Dynamique Des Paléoenvironnements À Huîtres Du

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Dynamique Des Paléoenvironnements À Huîtres Du Dynamique des paléoenvironnements à huîtres du Crétacé Supérieur nord-aquitain (SO France) et du Mio-Pliocène andalou (SE Espagne) : biodiversité, analyse séquentielle, biogéochimie Blaise Videt To cite this version: Blaise Videt. Dynamique des paléoenvironnements à huîtres du Crétacé Supérieur nord-aquitain (SO France) et du Mio-Pliocène andalou (SE Espagne) : biodiversité, analyse séquentielle, biogéochimie. Autre. Université Rennes 1, 2003. Français. tel-00006038 HAL Id: tel-00006038 https://tel.archives-ouvertes.fr/tel-00006038 Submitted on 7 May 2004 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. N° Ordre : 2939 THESE Présentée DEVANT L’ UNIVERSITE DE RENNES 1 pour obtenir le grade de : DOCTEUR DE L’ UNIVERSITE DE RENNES 1 Mention Sciences de la Terre PAR Blaise VIDET Equipe d’accueil : Géosciences Rennes Ecole Doctorale : Sciences de la Matière Composante universitaire : UFR Structure et Propriété de la Matière Titre de la Thèse : DYNAMIQUE DES PALEOENVIRONNEMENTS A HUITRES DU CRETACE SUPERIEUR NORD-AQUITAIN (SO FRANCE) ET DU MIO-PLIOCENE ANDALOU (SE ESPAGNE) : BIODIVERSITE, ANALYSE SEQUENTIELLE, BIOGEOCHIMIE Soutenue le 11 Décembre 2003 devant la commission d’Examen COMPOSITION DU JURY Ilhem Bentaleb Codirectrice de thèse Marc Floquet Rapporteur François Guillocheau Examinateur Agnès Lauriat-Rage Rapporteur Didier Néraudeau Directeur de thèse Jean-Pierre Platel Examinateur Remerciements Au terme de tout travail, il est d’usage de remercier tous ceux et celles qui, d’une manière ou d’une autre, ont contribué à sa réalisation. Cette tache n’est pas des plus aisée, en ce sens qu’il est souvent difficile d’identifier l’influence de chacun dans un travail souvent réalisé en équipe. Ainsi la liste ci-dessous est loin d’être exhaustive. Mes tous premiers remerciements s’adressent à Didier Néraudeau, mon directeur de thèse, qui a bien voulu me faire confiance, et qui a su m’encourager grâce à son dynamisme et à sa disponibilité. Je remercie également Ilhem Bentaleb, ma codirectrice, qui m’a initié au monde des isotopes stables. Je remercie également les personnes qui ont accepté d’être dans mon jury, Ilhem Bentaleb, Marc Floquet, François Guillocheau, Agnès Lauriat-Rage, Didier Néraudeau, Jean-Pierre Platel et qui ont fourni un important travail de relecture. Je suis également reconnaissant envers toutes les personnes qui ont put m’aider sur le terrain, m’apporter une aide technique au laboratoire, m’aider dans la reflexion de ce travail : Didier Néraudeau, Philippe Courville, François Guillocheau, Loïc Villier, Ilhem Bentaleb, Jean-Marie Rouchy, Evelyne Goubert, Vincent Perrichot, Romain Vullo, Ronan Alain, Yves Quété, Jean Cornichet, Xavier Le Coz, Philippe Landrain, Veronique Berrou… De manière plus indirecte, mais tout aussi essentielle, je fais un clin d’œil à tous les copains qui m’ont supportés durant ces trois années : Julien, Myriam, Stéphane, Castor, Charly, César, Rodrigo, Katia, Olivier, Nico, Delphine, Romain, Vincent, Ronan, Catherine, Flo, Ben, Nico, Christelle, Stéphanie, Yael, Eric, Vincent, Ludo, Dom, Yannick, Cyrille, NicoK, Thierry, Florence, Seb, Deb, virginie, Nico Ploem’, Pec, Vinc’, Benoît, Marie, Anne, Claude, Estelle, Pierrot, Karen, Loïg, Flo, Renaud…et tous ceux que j’oublie !!! Enfin je dédie cette thèse à ma famille qui m’a soutenu pendant mon parcours et à Catherine qui a su m’épauler tout au long de ces trois années… Résumé D’après l’étude du Crétacé supérieur du NE du bassin-aquitain (Charentes, France) et du Néogène supérieur du bassin de Sorbas (SE Espagne), les huîtres fossiles peuvent être étudiées à plusieurs échelles. (1) Les assemblages d’ostréidés présentent une grande diversité de géométries spatiales. Leurs accumulations peuvent être soit tridimensionnelles (« intérieur » d’un banc) ou bidimensionnelles, avec l’implantation des organismes à la surface du sédiment. Une analyse taphonomique permet alors de discriminer ces différentes structures d’assemblages d’ostréidés en fonction de leur signification paléoécologique. L’organisation de ces assemblages (géométrie, densité des assemblages…) est bien souvent révélatrice de la dynamique sédimentaire, permet d’identifier des surfaces stratigraphiques remarquables et contribue à l’analyse des corps sédimentaires pour des interprétations séquentielles des milieux de dépôts. (2) Les espèces se répartissent sur un large spectre de milieux où chacune d’entre elles et leurs morphotypes sont inféodés à des conditions environnementales