandis) near Coffs Habour, northern New South Wales, 'each tall because of each' (Les Murray (1991), Collected Poems) (Photograph by Ken Eldridge, supplied by Peter Burgess, CSIRO Forestry and Forest Products) With perspective phrasing, Les Murray (1991) summarises structural aspects of a gum forest as: 'Flooded gums on creek ground, each tall because of each' and on conceptualising water relations, 'Foliage builds like a layering splash: ground water drily upheld in edge-on, wax rolled, gall-puckered leaves upon leaves. The shoal life of parrots up there.' (Les Murray, Collected Poems, 1991) Introduction Life-giving water molecules, fundamental to our biosphere, are as remarkable as they are abundant. Hydrogen bonds, enhanced by dipole forces, confer extraordinary physical properties on liquid water that would not be expected from atomic structure alone. Water has the strongest surface tension, biggest specific heat, largest latent heat of vaporisation and, with the exception of mercury, the best thermal conductivity of any known natural liquid. A high specific grav-ity is linked to a high specific heat, and very few natural substances require 1 calorie to increase the temperature of 1 gram by 1ºC. Similarly, a high heat of vaporisation means that 500 calories are required to convert 1 gram of water from liquid to vapour at 100ºC. This huge energy requirement (latent heat of vaporisation, Section 14.5) ties up much heat so that massive bodies of water contribute to climatic stability, while tiny bodies of water are significant for heat budgets of organisms. -
Evolutionary Restoration Ecology
ch06 2/9/06 12:45 PM Page 113 189686 / Island Press / Falk Chapter 6 Evolutionary Restoration Ecology Craig A. Stockwell, Michael T. Kinnison, and Andrew P. Hendry Restoration Ecology and Evolutionary Process Restoration activities have increased dramatically in recent years, creating evolutionary chal- lenges and opportunities. Though restoration has favored a strong focus on the role of habi- tat, concerns surrounding the evolutionary ecology of populations are increasing. In this con- text, previous researchers have considered the importance of preserving extant diversity and maintaining future evolutionary potential (Montalvo et al. 1997; Lesica and Allendorf 1999), but they have usually ignored the prospect of ongoing evolution in real time. However, such contemporary evolution (changes occurring over one to a few hundred generations) appears to be relatively common in nature (Stockwell and Weeks 1999; Bone and Farres 2001; Kin- nison and Hendry 2001; Reznick and Ghalambor 2001; Ashley et al. 2003; Stockwell et al. 2003). Moreover, it is often associated with situations that may prevail in restoration projects, namely the presence of introduced populations and other anthropogenic disturbances (Stockwell and Weeks 1999; Bone and Farres 2001; Reznick and Ghalambor 2001) (Table 6.1). Any restoration program may thus entail consideration of evolution in the past, present, and future. Restoration efforts often involve dramatic and rapid shifts in habitat that may even lead to different ecological states (such as altered fire regimes) (Suding et al. 2003). Genetic variants that evolved within historically different evolutionary contexts (the past) may thus be pitted against novel and mismatched current conditions (the present). The degree of this mismatch should then determine the pattern and strength of selection acting on trait variation in such populations (Box 6.1; Figure 6.1). -
3 Limiting Factors and Threats
LOWER COLUMBIA SALMON RECOVERY & SUBBASIN PLAN December 2004 3 Limiting Factors and Threats 3 LIMITING FACTORS AND THREATS........................................................................3-1 3.1 HABITAT –STREAMS .....................................................................................................3-2 3.1.1 Background..........................................................................................................3-2 3.1.2 Limiting Factors...................................................................................................3-3 3.1.3 Threats................................................................................................................3-22 3.2 ESTUARY AND LOWER MAINSTEM HABITAT ..............................................................3-26 3.2.1 Background........................................................................................................3-26 3.2.2 Limiting Factors.................................................................................................3-27 3.2.3 Threats................................................................................................................3-36 3.3 HABITAT – OCEAN ......................................................................................................3-38 3.3.1 Background........................................................................................................3-38 3.3.2 Limiting Factors.................................................................................................3-38 -
Population Biology & Life Tables
