AAMERICAN MUSEUM NOVITATES Published by Number 1034 the AMERICAN MUSEUM of NATURAL HISTORY June 26, 1939 New York City

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AAMERICAN MUSEUM NOVITATES Published by Number 1034 the AMERICAN MUSEUM of NATURAL HISTORY June 26, 1939 New York City AAMERICAN MUSEUM NOVITATES Published by Number 1034 THE AMERICAN MUSEUM OF NATURAL HISTORY June 26, 1939 New York City A STUDY OF TETRAMERYX AND ASSOCIATED FOSSILS FROM PAPAGO SPRING CAVE, SONOITA, ARIZONA BY EDWIN H. COLBERT AND ROBERT G. CHAFFEE INTRODUCTION During the summer of 1934, Messrs. of its horns. Not only is Tetrameryx of Quentin Roosevelt and J. W. Burden ex- interest because of its anatomnical features, plored a limestone cave near Papago Spring, but also because of its association, for some two and one-half miles southwest of Roosevelt and Burden found pottery frag- Sonoita, Santa Cruz County, Arizona, and, ments and other artifacts in Papago Spring as a result of their field work, discovered cave that would seem to point to the fact various skull and skeletal remains of the that Tetrameryx onusrosagris was contem- Pleistocene antilocaprid, Tetrameryx. poraneous with early man in southwestern These fossils were sent to the American United States. Moreover, among the as- Museum, where they were prepared and sociated mammalian remains is a bone of stored. In November of the same year, the Pleistocene North American horse, Roosevelt and Burden' described some of Equus, which also once lived in the South- the material that they had collected, allot- west with early man. Evidently these ting it to a new species of the genus Tetra- fossil animals, now extinct in North Amer- meryx, namely: Tetrameryx onusrosagris. ica, persisted on this continent to within a In the spring of 1936, these two gentle- few thousands of years ago, as did many men made an additional exploration of the other typically "Pleistocene" mammals, Papago Spring cave, with the result that and the discovery at Papago Spring cave additional specimens of Tetrameryx were constitutes one more record of the post- discovered, so that we now have fairly Pleistocene persistence of Pleistocene types. complete skull and skeletal material of The purpose of this contribution is to Tetrameryx onusrosagris, as well as a few make a detailed study of Tetrameryx onus- other fossils that were associated with this rosagris, comparing it point for point with extinct antelope. the recent Antilocapra americana, and so Because of the complete series of fossil far as the known material permits, with skulls and skeletal remains from Sonoita, other fossil antilocaprids. The fossils as- we are able for the first time to make an sociated with Tetrameryx in the Papago adequate comparative study of this now Spring cave will also be described. extinct genus. This form is of particular Acknowledgments are due to Messrs. interest in that it resembles very closely Roosevelt and Burden for the interest they the modern pronghorn (as will be shown have shown in the Papago Spring cave de- in posit, and for their laborious and careful below) most of its structural characters, work of collecting the fossils hereinafter yet is quite different as to the development described. The American Museum of 1 Roosevelt, Quentin, and Burden, J. W., 1934, Natural History is also indebted to these "A New Species of Antilocaprine, Tetrameryx onus- rosagris, from a Pleistocene Cave Deposit in Southern gentlemen for their kindness in presenting Arizona." Amer. Mus. Novitates, No. 754. the fossils to this institution. OCCURRENCE OF THE FOSSILS Papago Spring cave is located in a lime- Arizona. This locality is in the basin stone hill in the Canelo Hills, some two range section of western United States, and a half miles southwest of Sonoita, characterized by block mountains an(d 2 AMERICAN MUSEUM NO VITA TES[[No. 1034 igneous cores separated by wide bolson is probably of a relatively recent geologic deposits. Faulting, folding and tilting of date, having been formed by solution dur- the sedimentary beds is common in this ing Pleistocene, or perhaps late Tertiary region, and the deposits in which Papago times. Certainly the deposits within the Spring cave is formed are tilted to an angle cave are all of Pleistocene or post-Pleisto- of some thirty or forty (legrees. The cene age. Fig. 1. (Above) View to the northwest from the summit of the cave-hill. (Below) View of the cave-hill from a point one-fourth of a mile southwest of the cave entrance. The cave entrance is on the side of the hill facing the observer, but is screened from view by the trees. limestones of the Canelo Hills are of De- Mr. Burdlen's (lescription of the cave is vonian, Carboniferous and Permian age,. presented at this point, to serve as an ex- relate(d to the Martin and Tornado lime- planation for the figure of the cave, shown