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Evaniidae Chrysididae (Chrysidinae) Bethylidae (Epyrinae) Dryinidae

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Evaniidae Chrysididae (Chrysidinae) Bethylidae (Epyrinae) Dryinidae Evaniidae Chrysididae (Chrysidinae) Bethylidae (Epyrinae) Dryinidae (Gonatopodinae) Scoliidae (Scoliinae)) Formicidae (Myrmicinae) Ampulicidae (Dolichurinae) Sphecidae (Ammophilinae) Crabronidae (Crabroninae) Crabronidae (Bembicinae) Mellitidae (Dasypodinae) Mellitidae (Macropidinae) Apidae (Apinae) Colletidae (Hylaeinae) Vespidae (Eumeninae) Vespidae (Polistinae) Vespidae (Vespinae) Tiphiidae (Tiphiinae) Chyphotidae* Thynnidae (Methocinae) Thynnidae (Diamminae) Pompilidae (Ceropalinae) Pompilidae (Pepsinae) Character 1: 3rd valvula Pompilidae (Pompilinae) Parsimony reconstruction (Unordered) [Steps: 6] Sapygidae (Sapyginae) Myrmosidae (Myrmosinae) entire Mutillidae (Dasylabrinae) subdivided Mutillidae (Mutillinae) Mutillidae (Myrmillinae) Evaniidae Chrysididae (Chrysidinae) Bethylidae (Epyrinae) Dryinidae (Gonatopodinae) Scoliidae (Scoliinae)) Formicidae (Myrmicinae) Ampulicidae (Dolichurinae) Sphecidae (Ammophilinae) Crabronidae (Crabroninae) Crabronidae (Bembicinae) Mellitidae (Dasypodinae) Mellitidae (Macropidinae) Apidae (Apinae) Colletidae (Hylaeinae) Vespidae (Eumeninae) Vespidae (Polistinae) Vespidae (Vespinae) Tiphiidae (Tiphiinae) Chyphotidae* Thynnidae (Methocinae) Thynnidae (Diamminae) Pompilidae (Ceropalinae) Pompilidae (Pepsinae) Character 2: Ratio of proximal and distal lobes Pompilidae (Pompilinae) Parsimony reconstruction (Unordered) [Steps: 7] Sapygidae (Sapyginae) Myrmosidae (Myrmosinae) proximal lobe shorter than distal Mutillidae (Dasylabrinae) more or less equal distal lobe shorter than proximal Mutillidae (Mutillinae) Mutillidae (Myrmillinae) Evaniidae Chrysididae (Chrysidinae) Bethylidae (Epyrinae) Dryinidae (Gonatopodinae) Scoliidae (Scoliinae)) Formicidae (Myrmicinae) Ampulicidae (Dolichurinae) Sphecidae (Ammophilinae) Crabronidae (Crabroninae) Crabronidae (Bembicinae) Mellitidae (Dasypodinae) Mellitidae (Macropidinae) Apidae (Apinae) Colletidae (Hylaeinae) Vespidae (Eumeninae) Vespidae (Polistinae) Vespidae (Vespinae) Tiphiidae (Tiphiinae) Chyphotidae* Thynnidae (Methocinae) Thynnidae (Diamminae) Pompilidae (Ceropalinae) Pompilidae (Pepsinae) Character 3: Degree of lobe sclerotisation Pompilidae (Pompilinae) Parsimony reconstruction (Unordered) [Steps: 3] Sapygidae (Sapyginae) Myrmosidae (Myrmosinae) proximal lobe less sclerotised than distal Mutillidae (Dasylabrinae) lobes more or less equally sclerotised proximal lobe more sclerotised than distal Mutillidae (Mutillinae) Mutillidae (Myrmillinae) Evaniidae Chrysididae (Chrysidinae) Bethylidae (Epyrinae) Dryinidae (Gonatopodinae) Scoliidae (Scoliinae)) Formicidae (Myrmicinae) Ampulicidae (Dolichurinae) Sphecidae (Ammophilinae) Crabronidae (Crabroninae) Crabronidae (Bembicinae) Mellitidae (Dasypodinae) Mellitidae (Macropidinae) Apidae (Apinae) Colletidae (Hylaeinae) Vespidae (Eumeninae) Vespidae (Polistinae) Vespidae (Vespinae) Tiphiidae (Tiphiinae) Chyphotidae* Thynnidae (Methocinae) Thynnidae (Diamminae) Pompilidae (Ceropalinae) Pompilidae (Pepsinae) Character 4: 2nd valvifer - 3rd valvula junction Pompilidae (Pompilinae) Parsimony reconstruction (Unordered) [Steps: 