EXERCISE 3 Population Biology: Life Tables & Theoretical Populations The purpose of this lab is to introduce the basic principles of population biology and to allow you to manipulate and explore a few of the most common equations using some simple Mathcad© wooksheets. A good introduction of this subject can be found in a general biology text book such as Campbell (1996), while a more complete discussion of pop- ulation biology can be found in an ecology text (e.g., Begon et al. 1990) or in one of the references listed at the end of this exercise. Exercise Objectives: After you have completed this lab, you should be able to: 1. Give deÞnitions of the terms in bold type. 2. Estimate population size from capture-recapture data. 3. Compare the following sets of terms: semelparous vs. iteroparous life cycles, cohort vs. static life tables, Type I vs. II vs. III survivorship curves, density dependent vs. density independent population growth, discrete vs. continuous breeding seasons, divergent vs. dampening oscillation cycles, and time lag vs. generation time in population models. 4. Calculate lx, dx, qx, R0, Tc, and ex; and estimate r from life table data. 5. Choose the appropriate theoretical model for predicting growth of a given population. 6. Calculate population size at a particular time (Nt+1) when given its size one time unit previous (Nt) and the corre- sponding variables (e.g., r, K, T, and/or L) of the appropriate model. 7. Understand how r, K, T, and L affect population growth. Population Size A population is a localized group of individuals of the same species. -
Equilibrium Theory of Island Biogeography: a Review
Equilibrium Theory of Island Biogeography: A Review Angela D. Yu Simon A. Lei Abstract—The topography, climatic pattern, location, and origin of relationship, dispersal mechanisms and their response to islands generate unique patterns of species distribution. The equi- isolation, and species turnover. Additionally, conservation librium theory of island biogeography creates a general framework of oceanic and continental (habitat) islands is examined in in which the study of taxon distribution and broad island trends relation to minimum viable populations and areas, may be conducted. Critical components of the equilibrium theory metapopulation dynamics, and continental reserve design. include the species-area relationship, island-mainland relation- Finally, adverse anthropogenic impacts on island ecosys- ship, dispersal mechanisms, and species turnover. Because of the tems are investigated, including overexploitation of re- theoretical similarities between islands and fragmented mainland sources, habitat destruction, and introduction of exotic spe- landscapes, reserve conservation efforts have attempted to apply cies and diseases (biological invasions). Throughout this the theory of island biogeography to improve continental reserve article, theories of many researchers are re-introduced and designs, and to provide insight into metapopulation dynamics and utilized in an analytical manner. The objective of this article the SLOSS debate. However, due to extensive negative anthropo- is to review previously published data, and to reveal if any genic activities, overexploitation of resources, habitat destruction, classical and emergent theories may be brought into the as well as introduction of exotic species and associated foreign study of island biogeography and its relevance to mainland diseases (biological invasions), island conservation has recently ecosystem patterns. become a pressing issue itself. -
COULD R SELECTION ACCOUNT for the AFRICAN PERSONALITY and LIFE CYCLE?
Person. individ.Diff. Vol. 15, No. 6, pp. 665-675, 1993 0191-8869/93 S6.OOf0.00 Printedin Great Britain.All rightsreserved Copyright0 1993Pergamon Press Ltd COULD r SELECTION ACCOUNT FOR THE AFRICAN PERSONALITY AND LIFE CYCLE? EDWARD M. MILLER Department of Economics and Finance, University of New Orleans, New Orleans, LA 70148, U.S.A. (Received I7 November 1992; received for publication 27 April 1993) Summary-Rushton has shown that Negroids exhibit many characteristics that biologists argue result from r selection. However, the area of their origin, the African Savanna, while a highly variable environment, would not select for r characteristics. Savanna humans have not adopted the dispersal and colonization strategy to which r characteristics are suited. While r characteristics may be selected for when adult mortality is highly variable, biologists argue that where juvenile mortality is variable, K character- istics are selected for. Human variable birth rates are mathematically similar to variable juvenile birth rates. Food shortage caused by African drought induce competition, just as food shortages caused by high population. Both should select for K characteristics, which by definition contribute to success at competition. Occasional long term droughts are likely to select for long lives, late menopause, high paternal investment, high anxiety, and intelligence. These appear to be the opposite to Rushton’s r characteristics, and opposite to the traits he attributes to Negroids. Rushton (1985, 1987, 1988) has argued that Negroids (i.e. Negroes) were r selected. This idea has produced considerable scientific (Flynn, 1989; Leslie, 1990; Lynn, 1989; Roberts & Gabor, 1990; Silverman, 1990) and popular controversy (Gross, 1990; Pearson, 1991, Chapter 5), which Rushton (1989a, 1990, 1991) has responded to. -
Population Growth Is a Critical Factor in Specie’S Ability to Maintain Homeostasis Within Its Environment