précises. Les nombreux paramètres climatiques et édaphiques contrôlant leur distribution ont ainsi pu être appréhendé (salinité, oxygénation, turbidité, nature du substratum, bathymétrie…). (3) Les coquilles enregistrent les fluctuations environnementales saisonnières via leur croissance accrétionaire. Une étude sclérochronologique et l’analyse de la composition en isotopes stables (C&O) de leurs stries de croissance permet de retrouver ce signal saisonnier. L’étude sclérochronologique permet également d’accéder à l’âge absolu des organismes et ainsi de reconstituer la dynamique des populations. Il est alors possible de quantifier le temps que représente une couche sédimentaire d’ordre annuel à pluriséculaire. L’étude isotopique stable a pu démontrer sur une espèce de Neopycnodonte actuelle que l’enregistrement par la coquille du signal isotopique de l’eau de mer n’était affecté que d’un effet biologique mineur. L’analyse de coquilles du Cénomanien supérieur et du Campanien supérieur témoigne de l’enregistrement d’un signal cyclique de la part des isotopes de l’oxygène démontre la présence d’une saisonnalité pour ces deux périodes. Abstract From the study of the Upper Cretaceous of the Aquitain Basin (Charentes, France) and of the Upper Neogene of the Sorbas basin (SE Spain), oysters can be useful at different scales of analyses: (1) Oyster assemblages have a great diversity of geometries. Their accumulation could be tridimensional, (in a shelly a bed) or bidimensional, with colonisation of a surface. A taphonomic analysis could be used to discriminate several find of assemblages according to their palaeoecological interests. Their organisation (geometry, population density…) often reveal sedimentary dynamics, can help to underline particular stratigraphic surfaces and can be used for the interpretation of sedimentary units. Such results could be useful for sequence stratigraphy analyses. (2) Species are distributed on numerous various environments. Each one of these species and their morphotypes are restricted to specific environmental conditions. Study of numerous parameters controlling their distributions (salinity, oxygenation, turbidity, quality of the substratum, bathymétry…) have be realised. Thus it is possible to reconstitute precisely climatic and edaphic parameters. (3) Shells record seasonal environmental fluctuations by sclerochronological studies and study of the stable isotopes composition (C&O) of their shell increments allow to characterise these seasonal variations. Sclerochronological studies allow to accede absolute age of each specimen too and then to reconstitute population dynamics. Therefore it is possible to characterise time of deposition of shellbeds (from several years to century time scale). Isotopic studies demonstrate that shell recording of sea water condition by Neopycnodonte is affected by a minor vital effect. The analyse of oxygene signal of Upper Cenomanian and Upper Campanian shells show a cyclic pattern that could be attributed to climatic seasonal variations. Sommaire Introduction générale 1 Chapitre I : Systématique des Ostréidés 5 1 Historique 7 2 Matériel & méthode 9 2.1 Modalité de définition d’une espèce 9 2.1.1 Caractères pris en compte 9 2.1.2 Plasticité et études morphométriques 12 2.2 Description 13 2.3 Répartition 13 2.4 Choix de la classification 16 3 Systématique des espèces étudiées 16 3.1 Ostreacea Rafinesque, 1815 16 3.1.1 Gryphaeidae Vyalov, 1936 16 3.1.1.1 Pycnodonteinae Stenzel, 1959 16 3.1.1.2 Exogyrinae Vyalov,1936 24 3.1.1.3 Gryphostreinae Conrad, 1865 33 3.1.2 Ostreidae Rafinesque, 1815 34 3.1.2.1 Crassostreinae Torigoe, 1981 34 3.1.2.2 Ostreinae Rafinesque, 1815 38 3.1.2.3 Lophinae Vyalov, 1936 39 4 Conclusions 45 4.1.1 Répartition stratigraphique des ostréidés 45 4.1.2 Répartition paléogéographique des ostréidés 46 Chapitre II : Cadre géologique 47 1 Introduction 49 2 Matériel & méthodes 49 2.1 Méthodes 49 2.1.1 Récolte et comptage des faunes 49 2.2 Cadre géologique 50 2.2.1 Le Cénomano-Turonien nord-aquitain 50 2.2.2 Le Campanien supérieur nord-aquitain 52 2.2.3 Le Mio-Pliocène andalous 54 2.2.3.1 Le bassin de Sorbas 54 2.2.3.2 Le bassin de Vera 57 3 Résultats 57 3.1 Le Cénomano-Turonien nord-aquitain 57 3.1.1 Cadeuil 57 3.1.2 Archingeay 59 3.1.3 Fouras « Bois-vert » 62 3.1.4 Fouras-Cadoret 64 3.1.5 Fouras « Fort Vaubant » 66 3.1.6 Ile d’Aix 68 3.1.7 Falaise de Piédemont 70 3.1.8 Ile d’Oléron 72 3.1.9 Ile Madame 75 3.1.10 Carrière de St Estèphe et ancienne carrière de Roulet 77 3.1.11 Falaise de Port-des-Barques 80 3.1.12 La Vallée 81 3.2 Le Campanien supérieur
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