9] Sapygidae (Sapyginae) Myrmosidae (Myrmosinae) narrow elastic cuticle Mutillidae (Dasylabrinae) wide elastic cuticle sclerotised bar Mutillidae (Mutillinae) Mutillidae (Myrmillinae) Evaniidae Chrysididae (Chrysidinae) Bethylidae (Epyrinae) Dryinidae (Gonatopodinae) Scoliidae (Scoliinae)) Formicidae (Myrmicinae) Ampulicidae (Dolichurinae) Sphecidae (Ammophilinae) Crabronidae (Crabroninae) Crabronidae (Bembicinae) Mellitidae (Dasypodinae) Mellitidae (Macropidinae) Apidae (Apinae) Colletidae (Hylaeinae) Vespidae (Eumeninae) Vespidae (Polistinae) Vespidae (Vespinae) Tiphiidae (Tiphiinae) Chyphotidae* Thynnidae (Methocinae) Thynnidae (Diamminae) Pompilidae (Ceropalinae) Pompilidae (Pepsinae) Character 5: Junction between 3rd valvula lobes Pompilidae (Pompilinae) Parsimony reconstruction (Unordered) [Steps: 6] Sapygidae (Sapyginae) Myrmosidae (Myrmosinae) articulation Mutillidae (Dasylabrinae) narrow elastic cuticle Mutillidae (Mutillinae) Mutillidae (Myrmillinae) Evaniidae Chrysididae (Chrysidinae) Bethylidae (Epyrinae) Dryinidae (Gonatopodinae) Scoliidae (Scoliinae)) Formicidae (Myrmicinae) Ampulicidae (Dolichurinae) Sphecidae (Ammophilinae) Crabronidae (Crabroninae) Crabronidae (Bembicinae) Mellitidae (Dasypodinae) Mellitidae (Macropidinae) Apidae (Apinae) Colletidae (Hylaeinae) Vespidae (Eumeninae) Vespidae (Polistinae) Vespidae (Vespinae) Tiphiidae (Tiphiinae) Chyphotidae* Thynnidae (Methocinae) Thynnidae (Diamminae) Pompilidae (Ceropalinae) Pompilidae (Pepsinae) Character 6: 3vv proximally during stinging Pompilidae (Pompilinae) Parsimony reconstruction (Unordered) [Steps: 4] Sapygidae (Sapyginae) Myrmosidae (Myrmosinae) lifted Mutillidae (Dasylabrinae) rotated touching Mutillidae (Mutillinae) Mutillidae (Myrmillinae) Evaniidae Chrysididae (Chrysidinae) Bethylidae (Epyrinae) Dryinidae (Gonatopodinae) Scoliidae (Scoliinae)) Formicidae (Myrmicinae) Ampulicidae (Dolichurinae) Sphecidae (Ammophilinae) Crabronidae (Crabroninae) Crabronidae (Bembicinae) Mellitidae (Dasypodinae) Mellitidae (Macropidinae) Apidae (Apinae) Colletidae (Hylaeinae) Vespidae (Eumeninae) Vespidae (Polistinae) Vespidae (Vespinae) Tiphiidae (Tiphiinae) Chyphotidae* Thynnidae (Methocinae) Thynnidae (Diamminae) Pompilidae (Ceropalinae) Pompilidae (Pepsinae) Character 7: Sting shape Pompilidae (Pompilinae) Parsimony reconstruction (Unordered) [Steps: 1] Sapygidae (Sapyginae) Myrmosidae (Myrmosinae) decurved Mutillidae (Dasylabrinae) coiled Mutillidae (Mutillinae) Mutillidae (Myrmillinae) Evaniidae Chrysididae (Chrysidinae) Bethylidae (Epyrinae) Dryinidae (Gonatopodinae) Scoliidae (Scoliinae)) Formicidae (Myrmicinae) Ampulicidae (Dolichurinae) Sphecidae (Ammophilinae) Crabronidae (Crabroninae) Crabronidae (Bembicinae) Mellitidae (Dasypodinae) Mellitidae (Macropidinae) Apidae (Apinae) Colletidae (Hylaeinae) Vespidae (Eumeninae) Vespidae (Polistinae) Vespidae (Vespinae) Tiphiidae (Tiphiinae) Chyphotidae* Thynnidae (Methocinae) Thynnidae (Diamminae) Pompilidae (Ceropalinae) Pompilidae (Pepsinae) Character 8: Sting function Pompilidae (Pompilinae) Parsimony reconstruction (Unordered) [Steps: 6] Sapygidae (Sapyginae) Myrmosidae (Myrmosinae) defense only Mutillidae (Dasylabrinae) defense and killing defense and immobilizing Mutillidae (Mutillinae) Mutillidae (Myrmillinae).
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