stones of a(djacent regions. In this con- in the accompanying illustration. nection it rnight be saidl that some pro- Entrance "A" is the largest of the two known ductids found in the wall of the cave would entrances and is simply a direct drop of approxi- mately twenty-five feet. Its walls bear signs indicate a probable Permian age for the of water erosion and it seems likely that a beds containing them. The cave, itself, stream once ran through it into the cave. At 1939 ] TETRAMERYX AND OTHER FOSSILS FROM ARIZONA 3 its northwest end big rocks and boulders have and then goes up at a 15 degree angle as far as fallen in. Its mouth is about sixty feet up the the one hundred and sixty-five foot mark. It hillside and is sheltered by overhanging rocks and then continues on a level as far as the two a tree. hundred and ninety-five foot mark, where it Entrance "B" is the one we used all the time, drops directly down in a corkscrew shaped and it is exceedingly narrow. Its mouth is tunnel. about forty feet above the ground (i.e., up the The height of the tunnel varies from three feet hill) and is almost hidden by a dense clump of to about twenty feet, while the main cavern is bushes. This entrance consists of a narrow about thirty feet high in its widest part. bending passage between large, loose boulders, The first fossil deposit contained the first and it descends at an angle of 45 to 60 degrees skull we ever discovered, together with three I Fig. 2. (Left) View from the top of the tunnel, showing fragmentary fossils exposed on a ledge. (Right) Interior of Papago Spring cave, near the main fossil deposit. for about twelve feet and then turns northeast other skulls and numerous bones. This deposit into a small cavern. This in turn opens first was apparently connected with deposit 2, be- into a still larger cavern, and, finally, into the cause the strata seemed to continue up to the main room of the cave. The floor, all the way tunnel mouth. The second deposit has ap- down the cave, from the first small cavern to parently been exposed to the air too long, for the sixty foot mark, descends at an angle of the bones it contains are exceedingly brittle. 25 to 30 degrees and is composed of loose rocks The third deposit has furnished us with most of and soft earth. From the sixty foot to the our bones. It is a "jutting-out" promontory eighty foot mark it dips still more to about of hard limestone-cemented rubble almost en- 45 degrees. It then levels out (down the tunnel) tirely undermined, and was probably a section to about a 10 degree angle as far as the one of the old cave floor. (Burden, J. W., Letter to hundred and ten foot mark, drops a bit more Walter Granger, dated April 29, 1936.) Tunnel continue-s approximately 80 feet and then becomes impassa6le 2' Ii0'- N MAP of PAPAGO SPRING CAVE by J.W. Burden and Quent;n Roosevel+ Loos0 6one fragments Pottery 6 First Fossil deposit .Se,.nvIA X \ Third I.I <> ~)Entfance B Distance in feet approximate Fig. 3. Map of Papago Spring cave, near Sonoita, Arizona. (Drawn by J. W. Burden and Quentin Roosevelt.) Ca) fti 4) cs .2 c3 k C; cq -. ~al -4- N Nq E.4- 0a) nN "3 ,0 6 AMERICAN MUSEUM NOVITATES [No. 1034 DESCRIPTIONS AND DISCUSSIONS Tetrameryx onusrosagris Roosevelt uppermost Pleistocene or subrecent. Papago and Burden Spring cave, south of Sonoita, Arizona. DIAGNOSIS.-Approximately equal to Anti- Tetrameryx onusrosagris, ROOSEVELT AND locapra in size, but with shorter, stockier limbs BURDEN, 1934, Amer. Mus. Novitates, No. 754. and a heavier body. Orbits less protruding Stockoceros onusrosagris, FRICK, 1937, Bull. than in the modern pronghorn, and but slightly Amer. Mus. Nat. Hist., LXIX, p. 527. turned anteriorly. Canine-premolar diastema TYPE.-Amer. Mus. No. 22488, a broken skull and muzzle relatively shorter than in Antilo- with the horn cores, cranium, basicranium, capra. Two pairs of subequal horn cores with cheek-teeth and nasals preserved. Maxillae elliptical cross-sections, arising on either side broken, premaxillae missing. from a common base above the orbit. These REFERRED SPEcIMENs.-(Referred to this horn cores branch in an antero-posterior plane species in the original type description.) and at an angle of about 45 degrees. (T. Amer. Mus. No. 22489, a broken skull with shuleri, the generic type, has a long posterior right horn cores, occiput, basicranium, right branch; T. (Stockocero8) conklingi has horn upper M3, left upper M2 and M3, maxillae cores more nearly parallel to each other; T. and nasals broken, premaxillae missing. onu8rosagris ref. from Burnett cave has a 27 Amer. Mus. No. 22484, fragment of occiput, degree angle between the branches; five detached horn cores, and four fragments Hayoceros of has an elongated cross-section of the anterior rami. core as in Antilocapra, and a sub-rounded Amer.
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