Chapter 4-B1 Population Ecology Population growth is a critical factor in specie’s ability to maintain homeostasis within its environment. Main themes ~Scientists study population characteristics to better understand growth and distribution of organisms. ~Populations have different distributions and densities depending on the species represented in the population ~Organisms in a population compete for energy sources, such as food and sunlight ~Homeostasis within a population is controlled by density-dependent and density-independent limiting factors ~Human population varies little overall, but can change greatly within small populations. I. Population dynamics A. 1. Population density a. defined- b. eg. deer in Newark deer in 5 mi. sq. area 2. a. defined-organism pattern in area (eg. more deer on Londondale than on Green Wave Drive) b. c. Types p. 93 -UNIFORM dispersion-relatively even distribution of a species. Eg. pine forest -CLUMPED dispersion-organism are clumped near food. Eg. schools of fish, eg. mushrooms on a fallen log -_________________-unpredictable & changing. Eg. deer, large fish, etc. 3. a. area where organisms CAN/ARE living b. Meaning of Distribution -in Science~it is area where something lives and reproduces -in everyday language ~it means distribution (passing out) eg. of report cards. B. 1. Limiting factors-biotic or abiotic factors that keep a population from increasing forever (indefinitely). a. b. amount of space=abiotic c. 2. Density-independent factors a. b. independent=doesn’t rely on the number of organisms, would happen regardless of the number of organisms c. it is usually abiotic (limits population growth) -flood (eg. no matter how many deer in an area) - -blizzard - -fire (Ponderosa pine needs fire to kill undergrowth which takes all the soil nutrients from them. -
The Basics of Population Dynamics Greg Yarrow, Professor of Wildlife Ecology, Extension Wildlife Specialist
The Basics of Population Dynamics Greg Yarrow, Professor of Wildlife Ecology, Extension Wildlife Specialist Fact Sheet 29 Forestry and Natural Resources Revised May 2009 All forms of wildlife, regardless of the species, will respond to changes in density dependence. These concepts are important for landowners habitat, hunting or trapping, and weather conditions with fluctuations and natural resource managers to understand when making decisions in animal numbers. Most landowners have probably experienced affecting wildlife on private land. changes in wildlife abundance from year to year without really knowing why there are fewer individuals in some years than others. How Many Offspring Can Wildlife Have? In many cases, changes in abundance are normal and to be expected. Most people realize that some wildlife species can produce more The purpose of the information presented here is to help landowners offspring than others. Bobwhite quail are genetically programmed to lay understand why animal numbers may vary or change. While a number an average of 14 eggs per clutch. Each species has a maximum genetic of important concepts will be discussed, one underlying theme should reproductive potential or biotic potential. always be remembered. Regardless of whether property is managed or not in any given year, there is always some change in the habitat, Biotic potential describes a population’s ability to grow over time however small. Wildlife must adjust to this change and, therefore, no through reproduction. Most bat species are likely to produce one population is ever the same from one year to the next. offspring per year. In contrast, a female cottontail rabbit will have a litter size of approximately 5. -
Trade-Off Between R-Selection and K-Selection in Drosophila Populations (Population Dynamics/Density-Dependent Selection/Evolution/Drosophila Melanogaster) LAURENCE D
Proc. Natl Acad. Sci. USA Vol. 78, No. 2, pp. 1303-1305, February 1981 Population Biology Trade-off between r-selection and K-selection in Drosophila populations (population dynamics/density-dependent selection/evolution/Drosophila melanogaster) LAURENCE D. MUELLERt AND FRANCISCO J. AYALA Department ofGenetics, University ofCalifornia, Davis, California 95616 Contributed by Franciscoj. Ayala, October 17, 1980 ABSTRACT Density-dependent genetic evolution was tested that show a trade-off-i.e., genotypes with high rs have low Ks in experimental populations of Drosophila melanogaster subject and vice versa-the outcome of evolution in Roughgarden's for eight generations to natural selection under high (K-selection) model is dependent on the environment. In stable environ- or low (r-selection) population density regimes. The test consisted of determining at high and at low densities the per capita rate of ments, the population reaches its carrying capacity and the gen- population growth of the selected populations. At high densities, otype with the largest value ofK ultimately becomes established the K-selected populations showed a higher per capita rate ofpop- and all others are eliminated. When the population is often be- ulation growth than did the r-selected populations, but the reverse low its carrying capacity owing to frequent episodes ofdensity- was true at lowdensities.These results corroborate the predictions independent mortality, the genotype with the highest r is fa- derived from formal models ofdensity-dependent selection. How- vored. Thus, according to this model evolution favors the gen- ever, no evidence of a trade-off in per capita rate of growth was observed in 25 populations ofD.